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域 (生物)

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生物分類法上八個主要的分類單元,生物被分成三個域,並進一步被劃分成其他分類單元。本圖沒有顯示較小的中間分類單元。

生物分類學中,(英語:domain、empire拉丁語regio)或總界superkingdom)是美國生物學家卡爾·沃斯1990年设计的三域系統中最高的分類單元,該系統中,生命之樹英语Tree of life (biology)包含細菌古菌真核生物等三個域,其中前兩者為原核生物,不具有細胞核;後者則具有細胞核和内膜系统包裹的细胞器[參⁠ 1]

三域生物特色

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基於16S 核糖體RNA序列的系統發生樹,顯示了可明顯區別的三支:細菌、古菌和真核生物

古菌

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古菌屬於單細胞原核生物,外形類似細菌,但部分分子序列與代謝途徑與真核生物較相近,其細胞膜醚酯英语Ether lipid所組成,不同於另兩域生物細胞膜主要成分為酯類[參⁠ 2]。早期發現的古菌多為嗜極生物,後來漸發現古菌普遍存在各種環境中[參⁠ 3]

細菌

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細菌也屬於單細胞原核生物,其細胞膜為酯類組成,廣泛分布於不同環境中,外形、營養方式與生長環境都有很高的多樣性。細菌的大小一般為0.5-5微米,少數特例如纳米比亚嗜硫珠菌[參⁠ 4]費氏刺骨魚菌英语Epulopiscium fishelsoni可大至0.5毫米而能以肉眼觀察[參⁠ 5],而小型者如黴漿菌僅有約0.3微米,與大型的病毒大小相仿[參⁠ 6],甚至還有更小的超微細菌英语Ultramicrobacteria[參⁠ 7]

真核生物

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真核生物具有細胞核以及多種膜結構組成的內膜系統,包括動物植物真菌[參⁠ 8]。2005年,國際原生生物學家協會英语International Society of Protistologists將真核生物分為六个超类群(supergroups),包括後鞭毛生物古蟲界變形蟲界囊泡藻界有孔蟲界原始色素體生物[參⁠ 9][參⁠ 10],但隨即有學者質疑囊泡藻界並非單系群,並有後續研究將其拆分[參⁠ 11][參⁠ 12]。近年來持續有許多真核生物的分類理論被提出[參⁠ 13][參⁠ 14]

分類

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2002年,英國生物學家湯瑪斯·卡弗利爾-史密斯提出了新壁總域的概念,即古菌與真核生物共同組成一演化支,為細菌中放線菌門姊妹群[參⁠ 15]。後續分子種系發生學的研究亦支持古菌與真核生物的親緣關係較與細菌的親緣關係近[參⁠ 16]

其他分類系統

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三域系統(上)與泉古菌假说英语Eocyte hypothesis(下)

除了三域系統外,其他較常用的分類系統還包括:

參考資料

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    參:

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  6. ^ Robertson J, Gomersall M, Gill P. Mycoplasma hominis: growth, reproduction, and isolation of small viable cells. Journal of Bacteriology. November 1975, 124 (2): 1007–18. PMC 235991可免费查阅. PMID 1102522. 
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  12. ^ Parfrey, Laura W.; Barbero, Erika; Lasser, Elyse; Dunthorn, Micah; Bhattacharya, Debashish; Patterson, David J. & Katz, Laura A. Evaluating support for the current classification of eukaryotic diversity. PLoS Genetics. 2006, 2 (12): e220. PMC 1713255可免费查阅. PMID 17194223. doi:10.1371/journal.pgen.0020220. 
  13. ^ Burki, Fabien; Shalchian-Tabrizi, Kamran; Minge, Marianne; Skjæveland, Åsmund; Nikolaev, Sergey I.; Jakobsen, Kjetill S. & Pawlowski, Jan. Butler, Geraldine , 编. Phylogenomics reshuffles the eukaryotic supergroups. PLoS ONE. 2007, 2 (8): e790. Bibcode:2007PLoSO...2..790B. PMC 1949142可免费查阅. PMID 17726520. doi:10.1371/journal.pone.0000790. 
  14. ^ Kim, E.; Graham, L. E. & Redfield, Rosemary Jeanne. Redfield, Rosemary Jeanne , 编. EEF2 analysis challenges the monophyly of Archaeplastida and Chromalveolata. PLoS ONE. 2008, 3 (7): e2621. Bibcode:2008PLoSO...3.2621K. PMC 2440802可免费查阅. PMID 18612431. doi:10.1371/journal.pone.0002621. 
  15. ^ Cavalier-Smith T. The phagotrophic origin of eukaryotes and phylogenetic classification of Protozoa. Int. J. Syst. Evol. Microbiol. March 2002, 52 (Pt 2): 297–354 [2013-07-13]. PMID 11931142. (原始内容存档于2017-07-29). 
  16. ^ Ciccarelli FD, Doerks T, von Mering C, Creevey CJ, Snel B, Bork P. Toward automatic reconstruction of a highly resolved tree of life (PDF). Science. 2006, 311 (5765): 1283–7 [2019-07-13]. Bibcode:2006Sci...311.1283C. PMID 16513982. doi:10.1126/science.1123061. (原始内容 (PDF)存档于2016-03-04). 
  17. ^ Mayr, Ernst. Two empires or three?. Proc Natl Acad Sci USA. 1998, 95 (17): 9720–9723 [5 September 2011]. Bibcode:1998PNAS...95.9720 请检查|bibcode=值 (帮助). PMC 33883可免费查阅. PMID 9707542. doi:10.1073/pnas.95.17.9720. (原始内容存档于2019-06-13). 
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  19. ^ Archibald, John M. The eocyte hypothesis and the origin of eukaryotic cells. PNAS. 23 December 2008, 105 (51): 20049–20050. Bibcode:2008PNAS..10520049A. PMC 2629348可免费查阅. PMID 19091952. doi:10.1073/pnas.0811118106. 
  20. ^ Lake, James A.; Henderson, Eric; Oakes, Melanie; Clark, Michael W. Eocytes: A new ribosome structure indicates a kingdom with a close relationship to eukaryotes. PNAS. June 1984, 81 (12): 3786–3790. Bibcode:1984PNAS...81.3786L. PMC 345305可免费查阅. PMID 6587394. doi:10.1073/pnas.81.12.3786. 
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  23. ^ 中國大百科全書 生物學分冊

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