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TUNDRA
TUNDRA
Peter D. Moore
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Bang FOF 10 9 8 7 6 5 4 3 2 1
Preface xi
Acknowledgments xiii
Introduction xv
What Is Tundra? xv
Where Is the Tundra? xvi
Why Is the Tundra Important? xviii
3 F Types of Tundra 64
Tundra Forms 64
Vegetation Structure 64
Polar Desert 65
Dwarf Shrub Tundra 66
Tall Shrub Tundra 71
Forest Tundra 71
Coastal Tundra 74
Tundra Wetlands 74
Antarctic Tundra 78
Alpine Tundra 79
Tropical Alpine Habitats 82
Conclusions 85
Increasingly, scientists, environmentalists, engineers, and Forests, Revised Edition; and Wetlands, Revised Edition—
land-use planners are coming to understand the living between them span the breadth of land-based and aquatic
planet in a more interdisciplinary way. The boundaries ecosystems on Earth. Each volume considers a specific
between traditional disciplines have become blurred as global ecosystem from many viewpoints: geographical, geo-
ideas, methods, and findings from one discipline inform logical, climatic, biological, historical, and economic. Such
and influence those in another. This cross-fertilization is broad coverage is vital if people are to move closer to under-
vital if professionals are going to evaluate and tackle the standing how the various ecosystems came to be, how they
environmental challenges the world faces at the beginning are changing, and, if they are being modified in ways that
of the 21st century. seem detrimental to humankind and the wider world, what
There is also a need for the new generation of adults, might be done about it.
currently students in high schools and colleges, to appreci- Many factors are responsible for the creation of Earth’s
ate the interconnections between human actions and envi- living mosaic. Climate varies greatly between Tropics and
ronmental responses if they are going to make informed poles, depending on the input of solar energy and the move-
decisions later, whether as concerned citizens or as inter- ments of atmospheric air masses and ocean currents. The
ested professionals. Providing this balanced interdisciplin- general trend of climate from equator to poles has resulted
ary overview—for students and for general readers as well in a zoned pattern of vegetation types, together with their
as professionals requiring an introduction to Earth’s major associated animals. Climate is also strongly affected by
environments—is the main aim of the Ecosystem set of the interaction between oceans and landmasses, resulting
volumes. in ecosystem patterns from east to west across continents.
The Earth is a patchwork of environments. The equato- During the course of geological time even the distribution
rial regions have warm seas with rich assemblages of cor- of the continents has altered, so the patterns of life currently
als and marine life, while the land is covered by tall forests, found on Earth are the outcome of dynamic processes and
humid and fecund, and containing perhaps half of all Earth’s constant change. The Ecosystem set examines the great eco-
living species. Beyond are the dry tropical woodlands and systems of the world as they have developed during this long
grassland, and then the deserts, where plants and animals history of climatic change, continental wandering, and the
face the rigors of heat and drought. The grasslands and for- recent meteoric growth of human populations.
ests of the temperate zone grow because of the increasing Each of the great global ecosystems has its own story to
moisture in these higher latitudes, but grade into coniferous tell: its characteristic geographical distribution; its pattern
forests and eventually scrub tundra as the colder conditions of energy flow and nutrient cycling; its distinctive soils or
of the polar regions become increasingly severe. The com- bottom sediments, vegetation cover, and animal inhabit-
plexity of diverse landscapes and seascapes can, neverthe- ants; and its own history of interaction with humanity. The
less, be simplified by considering them as the great global books in the Ecosystem set are structured so that the differ-
ecosystems that make up our patchwork planet. Each global ent global ecosystems can be analyzed and compared, and
ecosystem, or biome, is an assemblage of plants, animals, the relevant information relating to any specific topic can be
and microbes adapted to the prevailing climate and the quickly located and extracted.
associated physical, chemical, and biological conditions. The study of global ecosystems involves an examina-
The six volumes in the set—Deserts, Revised Edition; tion of the conditions that support the planet’s diversity.
Tundra; Oceans, Revised Edition; Tropical Forests; Temperate But environmental conditions are currently changing rap-
n xi n
xii F tundra
idly. Human beings have eroded many of the great global of the residue of Earth’s biodiversity. The starting point in
ecosystems as they have reclaimed land for agriculture and approaching these problems is to understand how the great
urban settlement and built roads that cut ecosystems into ecosystems of the world function, and how the species of
ever smaller units. The fragmentation of Earth’s ecosystems animals and plants within them interact to form stable and
is proving to be a serious problem, especially during times productive assemblages. If these great natural systems are to
of rapid climate change, itself the outcome of intensive survive, then humanity needs to develop greater respect and
industrial activities on the surface of the planet. The next concern for them, and this can best be achieved by under-
generation of ecologists will have to deal with the control standing better the remarkable properties of our patchwork
of global climate and also the conservation and protection planet. Such is the aim of the Ecosystem set.
Acknowledgments
I should like to record my gratitude to the editorial staff during the writing of this book, together with much needed
at Facts On File for their untiring support, assistance, and critical appraisal. I must also acknowledge the contribu-
encouragement during the preparation of this book. Frank tion of many generations of students in the Life Sciences
K. Darmstadt, executive editor, has been a constant source of Department of the University of London, King’s College,
advice and information and has been meticulous in check- who have been a constant source of stimulation, critical
ing the text and coordinating the final assembling of materi- comment, and new ideas. I also acknowledge a consider-
als. My gratitude also extends to the production department. able debt to my colleagues in teaching and research at King’s
I should also like to thank Richard Garratt for his excellent College, especially those who have accompanied me on field
illustrations and Elizabeth Oakes for her perceptive selection courses and research visits to many parts of the world. Their
of photographs. Particular thanks are due to my wife, who work, together with that of countless other dedicated ecolo-
has displayed a remarkable degree of patience and support gists, underlies the science presented in this book.
n xiii n
Introduction
The world is no longer rich in wilderness. In the course in the depths of winter when mountaintops offer recre-
of a few thousand years the human species has penetrated ational opportunities, such as skiing and snowboarding.
to all parts of the globe and has everywhere left an indel- Consequently, the tundra is a mystery to the vast majority
ible mark in the form of habitat destruction and pollution. of people, a land of ice and snow that is known only through
But of all the world’s biomes the tundra remains the closest television documentaries or adventure novels. It is not sur-
to its original, pristine condition. Tundra may be regarded prising, therefore, that there are many misconceptions about
as the last true wilderness. This book examines the struc- the tundra.
ture and the distribution of tundra ecosystems, including The tundra is cold. This is true as far as average annual
both the polar regions and the high mountain tops of the temperature is concerned, and it is certainly true with respect
world. It considers the development of the tundra during to the long, dark winters. But in summer temperatures can
the course of geological history and looks at the impact of become high, and the long summer days become favorable
people on this wild ecosystem. By understanding how the to many plants and animals. The tundra is low in biologi-
tundra has come into being and how it functions, future cal diversity. Again, this is true when compared to tropical
generations will be in a better position to protect its unique forests, but the range of creatures found in the tundra is still
features. considerable, and many of the plants and animals encoun-
Tundra is structured in a way that will make relevant tered there are found nowhere else on Earth. The tundra is
information both interesting and accessible to the student. wet in summer and snowy in winter. This is the immediate
Each chapter deals with a major feature of the tundra envi- impression of the landscape, but the actual quantity of pre-
ronment, including physical aspects, such as climate, geol- cipitation (rain and snow) is actually very low. The input of
ogy, and geography, and biological aspects, including the water from the atmosphere is similar to that of a desert, but
adaptations of organisms to extreme conditions. The stu- the low temperature and consequent low evaporation mean
dent is encouraged to integrate these living and nonliving that water or snow is often present. The tundra is a waste-
components by viewing the tundra as an interactive ecosys- land. From the point of view of human settlement and farm-
tem, operating like a delicately balanced machine. In order ing, this is clearly the case, but there are many inconspicuous
to appreciate the problems facing the tundra, the question of ways in which the tundra serves humanity. The balance of
environmental change is finally emphasized, looking at the gases in the Earth’s atmosphere and therefore global climate
possible consequences of climate change and increasing pol- are strongly affected by what goes on in the tundra. The tun-
lution in this remote and wild biome. Throughout the book dra has many hidden values.
diagrams, photographic illustrations, and explanatory side-
bars are used to ensure that the factual material is presented
with clarity and maximum impact.
The tundra regions of the world have the lowest density n What Is Tundra?
of human populations, with the possible exception of some The word tundra is derived from the Finnish language. The
deserts. It is the one biome that has resisted most attempts rounded, treeless hilltops of northern Finland are called
at agricultural exploitation and human settlement. Most tunturi, and the word has become modified to tundra. The
people, even those who spend much time in traveling the application of the word tundra has also become extended to
world, have never visited tundra habitats except perhaps cover all treeless landscapes found in generally cold places,
n xv n
xvi F tundra
such as the regions bordering upon the polar ice or oceans, season is particularly short, the soil may not thaw out
and the summits of high mountains, even in the Tropics. completely. Only the surface layers thaw, leaving the
The absence of trees in the tundra, which is its most deeper layers of the soil permanently frozen, which is
distinctive feature when first observed, is not confined to called the permafrost. Animal life in the soil is thus con-
this biome. Deserts and grasslands have few trees because fined to the upper layers, where conditions become warm
of drought conditions that restrict plant growth. The scrub enough for active life and growth in summer. Even the
vegetation of the chaparral is maintained free of trees hardy fungi and bacteria in the soil are restricted in
because it is subject to frequent fires. But the restricting their activity, spending the winter in a dormant state
factors for tree growth in the tundra are the combination and becoming active in the summer as they decompose
of low winter temperatures and high winds that carry abra- organic remains.
sive particles of ice, blasting and destroying all growing
parts of plants that raise themselves high above the ground.
All the vegetation found in the tundra hugs the soil closely.
Cushions of moss, flat plates of lichens on rocks, small n Where Is the Tundra?
domes of herbaceous plants, and low-lying stretches of There are three main regions of the Earth that support
dwarf shrubs form the general vegetation cover of the tun- the tundra biome: the Arctic region, together with some
dra. The only trees present are hardly recognizable as such, adjacent areas outside the Arctic Circle that can be termed
being reduced to contorted, dwarf forms that lie close to subarctic; some coastal locations on the continent of
the ground. Antarctica, together with neighboring islands in the south-
Beneath this cover of vegetation lies a sheltered and ern ocean, strictly subantarctic islands; and mountain
protected zone, where pockets of air remain still and unaf- summits throughout the world where conditions at high
fected by the wind above the plant canopy. Temperatures altitude are too severe for tree growth. These regions are
rise during the day, humidity stays high, and these two fac- shown on the map.
tors favor the many insects and other small invertebrates The Arctic and Antarctic tundra regions have much in
that manage to eke out a living, hidden from the bleak tun- common and can jointly be termed polar tundra. The moun-
dra landscape. Small mammals and birds can also find a taintop tundra, on the other hand, has many distinctive cli-
home in the low cover of vegetation, feeding on leaves, ber- matic features and contains some plants and animals that
ries, and insects. Some of the birds travel great distances to are not found in the polar regions. These montane sites can
take advantage of the productivity of the tundra together be classed as alpine tundra.
with the long days of summer in the polar regions. These The two types of tundra share a generally cold climate,
extended days enable them to feed their young more effec- often with only a short summer season that is suitable
tively and produce bigger families, so a surprising number for plant growth and animal activity. They are both sub-
of birds choose to breed in the tundra. Some larger mam- jected to ice and snow in winter, much of which may
mal herbivores, including caribou and musk ox, graze on persist through the summer. Wind is an important factor
the summer productivity. Both birds and mammals in turn in both types of tundra, restricting the height to which
may form a food source for the tundra predators, gyr fal- plants are able to grow. The vegetation of both alpine
con, arctic fox, wolf, and polar bear. Together the plants and polar tundra, therefore, has a superficial similarity.
and animals form the distinctive living community of the Some plant species are found in both types of tundra,
tundra. while others are restricted to either the alpine or the polar
The landscape of the tundra is not flat and uniform. tundra.
There are many microhabitats to be found within the tun- Some mountains are found within the Arctic and
dra, including peaty wetlands in the lower regions and Antarctic Circles, while other mountains are found close
raised ridges of shingle and rock in the more elevated to the equator. The polar mountains provide a home for
locations. Craggy cliffs and high mountains are found in a similar range of plants and animals to those found in
some of the polar tundra regions, providing impressive lowland polar tundra, while the mountains of equato-
vistas with permanent snow and ice. This varied landscape rial regions usually carry a very different assemblage of
offers opportunities for a range of specialized plants and species. Polar and equatorial mountains also differ con-
animals to find a home within the variety of microhabitats siderably in their respective climates, despite the fact
on offer. that both experience a cold overall average temperature.
The soils of the tundra are also distinctive in their Daily fluctuations in temperature are much greater on
form. With the coming of winter they freeze, while in an equatorial mountain, but seasonal variation is far less.
summer they thaw once again. When temperatures in the Precipitation is usually higher on mountains than is the
tundra region are especially low, or when the summer melt case in the lowland polar regions. Mountains are also
Introduction F xvii
Arctic Circle
equator
Antarctic Circle
The regions of the world in which tundra ecosystems are cific demands of their environment. When this is coupled
found. Much of Greenland and most of Antarctica are covered with the spatial isolation of mountains, which stimu-
by ice and so bear no vegetation. Tundra in regions outside lates evolutionary divergence, it is not surprising to find
the Arctic and Antarctic Circles is confined to high-altitude
that there are some very distinctive organisms found on
sites. This type of mountain tundra is called alpine tundra to
distinguish it from polar tundra. mountains, some of them very restricted in their global
distribution.
Tundra, therefore, is a globally widespread and varied
habitat. It is the product of extreme climatic conditions,
often widely separated from one another, and the inter- and it contains some organisms that are highly adapted in
vening lowlands may be impassable for animals with poor order to cope with the stress of extreme cold. In general,
dispersal ability. So the plants and animals of mountains tundra is not a rich biome; it does not contain a very high
may find themselves isolated, and this can lead to evolu- diversity of species, but it is the home of some very special-
tionary separation and development much like that found ized species that are found in no other type of habitat. The
on oceanic islands. structure, biochemistry, and behavior of tundra organisms
Mountain landscapes usually display a greater vari- have long attracted the interest of biologists and ecolo-
ety of topographical features than flatlands. High peaks gists, who are eager to discover how these creatures sur-
are interspersed with deep valleys, providing a wide range vive under such stress. The presence of whole communities
of habitats and a great variety of locations that differ in of these highly specialized creatures has made the tundra
their local climate, or microclimate. This diversity of habi- a living laboratory in which scientists can study the ways
tat and microclimate can lead to a great range of animals in which nature deals with the challenge of an extreme
and plants that have special requirements and make spe- environment.
xviii F tundra
the Tundra Important? expected elsewhere. There are many questions that need to
be answered about the role of the polar regions in global
It is evident that the tundra has much interest for scien- climate change, and this work is advancing rapidly. One of
tists, whether geographers who study the topography of these questions concerns the development of the so-called
ice-created landforms, biologists who examine cold tol- ozone hole, in which changes in atmospheric composition,
erance among plants and animals, or soil scientists who particularly in the gas ozone, can lead to potentially dan-
investigate the effect of freeze-thaw cycles on soil structure. gerous rays reaching the surface of the Earth. Knowledge
But the tundra is also important to people who may not be of this phenomenon has improved greatly in recent years.
directly involved in scientific investigations. In the future Remedial actions seem to be having some effect, although
the importance of the tundra may well become increas- there are considerable year-to-year variations and there
ingly appreciated as the many valuable features of this remain many unanswered questions.
biome become more widely known. This book is designed Tundra is one of Earth’s few remaining wilderness areas,
to assist in this process. and a consequence of this is that there is very little pollu-
The highly adapted plants and animals of the tundra tion resulting from human activities. But even here pesti-
contain within their cells genetic information that deter- cides manage to penetrate. The persistent pesticide DDT, for
mines their hardiness. There are genes, for example, that example, has been found in the flesh of Antarctic penguins,
provide plants with the ability to resist freezing or to which provides a salutary warning of just how widely dis-
survive even when cells become frozen. There are genes persed are the chemicals people introduce into their envi-
within polar bears that enable them to live a healthy and ronment. The relative purity of the tundra ecosystems makes
vigorous life despite the fact that their diet (mainly seals) them particularly useful in monitoring the movement of
contains extremely high levels of fat. Understanding how pesticides and their continued presence in the global envi-
either of these two biochemical systems operates could be ronment.
of great value in agriculture and human medicine, respec- The tundra is becoming increasingly accessible to peo-
tively. The animals and plants of the tundra, therefore, ple as tourism develops and the popularity of adventure
can be regarded as an enormous untapped resource of holidays and ecotourism grows. Undisturbed habitats will
information that may one day enable people to improve be increasingly exposed to stress from people, whether they
their lives. come to admire the scenery and wildlife or to exploit the
During the history of the Earth the tundra has expe- mineral reserves found in many tundra areas. Antarctica is
rienced changing fortunes. There have been long periods currently protected by international treaty, but the increas-
when this biome has been absent or found only on moun- ing scarcity of oil reserves and the continued demand for
taintops. There have also been times when the tundra has minerals will undoubtedly bring increased demand for
spread and expanded over continents, the ice ages. It is vital exploitation even in this last wilderness.
that the climatic mechanisms underlying these changes be It is this combination of potentials and problems that
understood. There is an apparent pattern to these events makes the tundra a biome that demands attention. Despite
that requires detailed study so that the future behavior of the its wide geographical distribution, the tundra has proved
tundra under ever-changing conditions can be predicted. hard to explore and understand. The severe conditions that
Geologists collect evidence for past changes in the distribu- make the tundra so distinctive have also made exploration
tion of tundra in order to detect patterns and understand and scientific research particularly difficult. There is much
the processes involved in these global climatic changes. that is yet to be discovered about the tundra, but such dis-
At the present day the climate of the polar regions covery depends upon our appreciating and understand-
appears to be changing more rapidly than that of other ing what is currently known about this remote and unique
parts of the world. It is not yet understood why this should biome. This is the task of the current book.
1
Climate of the Tundra
tundra distribution This is called its latitude. Lines of latitude run around
the Earth north and south of the equator, ranging
The map on page xvii shows the broad pattern of tundra dis- from 0° to 90°. Those with high values (close to the
tribution over the Earth’s land surface. As explained in the poles) are termed the high latitudes, and those close
introduction, there are two types of tundra present on this to the equator have low values and are called low
map: One is the polar tundra, restricted to the high latitudes
latitudes. Polar tundra is found in the high latitudes.
(see sidebar), and the other is the alpine tundra, found only
These lines are, of course, purely conceptual but are
on mountains but that can occur in any latitude, both high
an important means of defining any location on the
and low, depending simply on the presence of mountains.
surface of the Earth. A second coordinate is required
Polar tundra is found mainly in the Northern
to fix a precise position, and this is given by its lon-
Hemisphere, in the region called the Arctic. The word Arctic
gitude. Lines of longitude run from pole to pole over
refers to the northern constellation of stars called the Great
Bear (Arktos is Greek for “bear”). The northern distribu- the Earth’s surface and are numbered from 0° to
tion limit of the tundra, as shown in the diagram on page 359°, with the base line (0°) running through Green-
3, is defined by the Arctic Ocean, and the presence of an wich in London, England. This choice of the zero line
ice sheet over Greenland also restricts its occurrence on of longitude is a consequence of history, but it has
that large island. To the south it is bordered by the boreal proved convenient because the 180° line (the Inter-
forest, or taiga, where coniferous trees and birches become national Date Line) runs through the Pacific Ocean
the dominant feature of the landscape. The Arctic tundra rather than over land surface, where changing
covers approximately 29,000 square miles (7.5 million ha), date with location on either side of a line would be
which represents 5.5 percent of the total land surface of the extremely confusing for the local inhabitants.
Earth. It is divided into three blocks: North America with
n 1 n
F tundra
Eco-Tundra-002.eps
North Pole
90°N
66°23’N
Arctic Circle
60°N
30°N
equator 0°N
30°S
11,000 square miles (2.8 million ha), Eurasia with 10,000 Latitude is measured as the angle at the center of the Earth for
square miles (2.6 million ha), and Greenland and Iceland each location on the surface, taking the equator as 0° and the
with 8,000 square miles (2.1 million ha). poles as 90° north and south. The latitudes of the tropics of
Cancer and Capricorn together with the Arctic and Antarctic
Much of Alaska is covered by tundra, extending through
Circles are shown.
the Aleutian Islands, which form a chain out into the North
Pacific Ocean. The northern part of Canada has a tundra
cover running from the Northwest Territories around the Greenland is free from a permanent cover of ice, leaving lit-
southern shore of Hudson Bay and into northern Quebec tle room for tundra, but the more southerly island of Iceland
and Labrador. Extending out into the Arctic Ocean are in the North Atlantic Ocean has a covering of tundra. The
numerous islands, including Baffin Island and Ellesmere mainland of Europe has only a fringe of polar tundra in the
Island, all clothed with tundra. Only the coastal fringe of north of Scandinavia, mainly in northern Norway, but the
Climate of the Tundra F
ice-free areas of the islands of Svalbard in the Arctic Ocean lying off that coast. It also covers the eastern parts of Siberia,
bear tundra. This biome is also found along the fringe of the where the tundra reaches the Bering Sea, separating Russia
Arctic Ocean in northern Russia, together with the islands from Alaska. Effectively, the polar tundra forms a circle
around the Arctic Ocean; it has a circumpolar distribution.
Polar tundra is also found in the Southern Hemisphere
The Arctic tundra regions showing the extent of the Arctic
but is much less abundant. The main difference in the polar
polar tundra and the position of the magnetic North Pole.
Greenland is covered by an ice sheet, so tundra vegetation
regions of the Southern Hemisphere is the presence of a
occurs only around its coasts. Much of the Arctic Ocean is continental landmass, Antarctica. Unlike the North Pole,
covered by permanent sea ice, which extends yet farther as which is situated over the Arctic Ocean, the South Pole lies
fragmented pack ice, especially in winter. within the landmass. The main effect of the presence of this
Eco-Tundra-003.eps
Ellesmere Island
North Pole
Novaya Zemlya
Hudson Bay
Baffin Island
Svalbard
extent of tundra
Greenland Iceland
© Infobase Publishing
F tundra
Eco-Tundra-005.eps
0°
AT L A N T I C O C E A N
Antarctic peninsula
Weddell Sea
Transantarctic Mountains
Lesser Antarctica
PACIF
Greater Antarctica
IC O
CEA
EAN
N
OC
An alpine meadow in Switzerland shows the colorful flowers wherever mountains are high enough to result in restricted
for which such habitats are famous. (Peter D. Moore) forest growth. When conditions become too cold or wind
strengths too high for the survival of trees, tundra is able to
thrive. Precisely how high a mountain needs to be in order
to provide the right conditions for tundra varies with lati-
South America. This is called the Antarctic Peninsula, and tude.
this, together with a number of islands lying off its coast, is The mountain ranges of the world are shown in the map
referred to as Maritime Antarctica. This region extends far- on page xvii. As can be seen, they are quite widely scattered,
ther north than most of the Antarctic landmass and so pen- but where mountains occur they often form groups or long
etrates into relatively lower latitudes. Maritime Antactica chains. The reasons for this will be investigated later (see
is also surrounded by the sea, and these two factors, lower “Tectonic History of the Tundra Lands,” pages 33–35). In
latitude and oceanic conditions, result in a slightly warmer North and South America the main mountain systems lie on
climate, reducing ice cover and permitting the develop- the western sides of the continents and form roughly north-
ment of tundra vegetation. Most of the very limited tundra south running ridges. The mountain ranges of Alaska and
found in the Southern Hemisphere lies within the Maritime the Mackenzie Mountains of Canada merge in the south-
Antarctic. ern part of their range to create the Rocky Mountains, the
The polar tundra is thus restricted overall to the higher spine of western North America. The Cascade Mountains of
latitudes. Most tundra in the Northern Hemisphere lies Washington and Oregon and the Sierra Nevada of California
north of 70°N latitude. In the Southern Hemisphere the are essentially coastal branches and parallel systems along-
effect of the Antarctic ice sheet chills the southern ocean. side the main Rocky Mountain ridge. The range continues
Tundra is found on islands as far north as 55°S, but its limit south into Mexico as the Sierra Madre, which run through
is still clearly associated with relatively high latitude. This Central America and eventually link up with the Andes
is not the case for alpine tundra. Alpine tundra is found Mountains that line the west coast of South America. At
F tundra
the southern tip of that continent the Andes pass through the edge of the Ross Ice Shelf and the Ross Sea. In areas such
Patagonia and end in the extremity of Tierra del Fuego, as these, the polar and alpine tundras meet and are indistin-
where the alpine tundra and polar tundra meet. guishable from one another.
In Europe an extensive mountain chain runs approxi-
mately from northeast to southwest on the western fringe of
Scandinavia, mainly in Norway. The mountains of Scotland
in the British Isles are a southerly outlier of this system. n Global Climate Patterns
The remaining mountain chains of Europe are east-west in Understanding the distribution pattern of tundra demands
their orientation, a fact that has had considerable implica- a close study of climate patterns over the Earth. Why are the
tions for the biogeography of that continent (see “Glacial Arctic and Antarctic colder than the Tropics? And why is
History of the Earth,” pages 176–178). In the west of Europe, tundra found at lower latitudes in the Southern Hemisphere
the Pyrenees Mountains separate France from the Iberian than in the Northern Hemisphere? To answer these ques-
Peninsula (Spain and Portugal). To the north of Italy lie tions, it is necessary to consider the distribution of energy
the Alps, where the highest mountains in Europe are found around the Earth, and this requires information about the
(see illustration on page 5). Farther east, the Carpathian movements of air masses in the atmosphere and water
Mountains form a crescent on the western fringes of the masses in the oceans.
Black Sea. The first point to examine is the way in which energy
North Africa also has a mountain range, the Atlas arrives at the surface of the planet. Energy on Earth ulti-
Mountains, that lie in an east-west arrangement, running mately comes almost entirely from the Sun. There are some
through Morocco and Algeria. The remaining African exceptions to this statement: The hot interior of the Earth
mountains are concentrated in the eastern regions, running releases some geothermal energy, natural radioactive decay
south from the Ethiopian highlands. The Great Rift Valley also produces some energy, and the gravitational pull of the
systems of Kenya, Tanzania, Uganda, and Congo are rich Moon creates tidal energy. But the vast bulk of the energy
in high mountains, many of them volcanic, whose chains arriving on Earth is in the form of solar radiant energy.
run across the equator. In southern Africa the Drakensberg Energy comes in a variety of forms, which physicists dif-
Mountains lie along the eastern side of the continent. ferentiate in terms of wavelength. Light itself, for example,
In Asia the north-south running Ural Mountains sepa- consists of a spectrum of energy of different wavelengths,
rate this continent from Europe and extend from the Arctic from the very short violet end to the longer-wavelength
toward the Caspian Sea. It is in Asia that the highest moun- red end. This accounts only for the visible forms of energy;
tain in the world, Mount Everest (29,029 feet; 8,848 m) is there are many shorter-wavelength forms, such as ultra-
located, set within the Himalayas. This massive mountain violet and even shorter gamma rays, and also longer-wave-
block runs from Afghanistan in the west, where the moun- length forms, such as infrared, heat energy that can be felt
tains are called the Hindu Kush, through northern Pakistan, but not seen. The Sun emits energy over a very broad spec-
India, Tibet, and Nepal, ending in eastern China. They trum, but most of the solar radiation that reaches the sur-
extend northward around the Gobi Desert into Mongolia. In face of the Earth lies between 0.000001 and 0.00016 inches
eastern Russia lie the Altai Mountains, the Sayan Mountains, (0.00000025 and 0.0004 cm). Such minute numbers are dif-
and the Transbaikalian Mountains, reaching over 11,000 ficult to work with, so the measurements used by scientists
feet (3,400 m). Australia is not rich in high mountains but are metric, and the wavelength is expressed in micrometers
has the north-south orientated Great Dividing Range along (1 μm = 1 millionth of a meter). The Earth receives energy
its eastern coast. mainly in the 0.2 μm to 4.0 μm wavebands. The Earth’s
Alpine tundra is, therefore, much more widely dispersed atmosphere does not absorb energy within this range, so
than polar tundra, but separating the two is not always pos- it passes directly to the surface of the planet. Shorter wave-
sible because some high mountains lie within or adjacent to lengths, however, such as some ultraviolet radiation, are
the polar regions. In North America, for example, the Brooks absorbed by certain components of the atmosphere (see
Range of mountains lies along the southern edge of the polar “Ozone Holes,” page 226). In addition, dust in the atmo-
tundra in Alaska, and the Laurentian Mountains are situated sphere together with cloud cover can result in the absorp-
within the polar tundra of Labrador. In Europe the moun- tion or reflection of the some of the incoming energy.
tains of Norway merge with polar tundra at their northern When solar energy strikes the surface of the Earth,
extremity, as do the Ural Mountains of Asia. In eastern Asia much of it is likely to be absorbed. An object placed in sun-
the mountains of Siberia also merge with the tundra of the light becomes warm as a result of the absorption of energy,
Arctic. Antarctica has a ridge of mountains running along and as its temperature rises it radiates some energy back
the length of the Antarctic Peninsula and extending around into the atmosphere. In a similar way a land surface heated
Climate of the Tundra F
Eco-Tundra-007.eps
short wavelength
solar radiation space penetration
radiation to space
atmosphere
absorption
surface heating
penetration
long wavelength
Earth heat radiation
© Infobase Publishing
The greenhouse effect. Solar energy arriving at the Earth’s water vapor, carbon dioxide, oxides of nitrogen, and several
atmosphere consists mainly of short wavelength radiation. others, which have become known as greenhouse gases. It is
Some of this is reflected, a little is absorbed by the atmosphere, called the greenhouse effect because the entire Earth system
but much of the energy passes through to the Earth’s surface.
is acting like an enormous greenhouse. Light passes through
Here it heats the land and water, and energy is then radiated
back in the form of long wavelength heat. Some of the glass panes of a greenhouse without being absorbed to any
radiated heat penetrates the atmosphere and is lost to space, great extent by the glass. The glass does not itself become
but much is absorbed by the atmosphere, which then radiates hot. When light strikes the soil within the greenhouse it
heat in all directions, including some back to Earth. Certain causes the soil to become hot, and the reradiated heat does
gases in the atmosphere, the so-called greenhouse gases, not pass out through the glass roof because long-wave heat
are particularly efficient at trapping heat and act as a thermal
energy does not easily penetrate glass but remains within
blanket around the Earth.
the greenhouse, warming the air it contains. Without the
greenhouse effect the Earth would rapidly cool as soon as
the Sun set in the evening. Nights would become unbearably
by sunlight radiates heat energy, and heat energy has a lon- cold, and life on the planet would be much more difficult.
ger wavelength than light energy. Some of the heat energy As will be discussed later (see sidebar “Carbon Cycle and
radiated by land surfaces is taken up by the atmosphere, the Greenhouse Effect,” page 220), changes in the intensity
because the atmosphere is less transparent to longer-wave- of the greenhouse effect in the atmosphere have a profound
length heat than it is to light. By absorbing the reradiated influence on global climate.
heat energy in this way, the atmosphere acts like a ther- There is a complication in this process because the
mal blanket around the Earth, retaining heat that would amount of energy absorbed by a surface varies. A black
otherwise be rapidly lost, as shown in the diagram above. surface absorbs sunlight and becomes heated much more
This absorption, known as the greenhouse effect, is due to effectively than a white surface, and a dull, matte surface
the presence of certain gases in the atmosphere, including absorbs more efficiently than a shiny surface. The efficiency
8 F tundra
of a surface in absorbing light and becoming heated in the twice a year, when the length of day and night is equal
process is dependent on how much of the incoming energy over the whole planet. The Earth’s tilt on its axis of 23.5°,
is reflected, and the reflectivity of a surface is expressed as its however, means that this situation is not maintained. The
albedo (see sidebar below). Sun is overhead at the tropic of Cancer (66.5°N) during
As can be seen from the figures in the sidebar, differ- the Northern Hemisphere summer and at the tropic of
ent types of land surfaces and vegetation have a strong effect Capricorn (66.5°S) during the Southern Hemisphere sum-
on how much of the Sun’s energy is absorbed. The figures mer. Throughout the year, however, the angle of the Sun
for the surface of water also show that albedo depends quite within the low-latitude Tropics is high when compared
strongly on the angle of the Sun. These aquatic data apply with that of the higher latitudes. In the polar regions the
to a water body with a calm and smooth surface. Waves can angle of the Sun above the horizon is never high. This dif-
create very complex patterns of reflection. These variations ference in the solar angle between the poles and the Tropics
in albedo, depending on the nature of the surface, have an means that energy is spread over a wide area in the high
important impact on the world’s climate. Oceans and land latitudes, whereas it is concentrated in a smaller area in
surfaces, forests and deserts, ice sheets and wetlands all have the low latitudes. The heating potential of radiation in the
different capacities to absorb and reflect incoming radiation, polar regions is thus lower than that found in the Tropics,
which influences the degree to which they become warm in as shown in the diagram on page 9.
the sunshine and also the way they radiate heat back into the Seasonal differences in day length are also more pro-
atmosphere. nounced in the high latitudes. In the Tropics there is some
The angle at which radiation strikes the Earth’s sur- variation in day length depending on the position of the
face also influences absorption and reflection. The dia- overhead Sun at noon, but the variation is very small. Days
gram shows the way in which the Earth’s curvature results are roughly equal to nights in their length at all times of the
in energy being received at different angles depending year. At higher latitudes, however, there are increasingly wide
on latitude. The Sun is overhead at noon at the equator disparities in day length with season. At 60°N (the latitude
Albedo
When light falls upon a surface, some of the energy is of the light that falls upon it, which is why mirrors are usu-
reflected and some is absorbed. The absorbed energy is ally cold to the touch. The angle at which light falls upon
converted into heat and results in a rise in the temperature a surface also affects the proportion of light absorbed and
of the absorbing surface. What proportion of the light is reflected; light arriving vertically is reflected less than light
absorbed and what proportion reflected varies with the arriving from a low angle.
nature of the surface. A dark-colored surface absorbs a Scientists express the degree of reflectivity of a
greater proportion of the light energy and reflects less than surface as its albedo. It is stated as the amount of light
a light-colored surface; it therefore becomes warmer more reflected divided by the total incident light. In order of
rapidly. A dull, matte-textured surface absorbs energy efficiency of reflectivity, the following list gives examples
more effectively than a shiny one. A mirror reflects most of the albedo of various surfaces.
(above) Light from the Sun arrives at the Earth as parallel at high latitude. There are two means by which the Earth’s
beams, but the light heats different extents of surface area energy can be redistributed in this way, the atmosphere and
depending upon the angle at which it arrives. Polar regions the oceans.
receive less intense energy than the tropical areas. The tilt of
the Earth upon its axis means that the position of the overhead
noonday Sun varies with season.
(below) The circulation of the atmosphere occurs in a series of
cells, creating areas of low pressure near the equator and in
the region of the polar fronts, and areas of high pressure close
to the tropics of Cancer and Capricorn and at the North and
of Anchorage, Alaska, the southern tip of Greenland, and South Poles.
Oslo, Norway) the days of summer are extremely long, while Eco-Tundra-009.eps
the days of winter are extremely short. Beyond the Arctic
ITCZ = Intertropical Convergence Zone
Circle (66.5°N), there is at least one day in the year when the
Sun does not set. This difference in summer and winter day tropic of Cancer high
length results in a strong seasonal difference in temperature low polar front
in the high latitudes. The length of the summer season also
becomes shorter close to the poles. Within 450 miles (725
km) of the North Pole, for example, at Cape Morris Jessup in North Pole
northern Greenland (about 84°N), the length of time when
conditions are suitable for plant growth may be as little as high
three or four weeks.
When the three factors of solar angle, area of energy dis-
persal, and seasonal differences are put together, it is evident
equator
that the polar regions will have a smaller input of energy per
unit area of Earth’s surface than the Tropics. Lower energy
low ITCZ
levels result in lower temperatures overall, which means that
snowfall and persistent snow cover become more likely in
the high latitudes, and this has a feedback effect on albedo,
resulting in even more of the incoming energy being lost by
reflection.
The average temperature at the equator is 79°F (26°C), high
which varies very little with season, while at 80°N the January
South Pole
average is –22°F (–30°C) and the July average is 37°F (5°C).
At this high latitude the Earth is actually losing more energy
than it is receiving, so the Arctic can be regarded as an energy
sink. But the Arctic is not becoming colder year by year, so the tropic of Capricorn low polar front
energy it loses must be replaced by energy from other parts high
of the planet. There must be energy movement from areas with
© Infobase Publishing
an abundant supply, such as the Tropics, to the energy sinks
10 F tundra
n The Atmosphere and Climate surface. These belts are where the world’s deserts are mainly
found.
Heating in the Tropics and cooling at the poles results in a The hot, dry air of the desert belts may be deflected back
global energy imbalance, and this in turn gives rise to atmo- toward the equator, or they may travel toward the poles. If
spheric turbulence. Warm air has a lower density than cold they take the latter route, then these air masses meet cold,
air, so there is a tendency for cool, dense air to move toward polar air moving in the opposite direction. Where the two
the equator, pushing the warm, low-density air upward. Air air masses collide they create a boundary zone, called the
masses approach the equatorial regions from both north and polar front, and this region is characterized by unstable
south of the equator, creating an intertropical convergence weather conditions. The precise position of the polar front
zone where the two air masses meet. The warm, moisture- is very variable in space and time, but the mix of air masses
laden air rises and cools in the process (see sidebar “Lapse results in the cyclonic depressions and accompanying pre-
Rate,” page 22). Cooler air is less able to hold moisture, so cipitation typical of latitudes roughly between 40° and 60°
water condenses into raindrops, and the equatorial regions north and south of the equator. The polar front is also asso-
receive high levels of precipitation, resulting in the develop- ciated with a strong wind, the jet stream, which moves from
ment of the tropical rain forests. In the upper atmosphere, west to east at an altitude of around 35,000 feet (10,000 m)
around 10 miles (16 km) above the ground, the air that has at the boundary between the lower part of the atmosphere,
been forced upward moves out toward the poles, as shown
in the diagram on page 9, and since the air has now cooled
and become denser, it falls toward the Earth’s surface once
The atmosphere is divided into a series of layers, becoming
more, creating a high pressure belt between about 25° and less dense with altitude. The lights of the aurora occur in the
30° north and south of the equator. These falling air masses thermosphere, between 60 and 260 miles (100 and 420 km)
have lost their moisture over the equatorial regions, so they above the Earth’s surface. This is the layer of the atmosphere in
cause dry conditions when they arrive back at the Earth’s which satellites orbit the Earth.
Eco-Tundra-010.eps
400
600
satellite Exosphere
300
space shuttle
400
rocket
200
aurora
200
Thermosphere
100
balloon 100
meteors
40 airliner Mesosphere
Mt. Everest 5.5 miles (8.8 km) 50
20 Stratosphere
10 20
5 Troposphere 10
miles km
© Infobase Publishing
Climate of the Tundra F 11
the troposphere, and the upper part of the atmosphere, the latitude. In the region where the Southern Ocean meets the
stratosphere. This boundary, as shown in the diagram on page South Pacific, however, the southern polar front is located
10, is called the tropopause. The jet stream causes cyclones along latitude 60°S, as shown in the diagram on page 17.
(low-pressure systems) and anticyclones (high-pressure sys- Some of the air forced upward in the turbulent regions
tems) to move around the world in an easterly direction. The around the polar fronts moves on toward the poles at high
latitude of the polar front and the jet stream varies with the altitude. Here the air becomes very cold and dense, so it
season. In summer in North America the polar front and begins to lose altitude, and its descent creates another high
jet stream lie roughly along the Canadian border, dipping pressure zone over the poles, as shown in the diagram on
south to Washington, D.C., in the east. In winter it runs page 9. As in the high-pressure zone of the outer Tropics and
approximately from Los Angeles through northern Texas to sub-Tropics, the outcome is the production of low-precipi-
South Carolina. tation, desert conditions. But in the polar regions the lack
In the Southern Hemisphere the polar front encircles of precipitation does not result in drought at the surface of
Antarctica and lies within the Southern Ocean. For much of the Earth because the very low temperature at ground level
its length, where the Southern Ocean meets the South Atlantic restricts the degree of evaporation taking place, so water
Ocean and the Indian Ocean, it lies approximately along 50°S may accumulate over low-lying parts of the landscape even
though water input from the atmosphere is very restricted.
The oceanic currents play an important part in the
redistribution of energy around the world. In the North
Atlantic warm tropical waters from the Caribbean move up
the western edge of Europe and enter the Arctic Ocean. This
n The Oceans and Climate
penetration of warmth into the Arctic is much less strong in The atmosphere is not the only determinant of global pat-
the Pacific, so the north coast of Alaska is much colder than terns of climate; the oceans also have a profound influence.
the north of Scandinavia. Water has certain physical properties that affect climate. A
Eco-Tundra-011.eps
N. Atlantic C.
Gulf Stream
Guinea C.
S. Equatorial C.
S. Equatorial C.
warm currents
Humboldt C. Falkland C. Brazil C. Benguela C. Agulhas C. cool currents
© Infobase Publishing
12 F tundra
Eco-Tundra-012.eps
The global circulation of the Earth’s oceanic waters is called Archangel, at a latitude of 65°N in western Russia, lies on
the oceanic conveyor belt. Warm, low-salinity, and low-density the edge of the Barents Sea and has an average annual tem-
water moves along the surface of the ocean. On arrival in the perature of 33°F (0.4°C) and an average temperature for its
North Atlantic Ocean the water cools and forms a deep-water coldest month of 5°F (–15°C). But in the continental east
current that flows in the opposite direction.
of Russia, Verkhoyansk, in Siberia, which lies on the same
latitude, has an average annual temperature of 3°F (–16°C),
and the average for its coldest month is as low as –58°F
sample of water both gains and loses heat more slowly than (–50°C). The oceanic regions of Russia are kept warmer by
an equivalent weight of rock. Areas of land surrounded by the proximity of the sea. Oceanic regions also receive more
water are, therefore, protected to some extent from extreme precipitation because the sea is a source of atmospheric
variations in temperature; land close to the ocean is kept moisture. Thus, Archangel receives 18 inches (47 cm), while
cool in summer and relatively warm in winter. The proxim- Verkhoyansk has only five inches (13 cm) of precipitation
ity of the sea, therefore, has a considerable influence on the each year.
climate of a region. A site close to the sea is said to have an The oceans also act as channels for the redistribution of
oceanic climate, whereas one situated far from the ocean’s energy around the world. Surface water moves around and
influence has a continental climate. For example, the city of between the great oceans in the form of currents, as shown
Climate of the tundra F 1
of snow in the High Arctic and may amount to only about lack of snow means that vegetation is quickly able to take
four inches (10 cm) per year. The most northerly parts of advantage of rising light levels and increasing day length in
Arctic Canada have only two inches (5 cm) of precipitation the spring and begin its growth more rapidly. Early growth
per year. The very cold conditions, however, mean that the then attracts grazing animals, so the shallow snow cover has
evaporation rate is very low, so drought is experienced only extensive implications for the ecology of the tundra.
where there is very good soil drainage, such as on sand and The wind is itself an important climatic factor in polar
shingle ridges. tundra. High winds are frequent, and these bring with them
Where there is permanent ice cover, as in the case of the a wind chill effect. Wind constantly removes the warm layer
Greenland ice sheet, the climate is even more severe, with of air around an object or an organism, so heat energy is
the mean monthly temperature remaining below freezing taken from the object faster than in still air. From the point
throughout the year. Under such conditions there is no veg- of view of a living organism, the conditions feel much colder
etation. The shiny white surface of the ice has a very high when there is a wind blowing. Human beings subjected to
albedo, so almost all of the incident energy is reflected and a wind of 50 feet per second (15 m/sec) in air at 32°F (0°C)
very little is absorbed. The middle of the Greenland ice sheet experience the same degree of cold that they would in still
has a mean annual temperature of –18°F (–28°C), and even air at 14°F (–10°C). So conditions in the tundra are even
in summer the temperature rarely rises above freezing. Ice more unpleasant for living creatures than is apparent simply
sheets are maintained by the arrival of new material in the from the temperature data. An additional factor that makes
form of snowfall, replacing any loss from melting. In the things even worse is the way in which ice crystals are often
case of the center of the Greenland ice sheet, however, pre- suspended in the moving air, acting as an abrasive agent in
cipitation is very low. The ice mass survives because wast- the wind. Very few plants can survive exposure to ice-laden
age by melting is also extremely slow. Precipitation is higher winds that can strip the surface cells from a leaf.
only where the ice sheet lies close to the sea and the climate Although overall temperatures are low, and the grow-
is more oceanic. ing season is short in the Arctic, this does not mean that
The Low Arctic consists of the remaining regions of conditions are eternally cold. There are occasions when the
the tundra, mainly the northern parts of Siberia, Alaska, air temperature can be quite high. In the Taymyr Peninsula
Canada, coastal Greenland, and Iceland. The Low Arctic in the north of Siberia, for example, July temperatures often
has a climate that is cold but not quite as extreme as that of exceed 68°F (20°C) during the day, and temperatures as high
the High Arctic. The mean monthly temperature is below as 84°F (29°C) have been recorded. But the temperature is
freezing for between seven and nine months of the year, and strongly affected by local conditions of topography (see
the snow-free period is between three and four months in “Microclimate,” pages 18–21), and this means that the sum-
length. This means that plants may be able to grow for as mer climate is very variable over relatively short distances.
long as five months during the summer. As in the case of the This has a considerable effect on vegetation. The climate
High Arctic, the soil remains permanently frozen below the also depends on the distance to the ocean, especially if it is
surface layer, with the temperature at 100 feet (30 m) below influenced by the warm waters of the Gulf Stream (see “The
the surface remaining at about 25°F (–4°C). Precipitation is Oceans and Climate,” pages 11–14). The cooling effect of
generally still extremely low, at about eight inches (20 cm) the ocean during the summer is also an important factor in
per annum, but the ground is generally wet during the sum- determining the climate of the tundra. If the summer tem-
mer because of the low evaporation rate. In particularly oce- perature rises far enough, then trees can survive and tun-
anic regions, such as the coastal tundra around the Bering dra vegetation is lost. This is why all polar tundra habitats
Sea, precipitation may be as high as 30 inches (76 cm). lie relatively close to the sea, because the more continental
One result of the low precipitation in the Arctic tundra climates found deep within land masses have high summer
is that snow cover is generally thin. Vegetation is low-lying temperatures and become clothed with forest. High moun-
and uniform, so this does not influence the shallow covering tain tops are an exception, of course, but the lowland tundra
of snow. Wind speeds, on the other hand, can be very high, of the polar regions is almost always found within 300 miles
and this can lead to drifting and the development of deeper (500 km) of the sea or other large body of water.
snow patches in the lee of rocks and promontories. The So the polar tundra is oceanic in its distribution, and
shallow snow cover has important implications for the tem- the proximity of the ocean keeps summer temperatures cool
perature of the soil and the penetration of frost. Snow acts and winter temperatures less intensely cold than those at
as an insulating blanket over the surface of soil, so a poor locations deeper in the continental interiors. The seasonal
covering of snow leads to deeper frost penetration. This, in variation in temperature, therefore, is smaller than that
turn, may lead to the development of distinctive features in found in the continental regions of the subarctic lands. In
the topography, such as the ice mounds of palsa mires (see such locations the summer temperatures are higher (leading
“Tundra Wetlands,” pages 74–78). On the other hand, the to the development of forests), but the winter temperatures
1 F tundra