The vestibular system detects head motion and position through the vestibular apparatus in the inner ear. It sends signals to the vestibular nuclei which help maintain the position of the head. The semicircular canals detect rotation of the head while the utricle and saccule detect linear acceleration and orientation. When the head rotates, the endolymph in the canals moves to stimulate hair cells and trigger eye movements that compensate to maintain vision. Nystagmus occurs when this stimulation is unequal as rotation starts or stops.
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Vestibular Apparatus-2
The vestibular system detects head motion and position through the vestibular apparatus in the inner ear. It sends signals to the vestibular nuclei which help maintain the position of the head. The semicircular canals detect rotation of the head while the utricle and saccule detect linear acceleration and orientation. When the head rotates, the endolymph in the canals moves to stimulate hair cells and trigger eye movements that compensate to maintain vision. Nystagmus occurs when this stimulation is unequal as rotation starts or stops.
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vestibular system
The vestibular system can be divided into
the vestibular apparatus and central vestibular nuclei. The vestibular apparatus within the inner ear detects head motion and position and transduces this information to a neural signal. The vestibular nuclei are primarily concerned with maintaining the position of the head in space. The tracts that descend from these nuclei mediate head-on- neck and head-on-body adjustments. VESTIBULAR APPARATUS BONY AND MEMBRANOUS LABYRINTH • Bony labyrinth is the bony tubules situated in the petrous part of temporal bone • Membranous labyrinth are tubes lying within the bony labyrinth and constitutes the functional part of vestibular apparatus • ML is made up of • 1. three semicircular canals • 2. utricle and • 3. saccule • Bony canals are filled with perilymph • The membranous semicircular ducts are filled with endolymph Semicircular ducts • 3 in number (anterior, posterior and lateral) • Arranged in complimentary pairs bilaterally • Dilated part at one end is called the ampulla • Ampulla contains a crest called cristae ampularis • On top of this is a loose gelatinous tissue called Cupula contains hairs cells • SCD and ampulla are filled with endolymph Synergic pairs (mirror images) • L anterior with R anterior • L posterior with R posterior • L lateral with R lateral Position of SCD when the head is bent -the chin touching the sternum • The lateral SCD achieves the horizontal position • The ant project forward and 45o outward • The post project backward and 45o outwards Excitation of hair cells Macula of saccule and utricle • Macula is the sensory organ which detects the position of the head • Macula of utricle lies in horizontal plane and detects the orientation of head in upright posture • Macula of saccule lies in vertical position and detects the orientation of head in lying position Macula and statoconia • macula is covered by a gelatinous layer in which statoconia (calcium carbonate crystals) are present • Macula also contains hair cells • Cilia project from these hair cells which embed into the gelatinous layer • The base is connected to branches of vestibular nerve FUNCTIONS of SCD • When the head is bent and the body rotated to right by a machine around a horizontal axis • There will be bending of cupula in the right lateral duct towards the utricle producing excitation • Opposite effect on left SCD and simultaneous inhibition Detection of head rotation by SCD • SCD detects that the persons head is beginning or stopping to rotate • Example is rapid running in forward direction and then suddenly turns to one side • ????????????????????????????? • Fall • How is he prevented from falling Corrections • SCD informs the brain to make corrections • Cerebellum through vestibulocerebellum makes this possible Detection of linear acceleration by utricle and saccule maculae • When body suddenly pushed forwards • Tendency to fall backwards • Automatic response causes forward bending of the body • This is due to the changes in the statoconia in the macula Excitation of hair cells VESTIBULAR APPARATUS • while the right horizontal canal gets stimulated during head rotations to the right • the left horizontal canal gets stimulated (and thus predominantly signals) by head rotations to the left. When the rotation is stopped • What will be the result????????? • Reverse changes occur • How ???????? • After a while • The Endolymph continues to move along the direction of movement of the head due to inertia • Rotation is stopped • right cupula ----inhibited • Left cupula ------stimulated Nystagmus • This unequal stimulation of cupula is responsible for the production of nystagmus • The characteristic jerky movement of the eye observed at the start and end of a period of rotation is called nystagmus vestibulo-ocular reflex, VOR • reflex that maintains visual fixation on stationary points while the body rotates, • When rotation starts, the eyes move slowly in a direction opposite to the direction of rotation • When the limit of this movement is reached, the eyes quickly snap back to a new fixation point and then again move slowly in the other direction • Slow component • Fast component • The slow component is initiated by impulses from the labyrinths; • the quick component is triggered by a center in the brainstem. • Clinically, nystagmus is seen at rest in patients with lesions of the brainstem • By convention, the direction of eye movement in nystagmus is identified by the direction of the quick component • direction of the quick component is in the direction of rotation