List of Odonata species of Australia

Alpine darner, Austroaeschna flavomaculata

This is a list of species of damselflies and dragonflies recorded in Australia.

Common names of species are linked, beside their scientific names.

The list is split into two groups: damselflies (suborder Zygoptera) and other dragonflies (infraorder Anisoptera). Those groups are organized in Families and then Genera and Species.

Zygoptera (damselflies)

Argiolestidae

genus: Archiargiolestes

• Midget flatwing, Archiargiolestes parvulus
• Tiny flatwing, Archiargiolestes pusillissimus
• Little flatwing, Archiargiolestes pusillus

genus: Austroargiolestes

• New England flatwing, Austroargiolestes alpinus
• Flame flatwing, Austroargiolestes amabilis
• Tropical flatwing, Austroargiolestes aureus
• Barrington flatwing, Austroargiolestes brookhousei
• Powdered flatwing, Austroargiolestes calcaris
• Milky flatwing, Austroargiolestes christine
• Golden flatwing, Austroargiolestes chrysoides
• Azure flatwing, Austroargiolestes elke
• Common flatwing, Austroargiolestes icteromelas
• Sydney flatwing, Austroargiolestes isabellae

genus: Griseargiolestes

• Coastal flatwing, Griseargiolestes albescens
• Turquoise flatwing, Griseargiolestes bucki
• Grey-chested flatwing, Griseargiolestes eboracus
• Springs flatwing, Griseargiolestes fontanus
• Grey flatwing, Griseargiolestes griseus
• Alpine flatwing, Griseargiolestes intermedius
• Metallic flatwing, Griseargiolestes metallicus

genus: Miniargiolestes

• Stream flatwing, Miniargiolestes minimus

genus: Podopteryx

• Treehole flatwing, Podopteryx selysi

Coenagrionidae

genus: Aciagrion

• Blue slim, Aciagrion fragilis

genus: Agriocnemis

• Silver wisp, Agriocnemis argentea
• Tropical wisp, Agriocnemis dobsoni
• Pilbara wisp, Agriocnemis kunjina
• Pygmy wisp, Agriocnemis pygmaea
• Red-rumped wisp, Agriocnemis rubricauda
• Agriocnemis thoracalis

genus: Archibasis

• Blue-banded longtail, Archibasis mimetes

genus: Argiocnemis

• Red-tipped shadefly, Argiocnemis rubescens

genus: Austroagrion

• South-western billabongfly, Austroagrion cyane
• Northern billabongfly, Austroagrion exclamationis
• Pilbara billabongfly, Austroagrion pindrina
• Eastern billabongfly, Austroagrion watsoni

genus: Austrocnemis

• Tiny longlegs, Austrocnemis maccullochi
• Kimberley longlegs, Austrocnemis obscura
• Splendid longlegs, Austrocnemis splendida

genus: Caliagrion

• Large riverdamsel, Caliagrion billinghursti

genus: Ceriagrion

• Redtail, Ceriagrion aeruginosum

genus: Coenagrion

• Swamp bluet, Coenagrion lyelli

genus: Ischnura

• Aurora bluetail, Ischnura aurora
• Common bluetail, Ischnura heterosticta
• Colourful bluetail, Ischnura pruinescens

genus: Pseudagrion

• Gold-fronted riverdamsel, Pseudagrion aureofrons
• Northern riverdamsel, Pseudagrion cingillum
• Flame-headed riverdamsel, Pseudagrion ignifer
• Dusky riverdamsel, Pseudagrion jedda
• Citrine-headed riverdamsel, Pseudagrion lucifer
• Blue riverdamsel, Pseudagrion microcephalum

genus: Teinobasis

• Red-breasted longtail, Teinobasis rufithorax

genus: Xanthagrion

• Red and blue damsel, Xanthagrion erythroneurum

Hemiphlebiidae

genus: Hemiphlebia

• Ancient greenling, Hemiphlebia mirabilis

Isostictidae

genus: Austrosticta

• Northern pondsitter, Austrosticta fieldi
• Eastern pondsitter, Austrosticta frater
• Kimberley pondsitter, Austrosticta soror

genus: Eurysticta

• Pilbara pin, Eurysticta coolawanyah
• Coomalie pin, Eurysticta coomalie
• Kimberley pin, Eurysticta kununurra
• Queensland pin, Eurysticta reevesi

genus: Labidiosticta

• Large wiretail, Labidiosticta vallisi

genus: Lithosticta

• Rock narrow-wing, Lithosticta macra

genus: Neosticta

• Southern pinfly, Neosticta canescens
• Tropical pinfly, Neosticta fraseri
• Forest pinfly, Neosticta silvarum

genus: Oristicta

• Slender wiretail, Oristicta filicicola
• Elegant wiretail, Oristicta rosendaleorum

genus: Rhadinosticta

• Northern wiretail, Rhadinosticta banksi
• Wiretail, Rhadinosticta handschini
• Powdered wiretail, Rhadinosticta simplex

Lestidae

genus: Austrolestes

• Western ringtail, Austrolestes aleison
• Slender ringtail, Austrolestes analis
• Blue ringtail, Austrolestes annulosus
• Inland ringtail, Austrolestes aridus
• Metallic ringtail, Austrolestes cingulatus
• Northern ringtail, Austrolestes insularis
• Iota ringtail, Austrolestes io
• Wandering ringtail, Austrolestes leda
• Dune ringtail, Austrolestes minjerriba
• Cup ringtail, Austrolestes psyche

genus: Indolestes

• Small reedling, Indolestes alleni
• Cave reedling, Indolestes obiri
• Slender reedling, Indolestes tenuissimus

genus: Lestes

• Dusky spreadwing, Lestes concinnus

Lestoideidae

genus: Diphlebia

• Sapphire rockmaster, Diphlebia coerulescens
• Tropical rockmaster, Diphlebia euphaeoides
• Giant rockmaster, Diphlebia hybridoides
• Whitewater rockmaster, Diphlebia lestoides
• Arrowhead rockmaster, Diphlebia nymphoides

genus: Lestoidea

• Large bluestreak, Lestoidea barbarae
• Short-tipped bluestreak, Lestoidea brevicauda
• Common bluestreak, Lestoidea conjuncta
• Mount Lewis bluestreak, Lestoidea lewisiana

Platycnemididae

genus: Nososticta

• Black-winged threadtail, Nososticta baroalba
• Green-blue threadtail, Nososticta coelestina
• Northern threadtail, Nososticta fraterna
• Spot-winged threadtail, Nososticta kalumburu
• Koolpinyah threadtail, Nososticta koolpinyah
• Citrine threadtail, Nososticta koongarra
• Malachite threadtail, Nososticta liveringa
• Striped threadtail, Nososticta mouldsi
• Pilbara threadtail, Nososticta pilbara
• Orange threadtail, Nososticta solida
• Fivespot threadtail, Nososticta solitaria
• Melville Island threadtail, Nososticta taracumbi

Synlestidae

genus: Chorismagrion

• Pretty relict, Chorismagrion risi

genus: Episynlestes

• Southern whitetip, Episynlestes albicauda
• Tropical whitetip, Episynlestes cristatus
• Intermediate whitetip, Episynlestes intermedius

genus: Synlestes

• Forest needle, Synlestes selysi
• Tropical needle, Synlestes tropicus
• Bronze needle, Synlestes weyersii

Anisoptera (dragonflies)

Aeshnidae

genus: Acanthaeschna

• Thylacine darner, Acanthaeschna victoria (Coastal southern Queensland and northern New South Wales)

genus: Adversaeschna

• Blue-spotted hawker, Adversaeschna brevistyla (Coastal areas from Cooktown in northern Queensland, south to Victoria, and the Pilbara in Western Australia)

genus: Agyrtacantha

• Trifid duskhawker, Agyrtacantha dirupta (Cape York Peninsula)

genus: Anaciaeschna

• Australasian duskhawker, Anaciaeschna jaspidea (Coastal Queensland and Northern Territory)

genus: Anax

• Kimberley emperor, Anax georgius (The Kimberley)
• Green emperor, Anax gibbosulus (Coastal northern Australia from the Kimberley to the Gold Coast)
• Lesser green emperor, Anax guttatus (Coastal northern Australia from the Kimberley to the Gold Coast)
• Australian emperor, Anax papuensis (All Australia and islands)

genus: Antipodophlebia

• Terrestrial evening darner, Antipodophlebia asthenes

genus: Austroaeschna

Lesser Tasmanian darner Austroaeschna hardyi

• Western darner, Austroaeschna anacantha (Coastal south-western Western Australia)
• Mountain darner, Austroaeschna atrata (Australian Alps)
• S-spot darner, Austroaeschna christine (Central coast of Queensland)
• Wallum darner, Austroaeschna cooloola Southern coast of Queensland
• Eungella darner, Austroaeschna eungella (Central coast of Queensland)
• Alpine darner, Austroaeschna flavomaculata (Australian Alps)
• Lesser Tasmanian darner, Austroaeschna hardyi (Tasmania)
• Whitewater darner, Austroaeschna inermis (Coastal south-eastern Australia)
• Grampians darner, Austroaeschna ingrid (Western Victoria)
• Carnarvon darner, Austroaeschna muelleri (Inland southern Queensland)
• Multi-spotted darner, Austroaeschna multipunctata (Southern coastal New South Wales to Victoria)
• Sydney mountain darner, Austroaeschna obscura (Mountains around Sydney)
• Swamp darner, Austroaeschna parvistigma (Coastal New South Wales, Victoria, southern South Australia and Tasmania)
• Inland darner, Austroaeschna pinheyi (Carnarvon Gorge, Queensland)
• Forest darner, Austroaeschna pulchra (Coastal southern Queensland, New South Wales and Victoria)
• Sigma darner, Austroaeschna sigma (Coastal southern Queensland and New South Wales)
• Tropical unicorn darner, Austroaeschna speciosa (Central coast of Queensland)
• Conehead darner, Austroaeschna subapicalis (Coastal New South Wales and Victoria)
• Tasmanian darner, Austroaeschna tasmanica (Tasmania)
• Unicorn darner, Austroaeschna unicornis (Southern Queensland to coastal New South Wales, Victoria, Tasmania and South Australia)

genus: Austrogynacantha

• Australian duskhawker, Austrogynacantha heterogena (Across Australia north of about Sydney)

genus: Austrophlebia

• Southern giant darner, Austrophlebia costalis
• Northern giant darner, Austrophlebia subcostalis

genus: Dendroaeschna

• Wide-faced darner, Dendroaeschna conspersa

genus: Dromaeschna

• Green-striped darner, Dromaeschna forcipata (Coast of Cape York Peninsula, Queensland)
• Ochre-tipped darner, Dromaeschna weiskei (Coast of Cape York Peninsula, Queensland)

genus: Gynacantha

• Lesser duskhawker, Gynacantha dobsoni (Northern Australia)
• Slender duskhawker, Gynacantha kirbyi (Northern Queensland)
• Paddle-tipped duskhawker, Gynacantha mocsaryi (Northern Queensland)
• Cave duskhawker, Gynacantha nourlangie (Inland and northern Australia)
• Grey duskhawker, Gynacantha rosenbergi (Northern Queensland)

genus: Notoaeschna

• Northern riffle darner, Notoaeschna geminata
• Southern riffle darner, Notoaeschna sagittata

genus: Spinaeschna

• Southern cascade darner, Spinaeschna tripunctata
• Tropical cascade darner, Spinaeschna watsoni

genus: Telephlebia

• Southern evening darner, Telephlebia brevicauda
• Northern evening darner, Telephlebia cyclops
• Eastern evening darner, Telephlebia godeffroyi
• Tropical evening darner, Telephlebia tillyardi
• Coastal evening darner, Telephlebia tryoni
• Carnarvon evening darner, Telephlebia undia

Austropetaliidae

genus: Archipetalia

• Tasmanian redspot, Archipetalia auriculata (Tasmania)

genus: Austropetalia

• Northern redspot, Austropetalia annaliese (Eastern New South Wales)
• Waterfall redspot, Austropetalia patricia (Coastal New South Wales)
• Alpine redspot, Austropetalia tonyana (Southern New South Wales and Victoria between 600–1800 metres elevation)

Corduliidae

genus: Hemicordulia

• Australian emerald, Hemicordulia australiae (Widespread in eastern and south-western Australia)
• Fat-bellied emerald, Hemicordulia continentalis (Coastal Queensland)
• Desert emerald, Hemicordulia flava (Central Australia)
• Yellow-spotted emerald, Hemicordulia intermedia (Northern Australia)
• Slender emerald, Hemicordulia kalliste (Northern Australia)
• Pilbara emerald, Hemicordulia koomina (Pilbara, Western Australia)
• Superb emerald, Hemicordulia superba (Northern New South Wales and south-eastern Queensland)
• Tau emerald, Hemicordulia tau (Widespread across Australia)

genus: Pentathemis

• Metallic tigerhawk, Pentathemis membranulata (Northern Australia)

genus: Procordulia

• Western swamp emerald, Procordulia affinis (South-western Western Australia)
• Eastern swamp emerald, Procordulia jacksoniensis (Eastern New South Wales, Victoria, south-eastern South Australia, Tasmania)

Gomphidae

genus: Antipodogomphus

• Southern dragon, Antipodogomphus acolythus Queensland except for Cape York and south-west, New South Wales except south-east and Victoria
• Top End dragon, Antipodogomphus dentosus Top End of Northern Territory
• Cape York dragon, Antipodogomphus edentulus Cape York, Queensland
• Pilbara dragon, Antipodogomphus hodgkini Pilbara region of Western Australia
• Northern dragon, Antipodogomphus neophytus Northern Australia, from Western Australia to Queensland
• Spinehead dragon, Antipodogomphus proselythus Coastal Queensland

genus: Armagomphus

• Armourtail, Armagomphus armiger South-western Western Australia.

Austrogomphus guerini

genus: Austroepigomphus

subgenus Austroepigomphus

• Twinspot hunter, Austroepigomphus praeruptus (South-eastern South Australia, Victoria)

subgenus Xerogomphus

• Western red hunter, Austrogomphus gordoni (Western Australia except the Kimberleys and central Australia)
• Flame-tipped hunter, Austrogomphus turneri (Northern Australia, from Western Australia to Queensland)

genus: Austrogomphus

subgenus Austrogomphus

• Murray River hunter, Austrogomphus angelorum (South-eastern South Australia, Victoria, western New South Wales)
• Toothed hunter, Austrogomphus arbustorum (Northern Queensland)
• Inland hunter, Austrogomphus australis (Victoria, south-eastern South Australia, southern Queensland, north-eastern and western New South Wales)
• Western inland hunter, Austrogomphus collaris (South-western Western Australia)
• Unicorn hunter, Austrogomphus cornutus (Victoria, eastern New South Wales, mid- and southern Queensland)
• Northern river hunter, Austrogomphus doddi (North-eastern Queensland)
• Yellow-striped hunter, Austrogomphus guerini (South-eastern Queensland, eastern New South Wales, Victoria, south-eastern South Australia, Tasmania)
• Pimple-headed hunter, Austrogomphus mjobergi (Northern Northern Territory, north-western Western Australia, north-eastern Queensland)
• Kimberly hunter, Austrogomphus mouldsorum (North-western Western Australia)
• Jade hunter, Austrogomphus ochraceus (South-eastern Queensland, eastern New South Wales, southern Victoria)
• Tiny hunter, Austrogomphus pusillus (North-western Western Australia)

subgenus Pleiogomphus

• Pale hunter, Austrogomphus amphiclitus (Eastern New South Wales, eastern and south-western Queensland)
• Dark hunter, Austrogomphus bifurcatus (North-eastern Queensland)
• Fork hunter, Austrogomphus divaricatus (Eastern Queensland)
• Lemon-tipped hunter, Austrogomphus prasinus (North-eastern Queensland)

genus: Hemigomphus

• Black vicetail, Hemigomphus atratus Mid coastal Queensland
• Zebra vicetail, Hemigomphus comitatus Mid and north coastal Queensland
• Wallum vicetail, Hemigomphus cooloola Coastal New South Wales
• Southern vicetail, Hemigomphus gouldii Coastal South Australia, Victoria, New South Wales, southern Queensland
• Stout vicetail, Hemigomphus heteroclytus Coastal South Australia, Victoria, New South Wales, southern Queensland
• Kakadu vicetail, Hemigomphus magela Northern Territory
• Rainforest vicetail, Hemigomphus theischingeri Mid and northern coastal Queensland

genus: Ictinogomphus

• Australian tiger, Ictinogomphus australis
• Pilbara tiger, Ictinogomphus dobsoni
• Cape York tiger, Ictinogomphus paulini

genus: Odontogomphus

• Pinchtail, Odontogomphus donnellyi (Northern coastal Queensland)

genus: Zephyrogomphus

• Lilac hunter, Zephyrogomphus lateralis (South-western Western Australia)
• Rainforest hunter, Zephyrogomphus longipositor (North-eastern Queensland)

Libellulidae

genus: Aethriamanta

• Square-spot basker, Aethriamanta circumsignata (Northern Australia)
• L-spot basker, Aethriamanta nymphaeae (Northern Australia)

genus: Agrionoptera

• Red swampdragon, Agrionoptera insignis allogenes (Northern New South Wales, eastern Queensland, Northern Territory)
• Striped swampdragon, Agrionoptera longitudinalis biserialis (North-eastern Queensland)

genus: Austrothemis

• Swamp flat-tail, Austrothemis nigrescens (Southern Australia)

genus: Brachydiplax

• Palemouth, Brachydiplax denticauda
• Darkmouth, Brachydiplax duivenbodei

genus: Camacinia

• Black knight, Camacinia othello

genus: Crocothemis

• Black-headed skimmer, Crocothemis nigrifrons

genus: Diplacodes

• Wandering percher, Diplacodes bipunctata (Widespread across Australia)
• Scarlet percher, Diplacodes haematodes (Widespread across Australia)
• Black-faced percher, Diplacodes melanopsis (Coastal southern Queensland, coastal New South Wales, Victoria)
• Charcoal-winged percher, Diplacodes nebulosa (Northern Australia)
• Chalky percher, Diplacodes trivialis (Northern Australia)

genus: Huonia

• Forestwatcher, Huonia melvillensis

genus: Hydrobasileus

• Water prince, Hydrobasileus brevistylus

genus: Lathrecista

• Australasian slimwing, Lathrecista asiatica festa

genus: Macrodiplax

• Wandering pennant, Macrodiplax cora

genus: Nannodiplax

• Pygmy percher, Nannodiplax rubra

genus: Nannophlebia

• Elusive archtail, Nannophlebia eludens
• Pilbara archtail, Nannophlebia injibandi
• Top End archtail, Nannophlebia mudginberri
• Common archtail, Nannophlebia risi

genus: Nannophya

• Australian pygmyfly, Nannophya australis
• Eastern pygmyfly, Nannophya dalei
• Artesian pygmyfly, Nannophya fenshami
• Western pygmyfly, Nannophya occidentalis
• Scarlet pygmyfly, Nannophya paulsoni

genus: Neurothemis

• Spotted grasshawk, Neurothemis oligoneura
• Painted grasshawk, Neurothemis stigmatizans

genus: Notolibellula

• Bicoloured skimmer, Notolibellula bicolor

genus: Orthetrum

• Speckled skimmer, Orthetrum balteatum
• Brownwater skimmer, Orthetrum boumiera
• Blue skimmer, Orthetrum caledonicum
• Rosy skimmer, Orthetrum migratum
• Slender skimmer, Orthetrum sabina
• Green skimmer, Orthetrum serapia
• Fiery skimmer, Orthetrum villosovittatum

genus: Pantala

• Wandering glider, Pantala flavescens

genus: Potamarcha

• Swampwatcher, Potamarcha congener

genus: Raphismia

• Spiny-chested percher, Raphismia bispina

genus: Rhodothemis

• Red arrow, Rhodothemis lieftincki

genus: Rhyothemis

• Iridescent flutterer, Rhyothemis braganza
• Graphic flutterer, Rhyothemis graphiptera
• Yellow-striped flutterer, Rhyothemis phyllis
• Sapphire flutterer, Rhyothemis princeps
• Jewel flutterer, Rhyothemis resplendens

genus: Tetrathemis

• Rainforest elf, Tetrathemis irregularis

genus: Tholymis

• Twister, Tholymis tillarga

genus: Tramea

Male common glider - Darwin

• Dune glider, Tramea eurybia
• Common glider, Tramea loewii
• Northern glider, Tramea transmarina
• Narrow-lobed glider, Tramea stenoloba

genus: Urothemis

• Red baron, Urothemis aliena

genus: Zyxomma

• Short-tailed duskdarter, Zyxomma elgneri
• Large duskdarter, Zyxomma multinervorum
• Long-tailed duskdarter, Zyxomma petiolatum

Macromiidae

genus: Macromia

• Australian cruiser, Macromia tillyardi
• Rainforest cruiser, Macromia viridescens

Petaluridae

genus: Petalura

• South-eastern petaltail, Petalura gigantea
• Western petaltail, Petalura hesperia
• Giant petaltail, Petalura ingentissima
• Coastal petaltail, Petalura litorea
• Beautiful petaltail, Petalura pulcherrima

Synthemistidae

genus: Archaeosynthemis

• Twinspot tigertail, Archaeosynthemis leachii
• Western brown tigertail, Archaeosynthemis occidentalis
• Eastern brown tigertail, Archaeosynthemis orientalis
• Spiny tigertail, Archaeosynthemis spiniger

genus: Austrosynthemis

• Turquoise tigertail, Austrosynthemis cyanitincta

genus: Choristhemis

• Yellow-tipped tigertail, Choristhemis flavoterminata
• Delicate tigertail, Choristhemis olivei

genus: Eusynthemis

• Variable tigertail, Eusynthemis aurolineata
• Mount Lewis tigertail, Eusynthemis barbarae
• Small tigertail, Eusynthemis brevistyla
• Cooloola tigertail, Eusynthemis cooloola
• Carnarvon tigertail, Eusynthemis deniseae
• Southern tigertail, Eusynthemis guttata
• Pretty tigertail, Eusynthemis netta
• Black tigertail, Eusynthemis nigra
• Swift tigertail, Eusynthemis rentziana
• Rainforest tigertail, Eusynthemis tenera
• Mountain tigertail, Eusynthemis tillyardi
• Barrington tigertail, Eusynthemis ursa
• Beech tigertail, Eusynthemis ursula
• Golden tigertail, Eusynthemis virgula

genus: Parasynthemis

• Royal tigertail, Parasynthemis regina

genus: Synthemiopsis

• Tasmanian spotwing, Synthemiopsis gomphomacromioides

genus: Synthemis

• Swamp tigertail, Synthemis eustalacta
• Tasmanian swamp tigertail, Synthemis tasmanica

genus: Tonyosynthemis

• Clavicle tigertail, Tonyosynthemis claviculata
• Slender tigertail, Tonyosynthemis ofarrelli

Superfamily Libelluloidea

genus: Apocordulia

• Nighthawk, Apocordulia macrops

genus: Archaeophya

• Horned urfly, Archaeophya adamsi
• Magnificent urfly, Archaeophya magnifica

genus: Austrocordulia

• Sydney hawk, Austrocordulia leonardi
• Eastern hawk, Austrocordulia refracta
• Top End hawk, Austrocordulia territoria

genus: Austrophya

• Summit mystic, Austrophya monteithorum
• Rainforest mystic, Austrophya mystica

genus: Cordulephya

• Common shutwing, Cordulephya pygmaea
• Tropical shutwing, Cordulephya bidens
• Clubbed shutwing, Cordulephya divergens
• Mountain shutwing, Cordulephya montana

genus: Hesperocordulia

• Orange streamcruiser, Hesperocordulia berthoudi

genus: Lathrocordulia

• Queensland swiftwing, Lathrocordulia garrisoni
• Western swiftwing, Lathrocordulia metallica

genus: Micromidia

• Forest mosquitohawk, Micromidia atrifrons
• Early mosquitohawk, Micromidia convergens
• Thursday Island mosquitohawk, Micromidia rodericki

genus: Pseudocordulia

• Circle-tipped mistfly, Pseudocordulia circularis
• Ellipse-tipped mistfly, Pseudocordulia elliptica

References

• "Order ODONATA". Australian Faunal Directory. Australian Biological Resources Study. 2022. Retrieved 1 September 2024.
• "Theischinger, Günther; Hawking, John (2021). The Complete Field Guide to Dragonflies of Australia (2nd ed.). Melbourne, Australia: CSIRO Publishing. p. 406. ISBN 9781486313747.
• Watson, J.A.L.; Theischinger, G.; Abbey, H.M. (1991). The Australian Dragonflies: A Guide to the Identification, Distributions and Habitats of Australian Odonata. Melbourne: CSIRO. ISBN 0643051368.
• Theischinger, Gunther; Endersby, Ian (2009). Identification Guide to the Australian Odonata (PDF). Department of Environment, Climate Change and Water NSW. p. 241. ISBN 978 1 74232 475 3.
• Paulson, D.; Schorr, M.; Abbott, J.; Bota-Sierra, C.; Deliry, C.; Dijkstra, K.-D.; Lozano, F. (2024). "World Odonata List". OdonataCentral, University of Alabama.