138 The Fossil Record

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138 The Fossil Record


Key Idea: Fossils provide a record of the appearance and of the horse (below right) from the ancestral Hyracotherium
extinction of organisms. The fossil record can be used to to modern Equus is well documented in the fossil record.
establish the relative order of past events. The rich fossil record, which includes numerous transitional
Fossils are the remains of long-dead organisms that have fossils, has enabled scientists to develop a robust model
escaped decay, with their remains becoming mineralized. of horse phylogeny (evolutionary history). Horse evolution
Fossils provide a record of the appearance and extinction exhibits a complex tree-like lineage with many divergences.
of organisms, from species to whole taxonomic groups. Many species coexisted for some time over the 55 million
Once this record is calibrated against a time scale (by using year evolutionary period. The environmental transition from
dating techniques), it is possible to build up a picture of the forest to grasslands drove many of the changes observed in
evolutionary changes that have taken place. The evolution the equid fossil record.

Profile with Sedimentary Rocks Fossil Evidence of Horse Evolution


Containing Fossils 0
Ground surface Recent fossils are Equus (grassland)
found in more recent
sediments 5
The more recent the
Youngest layer of rock, the more
sediments resemblance there is
between the fossils found 10
in it and living forms.
Numerous extinct
species 15
The number of extinct
species is enormously
greater than the
number living today. 20
oga sraey fo snoilliM

Fossil types differ in 25


each stratum Merychippus
Fossils found in a given (grassland)
layer of sedimentary
rock generally differ in 30
significant respects from
those in other layers.
More primitive fossils 35
are found in older
sediments
Phyla are represented 40 Mesohippus
Oldest by more generalized (forest)
sediments forms in the older layers,
and not by specialized
forms (such as those 45
alive today).
Rock Strata are Layered Through Time 50
Rock strata are arranged in the order that they were deposited
(unless they have been disturbed by geological events). The most
recent layers are near the surface and the oldest are at the bottom.
55 Hyracotherium
New Fossil Types Mark Changes in Environment (Eohippus) (grassland)
In the rocks marking the end of one geological period, it is common
to find new fossils that become dominant in the next. Each geological The fossil record of the horse provides much evidence for evolution
period had an environment very different from those before and (change in body size, limb length, tooth structure, toe reduction). The
after. Their boundaries coincided with drastic environmental changes genus Equus (the modern horse) is the only living genus of what was
and the appearance of new niches. These produced new selection a large and diverse group of animals. Observation of the leg bones of
pressures resulting in new adaptive features in the surviving species various ancestors of Equus show a progressive loss of the outer toe
as their populations responded to the changes. bones, leaving the single middle bone and hoof supporting the animal.

1. How does the fossil record provide a record of evolutionary change over time:

2. In which way does the equid fossil record provide a good example of the evolutionary process?

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