Unit 2
Unit 2
• Essential oils: These are volatile oils produced by special glands and
cells. Aromatic flowers, leaves and bark are due to essential oils.
• Alkaloids: These are nitrogenous compounds, made up of carbon,
hydrogen, oxygen and nitrogen. They are found in storage organs
of plants such as seeds, bark and leaves. They are insoluble in
water but soluble in alcohol. They have sour taste and some are
poisonous. However, a large number of alkaloids, such as quinine,
reserpine, nicotine, caffeine, strychnine, morphine, atropine, are
used as medicines.
• Resins: These are produced by the oxidation of essential oils. These
are found in some special glands either alone or in combination
with essential oils. These are insoluble in water but soluble in ether
and alcohol. These are used in the manufacture of paints and
varnishes.
• Gums: Produced by the disintegration of cellulose cell wall.
They are soluble in water. Used for sticking purposes, and
also as medicine,
• Tannins: They are sour in taste and related to glycosides.
They occur in vacuolar sap, cell wall, bark and leaves of some
plants. They are found mostly in unripe fruits. They are used
on a large scale for hardening of leather, a process called
tanning of leather.
• Latex: It is a milky substance secreted by latex glands. Robber
secreted by the rubber tree Hevea brasiliensis is an important
example.
• Nectar: Nectar is secreted by nectaries in plants that attracts
insects for pollination because it is sweet and contains
sucrose, glucose and fructose.
3. Minerals:
Dermal Tissue
•Epidermis
•Periderm
•Guard cell
Meristematic Tissues
• Tissues where cells are constantly dividing are called
meristems or meristematic tissues.
• They are the mass of young and undifferentiated cells having
the power to divide.
• These regions produce new cells. These new cells are
generally small, isodiametric structures with a number of tiny
vacuoles and a large nucleus, by comparison. As the cells
mature the vacuoles will grow to many different shapes and
sizes, depending on the needs of the cell.
• There are three types of meristems:
• Apical Meristems
• Lateral Meristems
• Intercalary Meristems
• Apical meristems are located at or near the tips of roots and shoots. As
new cells form in the meristems, the roots and shoots will increase in
length. This vertical growth is also known as primary growth. A good
example would be the growth of a tree in height. Each apical meristem
will produce embryo leaves and buds as well as three types of primary
meristems: protoderm, ground meristems, and procambium. These
primary meristems will produce the cells that will form the primary
tissues.
• Intercalary meristems are found in grasses and related plants that do not
have a vascular cambium or a cork cambium, as they do not increase in
girth. These plants do have apical meristems and in areas of leaf
attachment, called nodes, they have the third type of meristematic
tissue. This meristem will also actively produce new cells and is
responsibly for increases in length. The intercalary meristem is
responsible for the regrowth of cut grass.
• Lateral meristems account for secondary growth in plants. Secondary
growth is generally horizontal growth. A good example would be the
growth of a tree trunk in girth. There are two types of lateral meristems
in the plants; vascular cambium and the cork cambium.
Permanent Tissue
Simple Permanent Tissue
• Parenchyma Tissue: Parenchyma cells form parenchyma tissue. Parenchyma cells are
the most abundant of cell types and are found in almost all major parts of higher
plants. These cells are basically sphere shaped when they are first made. However,
these cells have thin walls, which flatten at the points of contact when many cells are
packed together. These cells have large vacuoles and may contain various secretions
including starch, oils, tannins, and crystals. Parenchyma cells can divide if they are
mature, and this is vital in repairing damage to plant tissues. Parenchyma cells and
tissues comprise most of the edible portions of fruit.
• Some parenchyma cells have many chloroplasts and form the tissues found in leaves.
This type of tissue is called chlorenchyma. The chief function of this type of tissue is
photosynthesis, while parenchyma tissues without chloroplasts are generally used for
food or water storage.
• Additionally, some groups of cells are loosely packed together with connected air
spaces, such as in water lilies, this tissue is called aerenchyma tissue. These type of
cells can also develop irregular extensions of the inner wall which increases overall
surface area of the plasma membrane and facilitates transferring of dissolved
substances between adjacent cells.
• Functions: Synthesis, Storage, Conduction of water and food, photosynthesis
(chlorenchyma), buoyancy (aerenchyma)
• Sclerenchyma Tissue: Sclerenchyma cells form sclerenchyma tissue. These cells have
thick, tough secondary walls that are imbedded with lignin. At maturity, most
sclerenchyma cells are dead and function in structure and support. Sclerenchyma cells
can occur in two forms:
• Sclereids are sclerenchyma cells that are randomly distributed throughout other
tissues. Sometimes they are grouped within other tissues in specific zones or regions.
They are generally as long as they are wide. An example, would be the gritty texture in
some types of pears. The grittiness is due to groups of sclereid cells. Sclereids are
sometimes called stone cells.
• Fibers are found in a wide variety of tissues in roots, stems, leaves and fruits. Usually
fiber cells are much longer than they are wide and have a very tiny cavity in the center
of the cell. Currently, fibers from over 40 different plant families are used in the
manufacture of textiles, ropes, string and canvas goods to name a few.
• Collenchyma Tissue: Collenchyma cells form collenchyma tissue. They are mechanical
tissue with cell wall thickened by cellulose and pectin. These cells have a living
protoplasm, like parenchyma cells, and may also stay alive for a long period of time.
Their main distinguishing difference from parenchyma cells is the increased thickness
of their walls.
• Collenchyma cells are found just beneath the epidermis and generally they are
elongated and their walls are pliable and strong. As a plant grows these cells and the
tissues they form, provide flexible support for organs such as leaves and flower parts.
Good examples of collenchyma plant cells are the ‘strings’ from celery that get stuck in
our teeth.
Complex Permanent Tissue
• Xylem: Xylem is permanent tissue that conducts water along with minerals
from root to leaf. They are composed of tracheids, vessels, xylem fibers
and xylem parenchyma.
• Tracheids are the elongated, spindle shaped, thick lignified walled cells
which are dead at maturity.
• Vessels are multicellular elongated tubes formed by chain of elongated
cells.
• Pits are the areas of primary wall of tracheids and vessels through which
diffusion of fluid takes place from cell to cell.
• Xylem fibres are the sclerenchyma fibres associated with xylem.
• Xylem parenchyma are the parenchymatous cell of xylem that are living
elements in xylem.
• Phloem: Phloem is permanent tissue that conducts dissolved food from
leaves to storage organs and to growing regions. It occurs along with
xylem. This conduction system is composed of sieve elements, companion
cells, phloem fibres and phloem parenchyma.
• Sieve elements are composed of sieve cells and sieve tubes. Sieve cells
are living, elongated and slender that help for conduction. Sieve tubes are
combined to form sieve plate that consists of numerous perforating.
• Sieve-tube members that are alive contain a polymer called callose.
Callose stays in solution as long at the cell contents are under pressure. As
a repair mechanism, if an insect injures a cell and the pressure drops, the
callose will precipitate. However, the callose and a phloem protein will be
moved through the nearest sieve plate where they will form a plug.
• This prevents further leakage of sieve tube contents and the injury is not
necessarily fatal to overall plant turgor pressure.
• Companion cells are living, have dense granular cytoplasm and a
prominent nucleus.
• Phloem fibres are the sclerenchyma associated with phloem that are
lignified and having small and rounded simple pits.
• Phloem Parenchyma are the parenchymatous cell of phloem present in
dicots.
D. Microscopy and morphology of a plant
1. Leaves
2. Flower
3. Fruit
4. Seed
5. Stem
6. Bark
7. Root
8. Rhizome
1. Leaves
• Leaves are flat, thin, green appendages to the stem
that have important role in support and functions of
plant.
• Leaf includes leaf and leaflets.
• Medicinal leaves are collected during flowering season
of the plants, when plants reach maturity and they are
photosynthetically most active.
• Leaves containing volatile oils are collected when the
plant is rich in volatile oils.
• The weather and time of collection is important in
procurement of the leaves.
• Discolouration of leaves is considered as substandard.
Characterstics of leaf
1. Stomata: Stomata are minute epidermal openings in the
leaves of plant for gaseous exchange and transpiration.
Stomata consists of two kidney shaped cells and a minute
opening in between.
According to type and arrangement of cells the stomata are of 4
types:
a. Moss type
b. Gymnospermous type
c. Gramineous type
d. Dicotyledonous type
• Dicotyledonous stomata are further classified as:
• Paracytic or parallel celled stomata: It comprises of two guard
cells covered by two subsidiary cells, the long axes of which are
parallel to that of stoma.
• Diacytic or cross celled stomata: The guard cells are covered by
two subsidiary cells, the arrangement of subsidiary cells on guard
cells is at right angle to that of stoma.
• Anisocytic or unequal celled stomata: Two guard cells are
covered by three subsidiary cells of unequal size.
• Anomocytic or irregular celled stomata: Stomata is sorrounded
by varying number of subsidiary cells.
• Actinocytic or radiate celled stomata: Two guard cells are
sorrounded by a circle of radiating subsidiary cells.
Stomatal number: The average number of
stomata per square mm of the epidermis is
known as stomatal number.
Stomatal Index: The percentage proportion of
the number of stomata to the number of
epidermal cells of a leaf is known as stomatal
index.
Stomatal Index = S × 100
E+S
S= No. of stomata per unit area
E= No. of epidermal cells in the same area
2. Leaf constants: