Mechanism of Hearing
Mechanism of Hearing
Mechanism of Hearing
Ear
Consists of 3 parts
External ear
Transmits airborne sound waves to fluid-filled inner ear Amplifies sound energy
Middle ear
Transmits airborne sound waves to fluid-filled inner ear Amplifies sound energy
Inner ear
Houses 2 different sensory systems
Cochlea Contains receptors for conversion of sound waves into nerve impulses which makes hearing possible Vestibular apparatus Necessary for sense of equilibrium
Hearing
Pitch (tone) of sound
Depends on frequency of air waves
Intensity (loudness)
Depends on amplitude of air waves
Timbre (quality)
Determined by overtones
Hearing
Neural perception of sound energy Involves 2 aspects
Identification of the sounds (what) Localization of the sounds (where)
Sound waves
Traveling vibrations of air Consist of alternate regions of compression and rarefaction of air molecules
Equilibrium
Neural signals generated in response to mechanical deformation of hair cells by specific movement of fluid and related structures
Vestibular input goes to vestibular nuclei in brain stem and to cerebellum for use in maintaining balance and posture, controlling eye movement, perceiving motion and orientation
Interesting facts
The stapes or stirrup is the smallest bone in our body.
It is roughly the size of a grain of rice ~2.5mm
Inner ear reaches its full adult size when the fetus is 20-22 weeks old. The ears are responsible for keeping the body in balance Hearing loss is the number one disability in the world.
76.3% of people loose their hearing at age 19 and over
Specifications
Frequency range: 20Hz-20kHz Dynamic range: 0-130 dB Size of cochlea: smaller than a dime
A
N A T O M Y
Pinna /Auricle
Outer ear
Focuses sound waves (variations in pressure) into the ear canal
Pinna size: Inverse Square Law Larger pinna captures more of the wave Elephants: hear low frequency sound from up to 5 miles away Human Pinna structure: Pointed forward & has a number of curves Helps in sound localization More sensitive to sounds in front
Auditory Canal
Dogs/ Cats- Movable Pinna => focus on sounds from a particular direction
Pinna /Auricle
Outer ear
Auditory Canal
Horizontal localization
Sound Localization Vertical localization Is sound on your right or left side?
Interaural differences
- The signal needs to travel further to more distant ear - More distant ear partially occluded by the head
Left ear
Left ear
Left ear
intensity difference
Left ear
Thresholds
Interaural time differences (ITDs) Threshold ITD 10-20 ms (~ 0.7 cm) Interaural intensity differences (IIDs) Threshold IID 1 dB
Interaural time differences (ITDs) Low frequencies Up to around 1500 Hz; sensitivity declines rapidly above 1000 Hz
D U P L E X
H
E O
R
Y
Pinna /Auricle
Outer ear
Sound Localization
Auditory Canal
Horizontal localization
Vertical localization Is sound above or below? Pinna Directional Filtering
Pinna amplifies sound above and below differently
Curves in structure selective amplifies certain parts of the sound spectrum
Pinna /Auricle
Outer ear
Closed tube resonance: wave resonator Auditory canal length 2.7cm Resonance frequency ~3Khz Boosts energy between 2-5Khz upto 15dB
Auditory Canal
A
N A T O M Y
Eardrum
Middle Ear
Eardrum OssiclesOval Window Impedance matching Ossicles: Malleus, Incus, Stapes
Ossicles
Oval window
Amplification
By lever action < 3x Area amplification [55mm2 3.2mm2] 15x
Stapedius reflex
Protection against low frequency loud sounds Tenses muscles stiffens vibration of Ossicles Reduces sound transmitted (20dB)
A
N A T O M Y
Inner Ear
Body's balance organs Accelerometers in 3 perpendicular planes Hair cells detect fluid movements Connected to the auditory nerve
Inner Ear
Cochlea is a snail-shell like structure 2.5 turns 3 fluid-filled parts: Scala tympani Scala Vestibuli Cochlear duct (Organ of Corti)
(1) Organ of Corti (2) Scala tympani (3) Scala vestibulli (4) Spiral ganglion (5) auditory nerve fibres
Inner Ear
Organ of Corti Basilar membrane Inner hair cells and outer hair cells (16,000 -20,000) IHC:100 tiny stereocilia
The body's microphone: Vibrations of the oval window causes the cochlear fluid to vibrate Basilar membrane vibration produces a traveling wave Bending of the IHC cilia produces action potentials The outer hair cells amplify vibrations of the basilar membrane
Place Theory
4mm2 1mm2
32-35 mm long
Each position along the BM has a characteristic frequency for maximum vibration Frequency of vibration depends on the place along the BM At the base, the BM is stiff and thin (more responsive to high Hz) At the apex, the BM is wide and floppy (more responsive to low Hz)
Auditory Neuron
Carries impulses from both the cochlea and the semicircular canals Connections with both auditory areas of the brain Neurons encode
Steady state sounds Onsets or rapidly changing frequencies
Elephants
Sound Production
A a typical male elephants rumble is around an average minimum of 12 Hz, a female's rumble around 13 Hz and a calf's around 22 Hz. Produce sounds ranging over more than 10 octaves, from 5 Hz to over 9,000 Hz Produce very gentle, soft sounds as well as extremely powerful sounds. (112dB recorded a meter away)
Hearing Wider tympanic membranes Longer ear canals (20 cm) Spacious middle ears.
ear.
The fluid-filled semicircular canals play a role in balance, as hairs
Ear Bone: Middle Ear Figure 2: The auditory ossicles of the middle ear and the structures surrounding them.
Bony Labyrinth The two labyrinths of the inner ear. The bony labyrinth is partially cut away to show the membranous labyrinth within.
A cross section through one of the turns of the cochlea (inset) showing the scala tympani and scala vestibuli, which contain perilymph, and the cochlear duct, which is filled with endolymph.
Mechanism Of Hearing: 1. Sound waves enter the outer ear and travel through the external auditory canal until they reach the tympanic membrane, causing the membrane and the attached chain of auditory ossicles to vibrate.
(A) The fibres of the basilar membrane become progressively wider and more flexible from the base of the cochlea to the apex. As a result, each area of the basilar membrane vibrates preferentially to a particular sound frequency. (B) High-frequency sound waves cause maximum vibration of the area of the basilar membrane nearest to the base of the cochlea, (C) medium-frequency waves affect the centre of the membrane, (D) and low-frequency waves preferentially stimulate the apex of the basilar membrane. (The locations of cochlear frequencies along the basilar membrane shown are a composite drawn from different sources.)
The absolute threshold of hearing (ATH) is the minimum sound level of a pure tone that an average ear with normal hearing can hear in a noiseless environment. Minimum audibility curve is a standardised graph of the threshold of hearing versus frequency for an average human, and is used as the reference level when measuring hearing loss with an audiometer as shown on an audiogram An equal-loudness contour is a measure of sound pressure (dB SPL), over the frequency spectrum, for which a listener perceives a constant loudness when presented with pure steady tones. The unit of measurement for loudness levels is the phon, and is arrived at by reference to equal-loudness contours
An audiogram is a standard way of representing a person's hearing loss[1]. Most audiograms cover the limited range 100Hz to 8000Hz (8kHz) which is most important for clear understanding of speech, and they plot the threshold of hearing relative to a standardised curve that represents 'normal' hearing, in dBHL. Audiograms are set out with frequency in hertz (Hz) on the horizontal axis, most commonly on a logarithmic scale, and a linear dBHL scale on the vertical axis. Normal hearing is classified as being between -10dBHL and 15dBHL, although 0dB from 250Hz to 8kHz is deemed to be 'average' normal hearing.
Audiogram
MEASUREMENT Audiograms are produced using a piece of test equipment called an audiometer, and this allows different frequencies to be presented to the subject, usually over calibrated headphones, at any specified level. The levels are, however, not absolute, but weighted with frequency relative to a standard graph known as the minimum audibility curve which is intended to represent a 'normal' hearing. This is not the best threshold found for all subjects, under ideal test conditions, which is represented by around 0 Phon or the threshold of hearing on the equal-loudness contours.
Conductive hearing loss happens when there is a problem conducting sound waves through the outer ear, tympanic membrane (eardrum) or middle ear (ossicles) and the inner ear (oval window/ round window). This type of hearing loss may occur in conjunction with sensorineural hearing loss or alone. Sensorineural hearing loss is a type of hearing loss in which the root cause lies in the vestibulocochlear nerve (Cranial nerve VIII), the inner ear, or central processing centers of the brain.
During air conduction testing, small foam insert earphones are placed in your ear
canals. In some cases, headphones may be used instead. A series of tones are presented at various frequencies. You are required to respond by pressing a button or raising your hand whenever you hear a tone; even if it is very soft. Your results indicate to us the softest level at which you can hear a tone for each of ten different frequencies that make up the speech spectrum.
Bone conduction testing requires us to place a small oscillator attached to a headband behind your ear. Again you are asked to respond to the softest tone that you hear at various frequencies. By reviewing the results of the bone conduction testing, compared to the air conduction testing, the audiologist can determine if your hearing loss is the result of a problem in the outer, middle or inner ear.
Speech audiometry Subjective test in which subject repeats a standard list of words given through headphones at various loudness. Very useful in assessing need for hearing aid provision. Word recognition tests (also known as speech discrimination tests): involves reading a list of words to see if patients can discriminate words. Inferences can be made about central processing and central hearing deficits. Tympanometry Tympanometry is a measure of the stiffness of the eardrum and thus evaluates middle ear function
HEARING AID
A hearing aid is an electroacoustic body worn apparatus which typically fits in or behind the wearer's ear, and is designed to amplify and modulate sounds for the wearer. A hearing aid is an electronic, battery-operated device that amplifies and changes sound to allow for improved communication. Earlier devices, known as an "ear trumpet" or "ear horn", were passive funnel-like amplification cones designed to gather sound energy and direct it into the ear canal. Similar devices include the bone anchored hearing aid, and cochlear implant.
Receiver: Translates those electrical impulses into louder sounds. (delivers amplified sound into ear-miniature loudspeaker)
Battery: Serves as power source for device. Some hearing aids also have earmolds (earpieces) to direct the flow of sound into the ear and enhance sound quality.
Digital hearing aids use a microphone, receiver, battery, and computer chip. They transform the sound, convert it into bits, and manipulate it before amplifying the signal. A digital hearing aid can be programmed. This means that digital hearing aids can be individually adjusted to suit a specific user.
Behind-the-Ear (BTE)
BTE hearing aids are worn behind the ear and are connected to a plastic earmold that fits inside the outer ear. The components are held in a case behind the ear. Sound travels from the aid through the earmold into the ear. BTE aids are used by people of all ages for mild to profound hearing loss. Poorly fitting BTE earmolds may cause feedback, a whistle sound caused by the fit of the hearing aid or by buildup of earwax or fluid.
In-the-Ear (ITE)
ITE hearing aids fit completely in the outer ear and are used for mild to severe hearing loss. The case, which holds the components, is made of hard plastic. ITE aids can accommodate added technical mechanisms such as a telecoil, a small magnetic coil contained in the hearing aid that improves sound transmission during telephone calls. ITE aids can be damaged by earwax and ear drainage, and their small size can cause adjustment problems and feedback. They are not usually worn by children because the casings need to be replaced as the ear grows.
In-the-Canal (ITC)
ITC hearing aids are customized to fit the size and shape of the ear canal and are used for mild or moderately severe hearing loss. Small one-piece hearing aid All components contained in a custom-fit, hard-molded plastic shell Fits outside the ear canal Slightly visible in the ear
Completely-in-Canal (CIC)
CIC hearing aids are mostly concealed in the ear canal and are used for mild to moderately severe hearing loss. Not suitable for people with severe hearing losses. Because of their small size, canal aids may be difficult for the user to adjust and remove, and may not be able to hold additional devices, such as a telecoil. Canal aids can also be damaged by earwax and ear drainage. They are not recommended for children. All components contained in a custom-fit, hard-molded plastic shell
Cochlear implant
A cochlear implant (CI) is a surgically implanted electronic device that provides a sense of sound to a person who is profoundly deaf or severely hard of hearing. The cochlear implant is often referred to as a bionic ear. Unlike hearing aids, the cochlear implant does not amplify sound, but works by directly stimulating any functioning auditory nerves inside the cochlea with an electric field. External components of the cochlear implant include a microphone, speech processor and an RF transducer or primary headpiece coil.
Cochlear Implants
COCHLEAR IMPLANT
The headpiece coil has a magnet by which it attaches to another magnet placed on the secondary coil often beside the cochlear implant. The implant relays the incoming signal to the implanted electrodes in the cochlea. The speech processor allows an individual to adjust the sensitivity of the device. The implant gives recipients additional auditory information, which may include sound discrimination fine enough to understand speech in quiet environments. Post-implantation rehabilitative therapy is often critical to ensuring successful outcomes.
a microphone which picks up sound from the environment a speech processor which selectively filters sound to prioritize audible speech and sends the electrical sound signals through a thin cable to the transmitter, a transmitter, which is a coil held in position by a magnet placed behind the external ear, and transmits the processed sound signals to the internal device by electromagnetic induction,
INTERNAL:
a receiver and stimulator secured in bone beneath the skin, which converts the signals into electric impulses and sends them through an internal cable to electrodes,
an array of up to 22 electrodes wound through the cochlea, which send the impulses to the nerves in the scala tympani and then directly to the brain through the auditory nerve system. There are 4 manufacturers for Cochlear implants, and each one produces a different implant with different number of electrodes. Cochlear produces implants with 22 electrodes, Advanced Bionics produces implants with 16 electrodes and the use a technique called current stearing in which two electrodes are stimulated simulatenously with different current level to produce intermediate virtual channels. The number of channels is not a primary factor upon which an manufacturer is chosen, but the signal processing
The device is surgically implanted under a general anaesthetic, and the operation usually
takes from 1 to 5 hours. First a small area of the scalp directly behind the ear is shaven and cleaned. Then a small incision is made in the skin just behind the ear and the surgeon drills into the mastoid bone and the inner ear where the electrode array is inserted into the cochlea. The patient normally goes home the same day as the surgery, although some cochlear implant recipients stay in the hospital for 1 to 2 days. It is considered outpatient surgery. As with every medical procedure, the surgery involves a certain amount of risk; in
this case, the risks include skin infection, onset of tinnitus, damage to the vestibular
system, and damage to facial nerves that can cause muscle weakness, impaired facial sensation, or, in the worst cases, disfiguring facial paralysis.
There is also the risk of device failure, usually where the incision does not heal properly. This occurs in 2% of cases and
Cochlear Implants
Definition:
A device that electrically stimulates the auditory nerve of patients with severe-toprofound hearing loss to provide them with sound and speech information The first fully functional Brain Machine Interface (BMI)
Who is a candidate?
Severe-to profound sensorineural hearing loss Hearing loss did not reach severe-to-profound level until after acquiring oral speech and language skills Limited benefit from hearing aids
CI characteristics
1. Electrode design
2. Type of stimulation
3. Transmission link
4. Signal processing
Waveform representation or feature extraction