Tiger

The tiger (Panthera tigris) is a member of the genus Panthera and the largest living cat species native to Asia.
It has a powerful, muscular body with a large head and paws, a long tail and orange fur with black, mostly
vertical stripes. It is traditionally classified into nine recent subspecies, though some recognise only two
subspecies, mainland Asian tigers and the island tigers of the Sunda Islands.

Throughout the tiger's range, it inhabits mainly forests, from coniferous and temperate broadleaf and mixed
forests in the Russian Far East and Northeast China to tropical and subtropical moist broadleaf forests on the
Indian subcontinent and Southeast Asia. The tiger is an apex predator and preys mainly on ungulates such as
deer and wild boar, which it takes by ambush. It lives a mostly solitary life and occupies home ranges, which it
defends from individuals of the same sex. The range of a male tiger overlaps with that of multiple females with
whom he mates. Females give birth to usually two or three cubs that stay with their mother for about two
years. When becoming independent, they leave their mother's home range and establish their own.

Since the early 20th century, tiger populations have lost at least 93% of their historic range and are locally
extinct in West and Central Asia, in large areas of China and on the islands of Java and Bali. Today, the tiger's
range is severely fragmented. It is listed as Endangered on the IUCN Red List of Threatened Species, as its
range is thought to have declined by 53% to 68% since the late 1990s. Major threats to tigers are habitat
destruction and fragmentation due to deforestation, poaching for fur and the illegal trade of body parts for
medicinal purposes. Tigers are also victims of human–wildlife conflict as they attack and prey on livestock in
areas where natural prey is scarce. The tiger is legally protected in all range countries. National conservation
measures consist of action plans, anti-poaching patrols and schemes for monitoring tiger populations. In
several range countries, wildlife corridors have been established and tiger reintroduction is planned.

The tiger is among the most popular of the world's charismatic megafauna. It has been kept in captivity since
ancient times and has been trained to perform in circuses and other entertainment shows. The tiger featured
prominently in the ancient mythology and folklore of cultures throughout its historic range and has continued
to appear in culture worldwide.

Etymology

The Old English tigras derives from Old French tigre, from Latin tigris, which was a borrowing from Classical
Greek τίγρις 'tigris'.[4] Since ancient times, the word tigris has been suggested to originate from the Armenian
or Persian word for 'arrow', which may also be the origin of the name for the river Tigris.[5][6] However, today, the
connection between the animal and the river is doubted and they are likely to be homonyms.[6]
Taxonomy Tiger
Temporal range:

In 1758, Carl Linnaeus described the tiger in his work Systema

Naturae and gave it the scientific name Felis tigris, as the
genus Felis was being used for all cats at the time. His
scientific description was based on descriptions by earlier
naturalists such as Conrad Gessner and Ulisse Aldrovandi.[2]
In 1929, Reginald Innes Pocock placed the species in the
genus Panthera using the scientific name Panthera tigris.[7][8] A Bengal tigress in Kanha Tiger Reserve, India

Conservation status
Subspecies

Nine recent tiger subspecies have been proposed between the

early 19th and early 21st centuries, namely the Bengal, Endangered (IUCN 3.1)[1]
Malayan, Indochinese, South China, Siberian, Caspian, Javan,
Bali and Sumatran tigers.[9][10] The validity of several tiger CITES Appendix I (CITES)[1]

subspecies was questioned in 1999 as most putative

Scientific classification
subspecies were distinguished on the basis of fur length and
colouration, striping patterns and body size of specimens in Domain: Eukaryota

natural history museum collections that are not necessarily

Kingdom: Animalia
representative for the entire population. It was proposed to
recognise only two tiger subspecies as valid, namely P. t. tigris Phylum: Chordata
in mainland Asia and the smaller P. t. sondaica in the Greater
Class: Mammalia
Sunda Islands.[11]

Order: Carnivora
This two-subspecies proposal was reaffirmed in 2015 through
a comprehensive analysis of morphological, ecological and Suborder: Feliformia
mitochondrial DNA (mtDNA) traits of all putative tiger
Family: Felidae
subspecies.[10] In 2017, the Cat Classification Task Force of
the IUCN Cat Specialist Group revised felid taxonomy in
Subfamily: Pantherinae
accordance with the 2015 two-subspecies proposal and
recognised only P. t. tigris and P. t. sondaica.[12] Results of a Genus: Panthera

2018 whole-genome sequencing study of 32 samples from

Species: P. tigris
the six living putative subspecies—the Bengal, Malayan,
Indochinese, South China, Siberian and Sumatran tiger—found Binomial name
them to be distinct and separate clades.[13] These results were
Panthera tigris
corroborated in 2021 and 2023.[14][15] The Cat Specialist
(Linnaeus, 1758)[2]
Group states that "Given the varied interpretations of data, the
Subspecies
[subspecific] taxonomy of this species is currently under P. t. tigris
[16]
review by the IUCN SSC Cat Specialist Group."
P. t. sondaica
The following tables are based on the classification of the
†P. t. acutidens
tiger as of 2005,[9] and also reflect the classification
recognised by the Cat Classification Task Force in 2017.[12] †P. t. soloensis

†P. t. trinilensis

Tiger distribution as of 2022

Synonyms[3]

Felis tigris Linnaeus, 1758

Tigris striatus Severtzov, 1858

Tigris regalis Gray, 1867

Panthera tigris tigris (Linnaeus, 1758)[2]

Population Description Image

Bengal tiger formerly This population inhabits the Indian subcontinent.[17] The Bengal tiger has shorter fur than tigers
P. t. tigris (Linnaeus, further north,[8] with a light tawny to orange-red colouration,[8][18] and relatively long and narrow
[2]
1758) nostrils.[19]

†Caspian tiger This population occurred from Turkey to around the Caspian Sea.[17] It had bright rusty-red fur
formerly P. t. virgata with thin and closely spaced brownish stripes,[21] and a broad occipital bone.[11] Genetic analysis
[20]
(Illiger, 1815) revealed that it was closely related to the Siberian tiger.[22] It has been extinct since the 1970s.[23]

Siberian tiger This population lives in the Russian Far East, Northeast China and possibly North Korea.[17] The
formerly P. t. altaica Siberian tiger has long hair and dense fur.[24] Its ground colour varies widely from ochre-yellow in
[24]
(Temminck, 1844) winter to more reddish and vibrant after moulting.[25] The skull is shorter and broader than the
skulls of tigers further south.[19]

South China tiger This tiger historically lived in south-central China.[17] The skulls of the five type specimens had
formerly P. t. shorter carnassials and molars than tigers from India, a smaller cranium, orbits set closer
[26]
amoyensis (Hilzheimer, together and larger postorbital processes; skins were yellowish with rhombus-like stripes. It
[26] [12][27][28]
1905) has a unique mtDNA haplotype due to interbreeding with ancient tiger lineages. It is
[1]
extinct in the wild as there has not been a confirmed sighting since the 1970s, and survives only
in captivity.[15]

Indochinese tiger This tiger population occurs on the Indochinese Peninsula.[17] Indochinese tiger specimens are
formerly P. t. corbetti smaller with smaller skulls than specimens from India and appear to have darker fur with slightly
[29]
(Mazák, 1968) narrower stripes.[29][30]

Malayan tiger The Malayan tiger was proposed as a distinct subspecies on the basis of mtDNA and micro-
formerly P. t. jacksoni satellite sequences that differ from the Indochinese tiger.[31] It does not differ significantly in fur
[31]
(Luo et al., 2004) colour or skull size from Indochinese tigers.[30] There is no clear geographical barrier between
tiger populations in northern Malaysia and southern Thailand.[1]

Panthera tigris sondaica (Temminck, 1844)[12]

Population Description Image

†Javan tiger formerly This tiger was described based on an unspecified number of skins with short and smooth
P. t. sondaica hair.[24] Tigers from Java were small compared to tigers of the Asian mainland, had relatively
[24]
(Temminck, 1944) elongated skulls compared to the Sumatran tiger and longer, thinner and more numerous
stripes.[30] The Javan tiger is thought to have gone extinct by the 1980s.[23]

†Bali tiger formerly P. This tiger occurred on Bali and had brighter fur and a smaller skull than the Javan tiger.[32][33] A
t. balica (Schwarz, typical feature of Bali tiger skulls is the narrow occipital bone, which is similar to the Javan
[32]
1912) tiger's skull.[34] The tiger went extinct in the 1940s.[23]

Sumatran tiger The type specimen from Sumatra had dark fur.[35] The Sumatran tiger has particularly long hair
formerly P. t. sumatrae around the face,[17] thick body stripes,[30] and a broader and smaller nasal bone than other
[35]
(Pocock, 1929) island tigers.[30][19]

Evolution

The tiger shares the genus Panthera with the lion, leopard, jaguar and snow leopard. Results of genetic
analyses indicate that the tiger and snow leopard are sister species whose lineages split from each other
between 2.70 and 3.70 million years ago.[37] The tiger's whole genome
Snow leopard
sequencing shows repeated sequences that parallel those in other cat
genomes.[38]
Tiger
The fossil species Panthera palaeosinensis of early Pleistocene northern
China was described as a possible tiger ancestor when it was discovered
Jaguar
in 1924, but modern cladistics places it as basal to modern Panthera

Panthera.[39][40] Panthera zdanskyi, which lived around the same time and
Leopard
place, was suggested to be a sister species of the modern tiger when it
was examined in 2014.[40] However, as of 2023, at least two subsequent
studies considered P. zdanskyi likely to be a synonym of P. palaeosinensis, Lion

noting that its proposed differences from that species fell within the
Phylogeny of the Panthera based on
range of individual variation.[41][42] The earliest appearance of the modern
nuclear DNA[36]
tiger species in the fossil record are jaw fragments from Lantion in China
that are dated to the early Pleistocene.[40]

Middle- to late-Pleistocene tiger fossils have been found throughout China, Sumatra and Java. Prehistoric
subspecies include Panthera tigris trinilensis and P. t. soloensis of Java and Sumatra and P. t. acutidens of
China; late Pleistocene and early Holocene fossils of tigers have also been found in Borneo and Palawan,
Philippines.[43] Fossil specimens of tigers have also been reported from the Middle-Late Pleistocene of
Japan.[44] Results of a phylogeographic study indicate that all living tigers have a common ancestor that lived
between 108,000 and 72,000 years ago.[31] Genetic studies suggest that the tiger population contracted
around 115,000 years ago due to glaciation. Modern tiger populations originated from a refugium in Indochina
and spread across Asia after the Last Glacial Maximum. As they colonised northeastern China, the ancestors
of the South China tiger intermixed with a relict tiger population.[27][28]

Hybrids

Captive tigers have been bred with lions to create hybrids that share physical and behavioural qualities of both
parent species; the liger is the offspring of a female tiger and a male lion and the tigon the offspring of a male
tiger and a female lion.[45] The lion sire passes on a growth-promoting gene, but the corresponding growth-
inhibiting gene from the female tiger is absent, so that ligers grow far larger than either parent species. By
contrast, the male tiger does not pass on a growth-promoting gene while the lioness passes on a growth
inhibiting gene; hence, tigons are around the same size as their parents.[46] Since they often develop life
threatening birth defects and can easily become obese, breeding these hybrids is regarded as unethical.[45]
Characteristics

Tiger skeleton from Royal Natural

History Volume 1 (1839)

The tiger has a typical felid morphology, with a muscular body, shortened legs, strong forelimbs with wide front
paws, a large head and a tail that is about half the length of the rest of its body.[47][48] It has five digits,
including a dewclaw, on the front feet and four on the back, all of which have retractile claws that are compact
and curved, and can reach 10 cm (3.9 in) long.[47][49] The ears are rounded and the eyes have a round pupil.[47]
The nose ends in a triangular, pink tip with small black dots, the number of which increase with age.[50] The
tiger's skull is robust, with a constricted front region, proportionally small, elliptical orbits, long nasal bones, a
lengthened cranium with a large sagittal crest.[51][47] It resembles a lion's skull, but differs from it in the
concave or flattened (as opposed to convex) underside of the lower jaw and in its longer nasals.[51][43] The
tiger has 30 fairly robust teeth and its somewhat curved canines are the longest in the cat family at 6.4–
7.6 cm (2.5–3.0 in).[47][52]

The tiger is considered to be the largest living felid species. However, there is some debate over averages
compared to the lion. Since tiger populations vary greatly in size, the "average" size for a tiger may be less than
a lion, while the biggest tigers are bigger than their lion counterparts.[43] The tiger has a head-body length of
1.4–2.8 m (4.6–9.2 ft) with a 0.6–1.1 m (2.0–3.6 ft) tail and stands 0.8–1.1 m (2.6–3.6 ft) at the shoulder.[53]
The Siberian and Bengal tigers are the largest.[47] Male Bengal tigers weigh 200–260 kg (440–570 lb) and
females weigh 100–160 kg (220–350 lb). Island tigers are the smallest, likely due to insular dwarfism.[11] The
Sumatran tiger has a weight of 100–140 kg (220–310 lb) for males and 75–110 kg (165–243 lb) for
females.[54] The extinct Bali tiger was even smaller.[47]

Coat

Tiger coat
The tiger's coat usually has short hairs, reaching up to 35 mm (1.4 in), though the hairs of the northern-living
Siberian tiger can reach 105 mm (4.1 in). Belly hairs tend to be longer than back hairs. The density of their fur
is usually thin, though the Siberian tiger develops a particularly thick winter coat. The tiger has lines of fur
around the face and long whiskers, especially in males.[47] It has an orange colouration that varies from
yellowish to reddish.[55] White fur covers the underside, from head to tail, along with the inner surface of the
legs and parts of the face.[47][56] On the back of the ears, it has a prominent white spot, which is surrounded by
black.[47] The tiger is marked with distinctive black or dark brown stripes, which are uniquely shaped in each
individual.[47][57] The stripes are mostly vertical, but those on the limbs and forehead are horizontal. They are
more concentrated towards the back and those on the trunk may reach under the belly. The tips of stripes are
generally sharp and some may split up or split and fuse again. Tail stripes are thick bands and a black tip
marks the end.[56]

The tiger is one of only a few striped cat species.[58] Stripes are advantageous for camouflage in vegetation
with vertical patterns of light and shade, such as trees, reeds and tall grass.[57][59] This is supported by a
Fourier analysis study showing that the striping patterns line up with their environment.[60] The orange colour
may also aid in concealment, as the tiger's prey is colour blind and possibly perceives the tiger as green and
blended in with the vegetation.[61]

Colour variations

Pseudo-melanistic white tiger

The three colour variants – nearly stripeless snow-white, white and golden – are now virtually non-existent in
the wild due to the reduction of wild tiger populations but continue in captive populations. The white tiger has
a white background colour with sepia-brown stripes. The golden tiger is pale golden with reddish-brown
stripes. The snow-white tiger is a morph with extremely faint stripes and a pale reddish-brown ringed tail.
White and golden morphs are the result of an autosomal recessive trait with a white locus and a wideband
locus, respectively. The snow-white variation is caused by polygenes with both white and wideband loci.[62] The
breeding of white tigers is controversial, as they have no use for conservation. Only 0.001% of wild tigers have
the genes for this colour morph and the overrepresentation of white tigers in captivity is the result of
inbreeding. Hence, their continued breeding will risk both inbreeding depression and loss of genetic variability
in captive tigers.[63]

Pseudo-melanistic tigers with thick, merged stripes have been recorded in Simlipal National Park and three
Indian zoos; a population genetic analysis of Indian tiger samples revealed that this phenotype is caused by a
mutation of a transmembrane aminopeptidase gene. Around 37% of the Simlipal tiger population has this
feature, which has been linked to genetic isolation.[64]

Distribution and habitat

Camera trap of Siberian tiger in

Russia

The tiger historically ranged from eastern Turkey, northern Iran and Afghanistan to Central Asia and from
northern Pakistan through the Indian subcontinent and Indochina to southeastern Siberia, Sumatra, Java and
Bali.[47] As of 2022, it inhabits less than 7% of its historical distribution and has a scattered range in the Indian
subcontinent, the Indochinese Peninsula, Sumatra, northeastern China and the Russian Far East.[1] As of 2020,
India had the largest extent of global tiger habitat with 300,508 km2 (116,027 sq mi), followed by Russia with
195,819 km2 (75,606 sq mi).[65]

The tiger mainly lives in forest habitats and is highly adaptable.[54] Records in Central Asia indicate that it
primarily inhabited Tugay riverine forests and hilly and lowland forests in the Caucasus.[66] In the Amur-Ussuri
region of Russia and China, it inhabits Korean pine and temperate broadleaf and mixed forests; riparian forests
serve as dispersal corridors, providing food and water for both tigers and ungulates.[67] On the Indian
subcontinent, it inhabits mainly tropical and subtropical moist broadleaf forests, moist evergreen forests,
tropical dry forests, alluvial plains and the mangrove forests of the Sundarbans.[68] In the Eastern Himalayas, it
was documented in temperate forest up to an elevation of 4,200 m (13,800 ft) in Bhutan, of 3,630 m (11,910 ft)
in the Mishmi Hills and of 3,139 m (10,299 ft) in Mêdog County, southeastern Tibet.[69][70][71] In Thailand, it lives
in deciduous and evergreen forests.[72] In Sumatra, it inhabits lowland peat swamp forests and rugged
montane forests.[73]

Population density

Camera trapping during 2010–2015 in the deciduous and subtropical pine forest of Jim Corbett National Park,
northern India revealed a stable tiger population density of 12–17 individuals per 100 km2 (39 sq mi) in an area
of 521 km2 (201 sq mi).[74] In northern Myanmar, the population density in a sampled area of roughly
3,250 km2 (1,250 sq mi) in a mosaic of tropical broadleaf forest and grassland was estimated to be 0.21–0.44
tigers per 100 km2 (39 sq mi) as of 2009.[75] Population density in mixed deciduous and semi-evergreen
forests of Thailand's Huai Kha Khaeng Wildlife Sanctuary was estimated at 2.01 tigers per 100 km2 (39 sq mi);
during the 1970s and 1980s, logging and poaching had occurred in the adjacent Mae Wong and Khlong Lan
National Parks, where population density was much lower, estimated at only 0.359 tigers per 100 km2
(39 sq mi) as of 2016.[76] Population density in dipterocarp and montane forests in northern Malaysia was
estimated at 1.47–2.43 adult tigers per 100 km2 (39 sq mi) in Royal Belum State Park, but 0.3–0.92 adult
tigers per 100 km2 (39 sq mi) in the unprotected selectively logged Temengor Forest Reserve.[77]

Behaviour and ecology

Tiger bathing

Camera trap data show that tigers in Chitwan National Park avoided locations frequented by people and were
more active at night than during day.[78] In Sundarbans National Park, six radio-collared tigers were most active
in the early morning with a peak around dawn.[79] A three-year-long camera trap survey in Shuklaphanta
National Park revealed that tigers were most active from dusk until midnight.[80] In northeastern China, tigers
were crepuscular and active at night with activity peaking at dawn and dusk; they were largely active at the
same time as ungulate species.[81]

The tiger is a powerful swimmer and easily transverses rivers as wide as 8 km (5.0 mi); it immerses in water,
particularly on hot days.[57] In general, it is less capable of climbing trees than many other cats due to its size,
but cubs under 16 months old may routinely do so.[82] An adult was recorded climbing 10 m (33 ft) up a
smooth pipal tree.[47]

Social spacing

Adult tigers lead largely solitary lives within home ranges or territories, the size of which mainly depends on
prey abundance, geographic area and sex of the individual. Males and females defend their home ranges from
those of the same sex and the home range of a male encompasses that of multiple females.[47][57] Two
females in the Sundarbans had home ranges of 10.6 and 14.1 km2 (4.1 and 5.4 sq mi).[83] In Panna Tiger
Reserve, the home ranges of five reintroduced females varied from 53–67 km2 (20–26 sq mi) in winter to 55–
60 km2 (21–23 sq mi) in summer and to 46–94 km2 (18–36 sq mi) during the monsoon; three males had 84–
147 km2 (32–57 sq mi) large home ranges in winter, 82–98 km2 (32–38 sq mi) in summer and 81–118 km2
(31–46 sq mi) during monsoon seasons.[84] In Sikhote-Alin Biosphere Reserve, 14 females had home ranges
248–520 km2 (96–201 sq mi) and five resident males of 847–1,923 km2 (327–742 sq mi) that overlapped with
those of up to five females.[85] When tigresses in the same reserve had cubs of up to four months of age, they
reduced their home ranges and steadily enlarged them until their offspring were 13–18 months old.[86]

Tiger rubbing against tree to mark

territory

The tiger is a long-ranging species and individuals disperse over distances of up to 650 km (400 mi) to reach
tiger populations in other areas.[87] Young tigresses establish their first home ranges close to their mother's
while males migrate further than their female counterparts.[88] Four radio-collared females in Chitwan
dispersed between 0 and 43.2 km (0.0 and 26.8 mi) and 10 males between 9.5 and 65.7 km (5.9 and
40.8 mi).[89] A subadult male lives as a transient in another male's home range until he is older and strong
enough to challenge the resident male.[88][90] Tigers mark their home ranges by spraying urine on vegetation
and rocks, clawing or scent rubbing trees and marking trails with faeces, anal gland secretions and ground
scrapings.[57][91][92][93] Scent markings also allow an individual to pick up information on another's identity.
Unclaimed home ranges, particularly those that belonged to a deceased individual, can be taken over in days
or weeks.[57]

Male tigers are generally less tolerant of other males within their home ranges than females are of other
females. Disputes are usually solved by intimidation rather than fighting. Once dominance has been
established, a male may tolerate a subordinate within his range, as long as they do not come near him. The
most serious disputes tend to occur between two males competing for a female in oestrus.[94] Though tigers
mostly live alone, relationships between individuals can be complex. Tigers are particularly social at kills and a
male tiger will share a carcass with the females and cubs within this home range and unlike male lions, will
allow them to feed on the kill before he is finished with it. However, a female is more tense when encountering
another female at a kill.[95][96]
Communication

Tiger baring teeth as a

sign of aggression

Captive tiger roaring

During friendly encounters and bonding, tigers rub against each other's bodies.[97] Facial expressions include
the "defence threat", which involves a wrinkled face, bared teeth, pulled-back ears and widened pupils.[98][47]
Both males and females show a flehmen response, a characteristic grimace, when smelling urine markings.
Males also use the flehmen to detect the markings made by tigresses in oestrus.[47] Tigers will move their ears
around to display the white spots, particularly during aggressive encounters and between mothers and
cubs.[99] They also use their tails to signal their mood. To show cordiality, the tail sticks up and sways slowly,
while an apprehensive tiger lowers its tail or wags it side-to-side. When calm, the tail hangs low.[100]

Tigers are normally silent but can produce numerous vocalisations.[101][102] They roar to signal their presence
to other individuals over long distances. This vocalisation is forced through an open mouth as it closes and
can be heard 3 km (1.9 mi) away. They roar multiple times in a row and others respond in kind. Tigers also roar
during mating and a mother will roar to call her cubs to her. When tense, tigers moan, a sound similar to a roar
but softer and made when the mouth is at least partially closed. Moaning can be heard 400 m (1,300 ft)
away.[47][103] Aggressive encounters involve growling, snarling and hissing.[104] An explosive "coughing roar" or
"coughing snarl" is emitted through an open mouth and exposed teeth.[47][104][99] In friendlier situations, tigers
prusten, a soft, low-frequency snorting sound similar to purring in smaller cats.[105] Tiger mothers
communicate with their cubs by grunting, while cubs call back with miaows.[106] When startled they "woof".
They produce a deer-like "pok" sound for unknown reasons, but most often at kills.[107][108]
Hunting and diet

Tiger attacking a sambar deer in

Ranthambore Tiger Reserve

The tiger is a carnivore and an apex predator feeding mainly on ungulates, with a particular preference for
sambar deer, Manchurian wapiti, barasingha and wild boar. Tigers kill large ungulates like gaur[109] and
opportunistically smaller prey like monkeys, peafowl and other ground-based birds, porcupines and fish.[47][57]
Occasional attacks on Asian elephants and Indian rhinoceroses have also been reported.[110] More often,
tigers take the more vulnerable calves.[111] They sometimes prey on livestock and dogs in close proximity to
settlements.[47] Tigers occasionally consume vegetation, fruit and minerals for dietary fibre and
supplements.[112]

Tigers learn to hunt from their mothers, which is important but not essential for their success.[113] Depending
on the size of the prey, they typically kill weekly though mothers must kill more often.[54] They usually hunt
alone, but families hunt together when cubs are old enough.[114] They search for prey using vision and
hearing.[115] A tiger will also wait at a watering hole for prey to come by, particularly during hot summer
days.[116][117] It is an ambush predator and when approaching potential prey, it crouches with the head lowered
and hides in foliage. It switches between creeping forward and staying still. A tiger may even doze off and can
stay in the same spot for as long as a day, waiting for prey and launch an attack when the prey is close
enough,[118] usually within 30 m (98 ft).[54] If the prey spots it before then, the cat does not pursue further.[116] A
tiger can sprint 56 km/h (35 mph) and leap 10 m (33 ft);[119][120] it is not a long-distance runner and gives up a
chase if prey outpaces it over a certain distance.[116]

Two tigers attacking a wild boar

in Kanha Tiger Reserve

The tiger attacks from behind or at the sides and tries to knock the target off balance. It latches onto prey with
its forelimbs, twisting and turning during the struggle and tries to pull it to the ground. The tiger generally
applies a bite to the throat until its victim dies of strangulation.[47][121][122][123] It has an average bite force at the
canine tips of 1234.3 newtons.[124] Holding onto the throat puts the cat out of reach of horns, antlers, tusks
and hooves.[121][125] Tigers are adaptable killers and may use other methods, including ripping the throat or
breaking the neck. Large prey may be disabled by a bite to the back of the hock, severing the tendon. Swipes
from the large paws are capable of stunning or breaking the skull of a water buffalo.[126] They kill small prey
with a bite to the back of the neck or head.[127][54] Estimates of the success rate for hunting tigers range from a
low of 5% to a high of 50%. They are sometimes killed or injured by large or dangerous prey like gaur, buffalo
and boar.[54]

Tigers typically move kills to a private, usually vegetated spot no further than 183 m (600 ft), though they have
been recorded dragging them 549 m (1,801 ft). They are strong enough to drag the carcass of a fully grown
buffalo for some distance. They rest for a while before eating and can consume as much as 50 kg (110 lb) of
meat in one session, but feed on a carcass for several days, leaving little for scavengers.[128]

Predatory competitors

An 1807 illustration of dholes

attacking a tiger

In much of their range, tigers share habitat with leopards and dholes. They typically dominate both of them,
though with dholes it depends on their pack size. Interactions between them involve chasing, stealing kills and
direct killing.[129] Large dhole packs may kill tigers.[130] In Jigme Dorji National Park, tigers were found to
inhabit the deeper parts of forests while these smaller predators were pushed closer to the fringes.[131] The
three predators coexist by hunting different sized prey.[132] In Nagarhole National Park, the average weight for
tiger kills was found to be 91.5 kg (202 lb), compared to 37.6 kg (83 lb) for leopards and 43.4 kg (96 lb) for
dholes.[133]

Both leopards and dholes can live successfully in tiger habitat when there is abundant food and vegetation
cover.[132][134] Nevertheless, they appear to be less common where tigers are numerous. The recovery of the
tiger population in Rajaji National Park during the 2000s led to a reduction in leopard population densities.[135]
Similarly, at two sites in central India the size of dhole packs was negatively correlated with tiger densities.[136]
Leopard and dhole distribution in Kui Buri National Park correlated with both prey access and tiger
absence.[137]
Reproduction and life cycle

A tiger family in Kanha Tiger Reserve

The tiger generally mates all year round, particularly between November and April. A tigress is in oestrus for
three to six days at a time, separated by three to nine week intervals.[47] A resident male mates with all the
females within his home range, who signal their receptiveness by roaring and marking.[138][139] Younger,
transient males are also attracted, leading to a fight in which the more dominant male drives the usurper
off.[140][138] During courtship, the male is cautious with the female as he waits for her to show signs she is
ready to mate. She signals to him by positioning herself in lordosis with her tail to the side. Copulation is
generally 20 to 25 seconds long, with the male biting the female by the scruff of her neck. After it is finished,
the male quickly pulls away as the female may turn and slap him.[138] Tiger pairs may stay together for up to
four days and mate multiple times.[141] Gestation lasts around or over three months.[47]

A tigress gives birth in a secluded location, be it in dense vegetation, in a cave or under a rocky shelter.[142]
Litters consist of as many as seven cubs, but two or three are more typical.[140][142] Newborn cubs weigh 785–
1,610 g (27.7–56.8 oz) and are blind and altricial.[142] The mother licks and cleans her cubs, suckles them and
viciously defends them from any potential threat.[140] Cubs open their eyes at the age of three to 14 days and
their vision becomes clear after a few weeks.[142] They can leave the denning site after two months and around
the same time they start eating meat.[140][143] The mother only leaves them alone to hunt and even then she
does not travel far.[144] When she suspects an area is no longer safe, she moves her cubs to a new spot,
transporting them one by one by grabbing them by the scruff of the neck with her mouth.[145][146] A tigress in
Sikhote-Alin Biosphere Reserve maximised the time spent with her cubs by reducing her home range, killing
larger prey and returning to her den more rapidly than without cubs; when the cubs started to eat meat, she
took them to kill sites, thereby optimising their protection and access to food.[147] In the same reserve, one of
21 cubs died in over eight years of monitoring and mortality did not differ between male and female
juveniles.[148] Tiger monitoring over six years in Ranthambore Tiger Reserve indicated an average annual
survival rate of around 85 percent for 74 male and female cubs; survival rate increased to 97 percent for both
males and female juveniles of one to two years of age.[149] Causes of cub mortality include predators, floods,
fires, death of the mother and fatal injuries.[145][146][150][149]
 A tigress with her cub at Buffalo Zoo

After around two months, the cubs are able to follow their mother. They still hide in vegetation when she goes
hunting. Young bond through play fighting and practice stalking. A hierarchy develops in the litter, with the
biggest cub, often a male, being the most dominant and the first to eat its fill at a kill.[151] Around the age of six
months, cubs are fully weaned and have more freedom to explore their environment. Between eight and ten
months, they accompany their mother on hunts.[152] A cub can make a kill as early as 11 months and reach
independence as a juvenile of 18 to 24 months of age; males become independent earlier than
females.[153][149] Radio-collared tigers in Chitwan started leaving their natal areas at the age of 19 months.[89]
Young females are sexually mature at three to four years, whereas males are at four to five years.[47]
Generation length of the tiger is about 7–10 years.[154] Wild Bengal tigers live 12–15 years.[155] Data from the
International Tiger Studbook 1938–2018 indicate that captive tigers lived up to 19 years.[156]

The father does not play a role in raising the young, but he encounters and interacts with them. The resident
male appears to visit the female–cub families within his home range. They socialise and even share
kills.[157][158] One male was recorded looking after orphaned cubs whose mother had died.[159] By defending his
home range, the male protects the females and cubs from other males.[160] When a new male takes over,
dependent cubs are at risk of infanticide as the male attempts to sire his own young with the females.[161] A
seven-year long study in Chitwan National Park revealed that 12 of 56 detected cubs and juveniles were killed
by new males taking over home ranges.[150]

Health and diseases

Tigers are recorded as hosts for various parasites including tapeworms like Diphyllobothrium erinacei, Taenia
pisiformis in India and nematodes like Toxocara species in India and Physaloptera preputialis, Dirofilaria ursi and
Uiteinarta species in Siberia.[47] Canine distemper is known to occur in Siberian tigers.[162] A morbillivirus
infection was the likely cause of death of a tigress in the Russian Far East that was also tested positive for
feline panleukopenia and feline coronavirus.[163] Blood samples from 11 adult tigers in Nepal showed
antibodies for canine parvovirus-2, feline herpesvirus, feline coronavirus, leptospirosis and Toxoplasma
gondii.[164]

Threats

The tiger has been listed as Endangered on the IUCN Red List since 1986 and the global tiger population is
thought to have continuously declined from an estimated population of 5,000–8,262 tigers in the late 1990s to
3,726–5,578 individuals estimated as of 2022.[1] During 2001–2020, landscapes where tigers live declined
from 1,025,488 km2 (395,943 sq mi) to 911,901 km2 (352,087 sq mi).[65] Habitat destruction, habitat
fragmentation and poaching for fur and body parts are the major threats that contributed to the decrease of
tiger populations in all range countries.[1]

Protected areas in central India are highly fragmented due to linear infrastructure like roads, railway lines,
transmission lines, irrigation channels and mining activities in their vicinity.[165] In the Tanintharyi Region of
southern Myanmar, deforestation coupled with mining activities and high hunting pressure threatens the tiger
population in the area.[166] In Thailand, nine of 15 protected areas hosting tigers are isolated and fragmented,
offering a low probability for dispersal between them; four of these have no longer harboured tigers since at
least 2013.[167] In Peninsular Malaysia, 8,315.7 km2 (3,210.7 sq mi) of tiger habitat was cleared during 1988–
2012, most of it for industrial plantations.[168] Large-scale land acquisitions of about 23,000 km2 (8,900 sq mi)
for commercial agriculture and timber extraction in Cambodia contributed to the fragmentation of potential
tiger habitat, especially in the Eastern Plains.[169] Inbreeding depression coupled with habitat destruction,
insufficient prey resources and poaching is a threat to the small and isolated tiger population in the Changbai
Mountains along the China–Russia border.[170] In China, tigers became the target of large-scale 'anti-pest'
campaigns in the early 1950s, where suitable habitats were fragmented following deforestation and
resettlement of people to rural areas, who hunted tigers and prey species. Though tiger hunting was prohibited
in 1977, the population continued to decline and is considered extinct in South China since 2001.[171][172]

A Javan tiger skin on display

Tiger populations in India have been targeted by poachers since the 1990s and were extirpated in two tiger
reserves in 2005 and 2009.[173] Between March 2017 and January 2020, 630 activities of hunters using snares,
drift nets, hunting platforms and hunting dogs were discovered in a reserve forest of about 1,000 km2
(390 sq mi) in southern Myanmar.[174] Nam Et-Phou Louey National Park was considered the last important
site for the tiger in Laos, but it has not been recorded there at least since 2013; this population likely fell victim
to indiscriminate snaring.[175] Anti-poaching units in Sumatra's Kerinci Seblat landscape removed 362 tiger
snare traps and seized 91 tiger skins during 2005–2016; annual poaching rates increased with rising skin
prices.[176] Poaching is also the main threat to the tiger population in far eastern Russia, where logging roads
facilitate access for poachers and people harvesting forest products that are vital for prey species to survive in
winter.[177]

Body parts of 207 tigers were detected during 21 surveys in 1991–2014 in two wildlife markets in Myanmar
catering to customers in Thailand and China.[178] During the years 2000–2022, at least 3,377 tigers were
confiscated in 2,205 seizures in 28 countries; seizures encompassed 665 live and 654 dead individuals, 1,313
whole tiger skins, 16,214 body parts like bones, teeth, paws, claws, whiskers and 1.1 t (1.1 long tons; 1.2 short
tons) of meat; 759 seizures in India encompassed body parts of 893 tigers; and 403 seizures in Thailand
involved mostly captive-bred tigers.[179] Seizures in Nepal between January 2011 and December 2015 obtained
585 pieces of tiger body parts and two whole carcasses in 19 districts.[180] Seizure data from India during
2001–2021 indicate that tiger skins were the most often traded body parts, followed by claws, bones and
teeth; trafficking routes mainly passed through the states of Maharashtra, Karnataka, Tamil Nadu and
Assam.[181] A total of 292 illegal tiger parts were confiscated at US ports of entry from personal baggage, air
cargo and mail between 2003 and 2012.[182]

Demand for tiger parts for use in traditional Chinese medicine has also been cited as a major threat to tiger
populations.[183] Interviews with local people in the Bangladeshi Sundarbans revealed that they kill tigers for
local consumption and trade of skins, bones and meat, in retaliation for attacks by tigers and for
excitement.[184] Tiger body parts like skins, bones, teeth and hair are consumed locally by wealthy
Bangladeshis and are illegally trafficked from Bangladesh to 15 countries including India, China, Malaysia,
Korea, Vietnam, Cambodia, Japan and the United Kingdom via land borders, airports and seaports.[185] Tiger
bone glue is the prevailing tiger product purchased for medicinal purposes in Hanoi and Ho Chi Minh City.[186]
"Tiger farm" facilities in China and Southeast Asia breed tigers for their parts, but these appear to make the
threat to wild populations worse by increasing the demand for tiger products.[187]

Local people killing tigers in retaliation for attacking and preying on livestock is a threat in several tiger range
countries, as this consequence of human–wildlife conflict also contributes to the decline of the
population.[188][189][190][191][192]

Conservation

Internationally, the tiger is protected under CITES Appendix I, banning trade of live tigers and their body parts.[1]
In Russia, hunting the tiger has been banned since 1952.[197] In Bhutan, it has been protected since 1969 and
enlisted as totally protected since 1995.[198] Since 1972, it has been afforded the highest protection level under
India's Wild Life (Protection) Act, 1972.[199] In Nepal and Bangladesh, it has been protected since 1973.[199][185]
Since 1976, it has been totally protected under Malaysia's Protection of Wild Life Act,[200] and the country's
Wildlife Conservation Act enacted in 2010 increased punishments for wildlife-related crimes.[194] In Indonesia,
it has been protected since 1990.[201] In China, the trade in tiger body parts was banned in 1993.[202] The Thai
Wildlife Preservation and Protection Act was enacted in 2019 to combat poaching and trading of body
parts.[203]
In 1973, the National Tiger Conservation Authority and Project Tiger were Global wild tiger population
founded in India to gain public support for tiger conservation.[173] Since Country Year Estimate
2
then, 53 tiger reserves covering an area of 75,796 km (29,265 sq mi) India 2022 3,167–3,682[193]
have been established in the country up to 2022.[193] Myanmar's national
Russia 2022 573–600[194]
tiger conservation strategy developed in 2003 comprises management
Indonesia 2022 393[194]
tasks such as restoration of degraded habitats, increasing the extent of
Nepal 2022 316–355[195]
protected areas and wildlife corridors, protecting tiger prey species,
thwarting tiger killing and illegal trade of its body parts and promoting Thailand 2022 148–189[194]

public awareness through wildlife education programmes.[204] Bhutan's Malaysia 2022 <150[194]

first Tiger Action Plan implemented during 2006–2015 revolved around Bhutan 2022 131[196]

habitat conservation, human–wildlife conflict management, education Bangladesh 2018 114[194]

and awareness; the second Action Plan aimed at increasing the China 2022 >60[194]

country's tiger population by 20% until 2023 compared to 2015.[198] In Myanmar 2022 28[194]
2009, the Bangladesh Tiger Action Plan was initiated to stabilise the Total 5,634–5,891
country's tiger population, maintain habitat and a sufficient prey base,
improve law enforcement and foster cooperation between governmental agencies responsible for tiger
conservation.[205] The Thailand Tiger Action Plan ratified in 2010 envisioned increasing the country's tiger
populations by 50% in the Western Forest Complex and Dong Phayayen–Khao Yai Forest Complex and
reestablish populations in three potential landscapes until 2022.[206] The Indonesian National Tiger Recovery
Program ratified in 2010 aimed at increasing the Sumatran tiger population by 2022.[207] The third strategic
and action plan for the conservation of the Sumatran tiger for the years 2020–2030 revolves around
strengthening management of small tiger population units of less than 20 mature individuals and connectivity
between 13 forest patches in North Sumatra and West Sumatra provinces.[208]

Wild Sumatran tiger caught by camera

trap

Increases in anti-poaching patrol efforts in four Russian protected areas during 2011–2014 contributed to
reducing poaching, stabilising the tiger population and improving protection of ungulate populations.[209]
Poaching and trafficking were declared to be moderate and serious crimes in 2019.[194] Anti-poaching
operations were also established in Nepal in 2010, with increased cooperation and intelligence sharing
between agencies. These policies have led to many years of "zero poaching" and the country's tiger population
has doubled in a decade.[194] Anti-poaching patrols in the 1,200 km2 (460 sq mi) large core area of Taman
Negara lead to a decrease of poaching frequency from 34 detected incidents in 2015–2016 to 20 incidents
during 2018–2019; the arrest of seven poaching teams and removal of snares facilitated the survival of three
resident female tigers and at least 11 cubs.[210] Army and police officer are deployed for patrolling together
with staff of protected areas in Malaysia.[194]

Wildlife corridors are important conservation measures as they facilitate tiger populations to connect between
protected areas; tigers use at least nine corridors that were established in the Terai Arc Landscape and Sivalik
Hills in both Nepal and India.[211] Corridors in forested areas with low human encroachment are highly
suitable.[212][213] In West Sumatra, 12 wildlife corridors were identified as high priority for mitigating human–
wildlife conflicts.[214] In 2019, China and Russia signed a memorandum of understanding for transboundary
cooperation between two protected areas, Northeast China Tiger and Leopard National Park and Land of the
Leopard National Park, that includes the creation of wildlife corridors and bilateral monitoring and patrolling
along the Sino-Russian border.[215]

Rescued and rehabilitated problem tigers and orphaned tiger cubs have been released into the wild and
monitored in India, Sumatra and Russia.[84][216][217] In Kazakhstan, habitat restoration and reintroduction of
prey species in Ile-Balkash Nature Reserve have progressed and tiger reintroduction is planned for 2025.[218]
Reintroduction of tigers is considered possible in eastern Cambodia, once management of protected areas is
improved and forest loss stabilized.[219] South China tigers are kept and bred in Chinese zoos, with plans to
reintroduce their offspring into remote protected areas.[15][113] Coordinated breeding programs among zoos
have led to enough genetic diversity in tigers to act as "insurance against extinction in the wild".[220]

Relationship with humans

Tiger hunting on elephant-back in

India, 1808

Hunting

Tigers have been hunted by humans for millennia, as indicated by a painting on the Bhimbetka rock shelters in
India that is dated to 5,000–6,000 years ago. They were hunted throughout their range in Asia, chased on
horseback, elephant-back or even with sled dogs and killed with spears and later firearms. Such hunts were
conducted both by native governments and empires like the Mughal Empire, as well as European colonists.
Tigers were often hunted as trophies and because of their perceived danger.[221] An estimated 80,000 tigers
were killed between 1875 and 1925.[222][223]
Attacks

A Bengal tiger in the Sundarbans

Tigers are said to have directly killed more people than any other wild mammal.[53] In most areas, they avoid
humans, but attacks are a risk wherever people coexist with them.[224][225] Dangerous encounters are more
likely to occur in edge habitats between wild and agricultural areas.[224] Most attacks on humans are defensive,
including protection of young; however, tigers do sometimes see people as prey.[225] Man-eating tigers tend to
be old and disabled.[57] Tigers driven from their home ranges are also at risk of turning to man-eating.[226]

At the beginning of the 20th century, the Champawat Tiger was responsible for over 430 human deaths in
Nepal and India before she was shot by Jim Corbett.[227] This tigress suffered from broken teeth and was
unable to kill normal prey. Modern authors speculate that feeding on meagre human flesh forced the cat to kill
more and more.[228] Tiger attacks were particularly high in Singapore during the mid-19th century, when
plantations expanded into the tiger's habitat.[229] In the 1840s, the number of deaths ranged from 200 to 300
annually.[230]

Tiger attacks in the Sundarbans caused 1,396 human deaths in the period 1935–2006 according to official
records of the Bangladesh Forest Department.[231] Victims of attacks are local villagers who enter the tiger's
domain to collect resources like wood and honey. Fishermen have been particularly common targets. Methods
to counter tiger attacks have included face masks worn backwards, protective clothes, sticks and carefully
stationed electric dummies.[232]

Captivity

Tiger at Big Cat Rescue in 2014

 Publicity photo of animal trainer
Gunther Gebel-Williams with
several of his trained tigers, c.
1969

Tigers have been kept in captivity since ancient times. In ancient Rome, tigers were displayed in
amphitheatres; they were slaughtered in venatio hunts and used to kill criminals.[233] The Mongol ruler Kublai
Khan is reported to have kept tigers in the 13th century. Starting in the Middle Ages, tigers were being kept in
European menageries.[234] Tigers and other exotic animals were mainly used for the entertainment of elites but
from the 19th century onward, they were exhibited more to the public. Tigers were particularly big attractions
and their captive population soared.[235] In 2020, there were over 8,000 captive tigers in Asia, over 5,000 in the
US and no less than 850 in Europe.[236] Captive tigers may display stereotypical behaviours such as pacing or
inactivity. Modern zoos are able to reduce such behaviours with exhibits designed so the animals can move
between separate but connected enclosures.[237] Enrichment items are also important for the cat's welfare and
the stimulation of its natural behaviours.[238]

Tigers have played prominent roles in circuses and other live performances. Ringling Bros included many tiger
tamers in the 20th century including Mabel Stark, who became a big draw and had a long career. She was well
known for being able to control the tigers despite being a small woman; using "manly" tools like whips and
guns. Another trainer was Clyde Beatty, who used chairs, whips and guns to provoke tigers and other beasts
into acting fierce and allowed him to appear courageous. He would perform with as many as 40 tigers and
lions in one act. From the 1960s onward, trainers like Gunther Gebel-Williams would use gentler methods to
control their animals. Sara Houcke was dubbed "the Tiger Whisperer" as she trained the cats to obey her by
whispering to them.[239] Siegfried & Roy became famous for performing with white tigers in Las Vegas. The act
ended in 2003 when a tiger attacked Roy during a performance.[240] In 2009, tigers were the most traded circus
animals.[241] The use of tigers and other animals in shows would eventually decline in many countries due to
pressure from animal rights groups and greater desires from the public to see them in more natural settings.
Several countries restrict or ban such acts.[242]

Tigers have become popular in the exotic pet trade, particularly in the United States[243] where only 6% of the
captive tiger population in 2020 were being housed in zoos and other facilities approved by the Association of
Zoos and Aquariums.[187] Private collectors are thought to be ill-equipped to provide proper care for tigers,
which compromises their welfare. They can also threaten public safety by allowing people to interact with
them.[187] The keeping of tigers and other big cats by private people was banned in the US in 2022.[244] Most
countries in the European Union have banned breeding and keeping tigers outside of licensed zoos and rescue
centres, but some still allow private holdings.[245]
Cultural significance

Tiger-shaped jie (badge of authority)

with gold inlays, from the tomb of
Zhao Mo

The tiger is among the most famous of the charismatic megafauna. Kailash Sankhala has called it "a rare
combination of courage, ferocity and brilliant colour",[140] while Candy d'Sa calls it "fierce and commanding on
the outside, but noble and discerning on the inside". In a 2004 online poll involving more than 50,000 people
from 73 countries, the tiger was voted the world's favourite animal with 21% of the vote, narrowly beating the
dog.[246] Similarly, a 2018 study found the tiger to be the most popular wild animal based on surveys, as well as
appearances on websites of major zoos and posters of some animated movies.[247]

While the lion represented royalty and power in Western culture, the tiger played such a role in various Asian
cultures. In ancient China, the tiger was seen as the "king of the forest" and symbolised the power of the
emperor.[248] In Chinese astrology, the tiger is the third out of 12 symbols in the Chinese zodiac and controls
the period between 15:00 and 17:00 o'clock in the afternoon. The Year of the Tiger is thought to bring "dramatic
and extreme events". The White Tiger is one of the Four Symbols of the Chinese constellations, representing
the west along with the yin and the season of autumn. It is the counterpart to the Azure Dragon, which
conversely symbolises the east, yang and springtime.[249] The tiger is one of the animals displayed on the
Pashupati seal of the Indus Valley civilisation. The big cat was depicted on seals and coins during the Chola
Dynasty of southern India, as it was the official emblem.[250]

The Hindu goddess Durga

riding a tiger. Guler school,
early 18th century
Tigers have had religious and folkloric significance. In Buddhism, the tiger, monkey and deer are the Three
Senseless Creatures, with the tiger symbolising anger.[251] In Hinduism, the tiger is the vehicle of Durga, the
goddess of feminine power and peace, whom the gods created to fight demons. Similarly, in the Greco-Roman
world, the tiger was depicted being ridden by the god Dionysus. In Korean mythology, tigers are messengers of
the Mountain Gods.[252] In both Chinese and Korean culture, tigers are seen as protectors against evil spirits
and their image was used to decorate homes, tombs and articles of clothing.[248][253][254] In the folklore of
Malaysia and Indonesia, "tiger shamans" heal the sick by invoking the big cat. People turning into tigers and
the inverse has also been widespread; in particular weretigers are people who could change into tigers and
back again. The Mnong people of Indochina believed that tigers could shapeshift into humans.[254] Among
some indigenous peoples of Siberia, it was believed that men would seduce women by transforming into
tigers.[248]

William Blake's 1794 poem "The Tyger" portrays the animal as the duality of beauty and ferocity. It is the sister
poem to "The Lamb" in Blake's Songs of Innocence and of Experience and he ponders why God would create
such different creatures. The tiger is featured in the mediaeval Chinese novel Water Margin, where the cat
battles and is slain by the bandit Wu Song, while the tiger Shere Khan in Rudyard Kipling's The Jungle Book
(1894) is the mortal enemy of the human protagonist Mowgli. The image of the friendly tame tiger has also
existed in culture, notably Tigger, the Winnie-the-Pooh character and Tony the Tiger, the Kellogg's cereal
mascot.[255]

See also

List of largest cats

Tiger Temple

References

1. Goodrich, J.; Wibisono, H.; Miquelle, D.; Lynam, A.J; 2. Linnaeus, C. (1758). "Felis tigris" (https://archive.org/str
Sanderson, E.; Chapman, S.; Gray, T.N.E.; Chanchani, P. eam/mobot31753000798865#page/41/mode/2up) .
& Harihar, A. (2022). "Panthera tigris" (https://www.iucn Caroli Linnæi Systema naturæ per regna tria naturæ,
redlist.org/species/15955/214862019) . IUCN Red secundum classes, ordines, genera, species, cum
List of Threatened Species. 2022: characteribus, differentiis, synonymis, locis (in Latin).
e.T15955A214862019. doi:10.2305/IUCN.UK.2022- Vol. Tomus I (decima, reformata ed.). Holmiae:
1.RLTS.T15955A214862019.en (https://doi.org/10.230 Laurentius Salvius. p. 41.
5%2FIUCN.UK.2022-1.RLTS.T15955A214862019.en) .
3. Ellerman, J.R. & Morrison-Scott, T.C.S. (1951).
Retrieved 31 August 2022.
"Panthera tigris, Linnaeus, 1758" (https://archive.org/de
tails/checklistofpalae00elle/page/318/mode/2up) .
Checklist of Palaearctic and Indian mammals 1758 to
1946. London: British Museum. pp. 318–319.
4. Liddell, H. G. & Scott, R. (1940). "τίγρις" (https://www.p 10. Wilting, A.; Courtiol, A.; Christiansen, P.; Niedballa, J.;
erseus.tufts.edu/hopper/text?doc=Perseus%3Atext%3 Scharf, A. K.; Orlando, L.; Balkenhol, N.; Hofer, H.;
A1999.04.0057%3Aentry%3Dti%2Fgris) . A Greek- Kramer-Schadt, S.; Fickel, J. & Kitchener, A. C. (2015).
English Lexicon (Revised and augmented ed.). Oxford: "Planning tiger recovery: Understanding intraspecific
Clarendon Press. Archived (https://web.archive.org/we variation for effective conservation" (https://www.ncbi.
b/20201021200154/http://www.perseus.tufts.edu/hop nlm.nih.gov/pmc/articles/PMC4640610) . Science
per/text?doc=Perseus:text:1999.04.0057:entry=ti/gri Advances. 11 (5): e1400175.
s) from the original on 21 October 2020. Retrieved Bibcode:2015SciA....1E0175W (https://ui.adsabs.harva
21 February 2021. rd.edu/abs/2015SciA....1E0175W) .
doi:10.1126/sciadv.1400175 (https://doi.org/10.1126%
5. Varro, M. T. (1938). "XX. Ferarum vocabula" (https://arc
2Fsciadv.1400175) . PMC 4640610 (https://www.ncb
hive.org/details/onlatinlanguage01varruoft/page/96/m
i.nlm.nih.gov/pmc/articles/PMC4640610) .
ode/2up) [XX. The names of wild beasts]. De lingua
PMID 26601191 (https://pubmed.ncbi.nlm.nih.gov/266
latina [On the Latin language]. Translated by Kent, R. G.
01191) .
London: W. Heinemann. pp. 94–97.
11. Kitchener, A. (1999). "Tiger distribution, phenotypic
6. Thorley, D. (2017). "Naming the tiger in the Early
variation and conservation issues" in Seidensticker,
Modern world". Renaissance Quarterly. 70 (3): 977–
Christie & Jackson 1999, pp. 19–39
1006. doi:10.1086/693884 (https://doi.org/10.1086%2
F693884) . JSTOR 26560471 (https://www.jstor.org/s 12. Kitchener, A. C.; Breitenmoser-Würsten, C.; Eizirik, E.;
table/26560471) . S2CID 165388712 (https://api.sem Gentry, A.; Werdelin, L.; Wilting, A.; Yamaguchi, N.;
anticscholar.org/CorpusID:165388712) . Abramov, A. V.; Christiansen, P.; Driscoll, C.; Duckworth,
J. W.; Johnson, W.; Luo, S.-J.; Meijaard, E.; O’Donoghue,
7. Pocock, R. I. (1929). "Tigers" (https://archive.org/detail
P.; Sanderson, J.; Seymour, K.; Bruford, M.; Groves, C.;
s/journalofbomb33341929bomb/page/n133) .
Hoffmann, M.; Nowell, K.; Timmons, Z. & Tobe, S.
Journal of the Bombay Natural History Society. 33 (3):
(2017). "A revised taxonomy of the Felidae: The final
505–541.
report of the Cat Classification Task Force of the IUCN
8. Pocock, R. I. (1939). "Panthera tigris" (https://archive.or Cat Specialist Group" (https://repository.si.edu/bitstrea
g/stream/PocockMammalia1/pocock1#page/n247/m m/handle/10088/32616/A_revised_Felidae_Taxonomy
ode/2up) . The Fauna of British India, Including Ceylon _CatNews.pdf?sequence=1&isAllowed=y#page=66)
and Burma. Mammalia: Volume 1. London: T. Taylor and (PDF). Cat News (Special Issue 11): 66–68. Archived (h
Francis, Ltd. pp. 197–210. ttps://web.archive.org/web/20200117172708/https://r

9. Wozencraft, W. C. (2005). "Species Panthera tigris" (htt epository.si.edu/bitstream/handle/10088/32616/A_rev

p://www.departments.bucknell.edu/biology/resources/ ised_Felidae_Taxonomy_CatNews.pdf?sequence=1&is

msw3/browse.asp?id=14000259) . In Wilson, D. E.; Allowed=y#page=66) (PDF) from the original on 17

January 2020. Retrieved 27 August 2019.
Reeder, D. M. (eds.). Mammal Species of the World: A
Taxonomic and Geographic Reference (http://www.goog
le.com/books?id=JgAMbNSt8ikC&pg=PA546)
(3rd ed.). Johns Hopkins University Press. p. 546.
ISBN 978-0-8018-8221-0. OCLC 62265494 (https://ww
w.worldcat.org/oclc/62265494) .
13. Liu, Y.-C.; Sun, X.; Driscoll, C.; Miquelle, D. G.; Xu, X.; 17. Nowell, K. & Jackson, P. (1996). "Tiger, Panthera tigris
Martelli, P.; Uphyrkina, O.; Smith, J. L. D.; O’Brien, S. J. & (Linnaeus, 1758)" (https://portals.iucn.org/library/site
Luo, S.-J. (2018). "Genome-wide evolutionary analysis s/library/files/documents/1996-008.pdf#page=80)
of natural history and adaptation in the world's tigers" (PDF). Wild Cats: Status Survey and Conservation Action
(https://doi.org/10.1016%2Fj.cub.2018.09.019) . Plan. Gland, Switzerland: IUCN. pp. 55–65. ISBN 2-
Current Biology. 28 (23): 3840–3849. 8317-0045-0. Archived (https://web.archive.org/web/2
Bibcode:2018CBio...28E3840L (https://ui.adsabs.harva 0240125121859/https://portals.iucn.org/library/sites/l
rd.edu/abs/2018CBio...28E3840L) . ibrary/files/documents/1996-008.pdf#page=80)
doi:10.1016/j.cub.2018.09.019 (https://doi.org/10.101 (PDF) from the original on 25 January 2024. Retrieved
6%2Fj.cub.2018.09.019) . PMID 30482605 (https://pu 25 January 2024.
bmed.ncbi.nlm.nih.gov/30482605) .
18. Srivastav, A.; Malviya, M.; Tyagi, P. C. & Nigam, P.
14. Armstrong, E. E.; Khan, A.; Taylor, R. W.; Gouy, A.; (2011). Indian National Studbook of the Bengal Tiger
Greenbaum, G.; Thiéry, A; Kang, J. T.; Redondo, S. A.; (Panthera tigris tigris) (https://cza.nic.in/uploads/docu
Prost, S.; Barsh, G.; Kaelin, C.; Phalke, S.; Chugani, A.; ments/studbooks/english/Bengal%20Tiger%20Studbo
Gilbert, M.; Miquelle, D.; Zachariah, A.; Borthakur, U.; ok%202011.pdf) (PDF) (Report). Central Zoo
Reddy, A.; Louis, E.; Ryder, O. A.; Jhala, Y. V.; Petrov, D.; Authority, Wildlife Institute of India. Retrieved 27 May
Excoffier, L.; Hadly, E. & Ramakrishnan, U. (2021). 2024.
"Recent evolutionary history of tigers highlights
19. Mazák, J. H. (2010). "Craniometric variation in the tiger
contrasting roles of genetic drift and selection" (http
(Panthera tigris): Implications for patterns of diversity,
s://www.ncbi.nlm.nih.gov/pmc/articles/PMC813651
taxonomy and conservation". Mammalian Biology. 75
3) . Molecular Biology and Evolution. 38 (6): 2366–
(1): 45–68. Bibcode:2010MamBi..75...45M (https://ui.a
2379. doi:10.1093/molbev/msab032 (https://doi.org/1
dsabs.harvard.edu/abs/2010MamBi..75...45M) .
0.1093%2Fmolbev%2Fmsab032) . PMC 8136513 (htt
doi:10.1016/j.mambio.2008.06.003 (https://doi.org/10.
ps://www.ncbi.nlm.nih.gov/pmc/articles/PMC813651
1016%2Fj.mambio.2008.06.003) .
3) . PMID 33592092 (https://pubmed.ncbi.nlm.nih.go
v/33592092) . 20. Illiger, C. (1815). "Überblick der Säugethiere nach ihrer
Verteilung über die Welttheile" (https://bibliothek.bbaw.
15. Wang, C.; Wu, D. D.; Yuan, Y. H.; Yao, M. C.; Han, J. L.;
de/digitalisierte-sammlungen/akademieschriften/ansi
Wu, Y. J.; Shan, F.; Li, W. P.; Zhai, J. Q.; Huang, M; Peng,
cht-akademieschriften?tx_bbaw_academicpublications
S. H.; Cai, Q .H.; Yu, J. Y.; Liu, Q. X.; Lui, Z. Y.; Li, L. X.;
how%5Baction%5D=show&tx_bbaw_academicpublicati
Teng, M. S.; Huang, W.; Zhou, J. Y.; Zhang, C.; Chen, W.
onshow%5Bcontroller%5D=AcademicPublication%5CV
& Tu, X. L. (2023). "Population genomic analysis
olume&tx_bbaw_academicpublicationshow%5Bpage%
provides evidence of the past success and future
5D=195&tx_bbaw_academicpublicationshow%5Bvolu
potential of South China tiger captive conservation" (htt
me%5D=85&cHash=f015ec3f9a13240a9559ebdcb88a
ps://www.ncbi.nlm.nih.gov/pmc/articles/PMC1011177
afa4) . Abhandlungen der Königlichen Preußischen
2) . BMC Biology. 21 (1): 64. doi:10.1186/s12915-023-
Akademie der Wissenschaften zu Berlin. 1804–1811:
01552-y (https://doi.org/10.1186%2Fs12915-023-0155
39–159.
2-y) . PMC 10111772 (https://www.ncbi.nlm.nih.gov/
pmc/articles/PMC10111772) . PMID 37069598 (http 21. Sludskii 1992, p. 137.

s://pubmed.ncbi.nlm.nih.gov/37069598) .

16. "Tiger" (http://www.catsg.org/index.php?id=124) .

CatSG. Retrieved 14 June 2024.
22. Driscoll, C. A.; Yamaguchi, N.; Bar-Gal, G. K.; Roca, A. L.; 28. Hu, J.; Westbury, M. V.; Yuan, J.; Wang, C.; Xiao, B.;
Luo, S.; MacDonald, D. W. & O'Brien, S. J. (2009). Chen, S.; Song, S.; Wang, L.; Lin, H.; Lai, X. & Sheng, G.
"Mitochondrial phylogeography illuminates the origin of (2022). "An extinct and deeply divergent tiger lineage
the extinct Caspian Tiger and its relationship to the from northeastern China recognized through
Amur Tiger" (https://www.ncbi.nlm.nih.gov/pmc/article palaeogenomics" (https://www.ncbi.nlm.nih.gov/pmc/
s/PMC2624500) . PLOS ONE. 4 (1): e4125. articles/PMC9326283) . Proceedings of the Royal
Bibcode:2009PLoSO...4.4125D (https://ui.adsabs.harva Society B: Biological Sciences. 289 (1979).
rd.edu/abs/2009PLoSO...4.4125D) . doi:10.1098/rspb.2022.0617 (https://doi.org/10.1098%
doi:10.1371/journal.pone.0004125 (https://doi.org/10. 2Frspb.2022.0617) . PMC 9326283 (https://www.ncb
1371%2Fjournal.pone.0004125) . PMC 2624500 (http i.nlm.nih.gov/pmc/articles/PMC9326283) .
s://www.ncbi.nlm.nih.gov/pmc/articles/PMC262450 PMID 35892215 (https://pubmed.ncbi.nlm.nih.gov/358
0) . PMID 19142238 (https://pubmed.ncbi.nlm.nih.go 92215) .
v/19142238) .
29. Mazák, V. (1968). "Nouvelle sous-espèce de tigre
23. Seidensticker, J.; Christie, S. & Jackson, P. (1999). provenant de l'Asie du sud-est". Mammalia. 32 (1):
"Preface" in Seidensticker, Christie & Jackson 1999, 104–112. doi:10.1515/mamm.1968.32.1.104 (https://d
pp. xv–xx oi.org/10.1515%2Fmamm.1968.32.1.104) .
S2CID 84054536 (https://api.semanticscholar.org/Cor
24. Temminck, C. J. (1844). "Aperçu général et spécifique
pusID:84054536) .
sur les Mammifères qui habitent le Japon et les Iles qui
en dépendent" (https://archive.org/details/faunajaponi 30. Mazák, J. H. & Groves, C. P. (2006). "A taxonomic
casi00sieb/page/43) . In Siebold, P. F. v.; Temminck, revision of the tigers (Panthera tigris) of Southeast
C. J.; Schlegel, H. (eds.). Fauna Japonica sive Descriptio Asia" (http://www.dl.edi-info.ir/A%20taxonomic%20revi
animalium, quae in itinere per Japoniam, jussu et sion%20of%20the%20tigers%20of%20Southeast%20As
auspiciis superiorum, qui summum in India Batava ia.pdf) (PDF). Mammalian Biology. 71 (5): 268–287.
imperium tenent, suscepto, annis 1825–1830 collegit, Bibcode:2006MamBi..71..268M (https://ui.adsabs.harv
notis, observationibus et adumbrationibus illustravit Ph. ard.edu/abs/2006MamBi..71..268M) .
Fr. de Siebold. Leiden: Lugduni Batavorum. doi:10.1016/j.mambio.2006.02.007 (https://doi.org/10.
1016%2Fj.mambio.2006.02.007) . Archived (https://w
25. Sludskii 1992, pp. 131, 134.
eb.archive.org/web/20230531133543/http://www.dl.e
26. Hilzheimer, M. (1905). "Über einige Tigerschädel aus di-info.ir/A%20taxonomic%20revision%20of%20the%20
der Straßburger zoologischen Sammlung" (https://archi tigers%20of%20Southeast%20Asia.pdf) (PDF) from
ve.org/details/zoologischeranze28deut/page/596) . the original on 31 May 2023. Retrieved 15 January
Zoologischer Anzeiger. 28: 594–599. 2024.

27. Sun, X; Liu, Y-C; Tiunov, M. P.; Gimranov, D. O.; Zhuang,

Y; Han, Y; Driscoll, C. A.; Driscoll, C. A.; Pang, Y; Li, C;
Pan, Y; Velasco, M. S.; Gopalakrishnan, S; Yang, R-Z; Li,
B-G; Jin, K; Xu, X; Uphyrkina, O; Huang, Y; Wu, X-H;
Gilbert, M. T. P.; O’Brien, S. J.; Yamaguchi, N; Luo, S-J
(2023). "Ancient DNA reveals genetic admixture in
China during tiger evolution". Nature Ecology &
Evolution. 7: 914–1929. doi:10.1038/s41559-023-
02185-8 (https://doi.org/10.1038%2Fs41559-023-0218
5-8) .
31. Luo, S.-J.; Kim, J.-H.; Johnson, W. E.; van der Walt, J.; 37. Davis, B. W.; Li, G. & Murphy, W. J. (2010). "Supermatrix
Martenson, J.; Yuhki, N.; Miquelle, D. G.; Uphyrkina, O.; and species tree methods resolve phylogenetic
Goodrich, J. M.; Quigley, H. B.; Tilson, R.; Brady, G.; relationships within the big cats, Panthera (Carnivora:
Martelli, P.; Subramaniam, V.; McDougal, C.; Hean, S.; Felidae)". Molecular Phylogenetics and Evolution. 56 (1):
Huang, S.-Q.; Pan, W.; Karanth, U. K.; Sunquist, M.; 64–76. Bibcode:2010MolPE..56...64D (https://ui.adsab
Smith, J. L. D. & O'Brien, S. J. (2004). "Phylogeography s.harvard.edu/abs/2010MolPE..56...64D) .
and genetic ancestry of tigers (Panthera tigris)" (http doi:10.1016/j.ympev.2010.01.036 (https://doi.org/10.1
s://www.ncbi.nlm.nih.gov/pmc/articles/PMC53481 016%2Fj.ympev.2010.01.036) . PMID 20138224 (http
0) . PLOS Biology. 2 (12): e442. s://pubmed.ncbi.nlm.nih.gov/20138224) .
doi:10.1371/journal.pbio.0020442 (https://doi.org/10.1
38. Cho, Y. S.; Hu, L.; Hou, H.; Lee, H.; Xu, J.; Kwon, S.; Oh,
371%2Fjournal.pbio.0020442) . PMC 534810 (https://
S.; Kim, H. M.; Jho, S.; Kim, S.; Shin, Y. A.; Kim, B. C.;
www.ncbi.nlm.nih.gov/pmc/articles/PMC534810) .
Kim, H.; Kim, C. U.; Luo, S. J.; Johnson, W. E.; Koepfli, K.
PMID 15583716 (https://pubmed.ncbi.nlm.nih.gov/155
P.; Schmidt-Küntzel, A.; Turner, J. A.; Marker, L.; Harper,
83716) .
C.; Miller, S. M.; Jacobs, W.; Bertola, L. D.; Kim, T. H.;
32. Schwarz, E. (1912). "Notes on Malay tigers, with Lee, S.; Zhou, Q.; Jung, H. J.; Xu, X. & Gadhvi, P. (2013).
description of a new form from Bali" (https://archive.or "The tiger genome and comparative analysis with lion
g/stream/annalsmagazineof8101912lond#page/324/ and snow leopard genomes" (https://www.ncbi.nlm.ni
mode/2up) . Annals and Magazine of Natural History. h.gov/pmc/articles/PMC3778509) . Nature
8. 10 (57): 324–326. Communications. 4: 2433.
doi:10.1080/00222931208693243 (https://doi.org/10. Bibcode:2013NatCo...4.2433C (https://ui.adsabs.harva
1080%2F00222931208693243) . rd.edu/abs/2013NatCo...4.2433C) .
doi:10.1038/ncomms3433 (https://doi.org/10.1038%2
33. Mazak, V. (2004). Der Tiger (in German). Westarp
Fncomms3433) . hdl:2263/32583 (https://hdl.handle.
Wissenschaften Hohenwarsleben. ISBN 978-3-89432-
net/2263%2F32583) . PMC 3778509 (https://www.nc
759-0.
bi.nlm.nih.gov/pmc/articles/PMC3778509) .
34. Mazák, V.; Groves, C. P. & Van Bree, P. (1978). "Skin and PMID 24045858 (https://pubmed.ncbi.nlm.nih.gov/240
Skull of the Bali Tiger, and a list of preserved 45858) .
specimens of Panthera tigris balica (Schwarz, 1912)".
39. Tseng, Z. J.; Wang, X.; Slater, G. J.; Takeuchi, G. T.; Li, Q.;
Zeitschrift für Säugetierkunde. 43 (2): 108–113.
Liu, J. & Xie, G. (2014). "Himalayan fossils of the oldest
35. Pocock, R. I. (1929). "Tigers" (https://archive.org/detail known pantherine establish ancient origin of big cats"
s/journalofbomb33341929bomb/page/n185) . (https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3843
Journal of the Bombay Natural History Society. 33: 505– 846) . Proceedings of the Royal Society B: Biological
541. Sciences. 281 (1774): 20132686.
doi:10.1098/rspb.2013.2686 (https://doi.org/10.1098%
36. Li, G.; Davis, B. W.; Eizirik, E.; Murphy, W. J. (2016).
"Phylogenomic evidence for ancient hybridization in 2Frspb.2013.2686) . PMC 3843846 (https://www.ncb

the genomes of living cats (Felidae)" (https://www.ncb i.nlm.nih.gov/pmc/articles/PMC3843846) .

i.nlm.nih.gov/pmc/articles/PMC4691742) . Genome PMID 24225466 (https://pubmed.ncbi.nlm.nih.gov/242

25466) .
Research. 26 (1): 1–11. doi:10.1101/gr.186668.114 (htt
ps://doi.org/10.1101%2Fgr.186668.114) .
PMC 4691742 (https://www.ncbi.nlm.nih.gov/pmc/arti
cles/PMC4691742) . PMID 26518481 (https://pubme
d.ncbi.nlm.nih.gov/26518481) .
40. Mazák, J. H.; Christiansen, P. & Kitchener, A. C. (2011). 45. Gabryś, J.; Kij, B.; Kochan, J. & Bugno-Poniewierska, M.
"Oldest Known Pantherine Skull and Evolution of the (2021). "Interspecific hybrids of animals-in nature,
Tiger" (https://www.ncbi.nlm.nih.gov/pmc/articles/PM breeding and science–a review" (https://doi.org/10.247
C3189913) . PLOS ONE. 6 (10): e25483. 8%2Faoas-2020-0082) . Annals of Animal Science. 21
Bibcode:2011PLoSO...625483M (https://ui.adsabs.harv (2): 403–415. doi:10.2478/aoas-2020-0082 (https://do
ard.edu/abs/2011PLoSO...625483M) . i.org/10.2478%2Faoas-2020-0082) .
doi:10.1371/journal.pone.0025483 (https://doi.org/10.
46. "Genomic Imprinting" (https://learn.genetics.utah.edu/
1371%2Fjournal.pone.0025483) . PMC 3189913 (http
content/epigenetics/imprinting/) . Genetic Science
s://www.ncbi.nlm.nih.gov/pmc/articles/PMC318991
Learning Center, Utah.org. Archived (https://web.archiv
3) . PMID 22016768 (https://pubmed.ncbi.nlm.nih.go
e.org/web/20190904215316/https://learn.genetics.uta
v/22016768) .
h.edu/content/EPIGENETICS/imprinting/) from the
41. Hemmer, H. (2023). "The evolution of the original on 4 September 2019. Retrieved 26 August
palaeopantherine cats, Palaeopanthera gen. nov. 2018.
Blytheae (Tseng et al., 2014) and Palaeopanthera
47. Mazák, V. (1981). "Panthera tigris" (https://doi.org/10.2
pamiri (Ozansoy, 1959) comb. Nov. (Mammalia,
307%2F3504004) . Mammalian Species (152): 1–8.
Carnivora, Felidae)". Palaeobiodiversity and
doi:10.2307/3504004 (https://doi.org/10.2307%2F350
Palaeoenvironments. 103 (4): 827–839.
4004) . JSTOR 3504004 (https://www.jstor.org/stabl
Bibcode:2023PdPe..103..827H (https://ui.adsabs.harva
e/3504004) .
rd.edu/abs/2023PdPe..103..827H) .
doi:10.1007/s12549-023-00571-5 (https://doi.org/10.1 48. Sludskii 1992, p. 98.

007%2Fs12549-023-00571-5) . S2CID 257842190 (htt

49. Thapar 2004, p. 26.
ps://api.semanticscholar.org/CorpusID:257842190) .
50. Zhao, C; Dai, W; Liu, Q; Liu, D; Roberts, N. J.; Liu, Z;
42. Jiangzuo, Q.; Wang, Y.; Ge, J.; Liu, S.; Song, Y.; Jin, C.; Gong, M; Qiu, H; Liu, C; Liu, D; Ma, G; Jiang, G (2024).
Jiang, H. & Liu, J. (2023). "Discovery of jaguar from "Combination of facial and nose features of Amur
northeastern China middle Pleistocene reveals an
tigers to determine age". Integrative Zoology.
intercontinental dispersal event". Historical Biology. 35 doi:10.1111/1749-4877.12817 (https://doi.org/10.111
(3): 293–302. Bibcode:2023HBio...35..293J (https://ui. 1%2F1749-4877.12817) .
adsabs.harvard.edu/abs/2023HBio...35..293J) .
51. Sludskii 1992, p. 103.
doi:10.1080/08912963.2022.2034808 (https://doi.org/
10.1080%2F08912963.2022.2034808) . 52. Thapar 2004, p. 25.
S2CID 246693903 (https://api.semanticscholar.org/Co
53. Novak, R. M. & Walker, E. P. (1999). "Panthera tigris
rpusID:246693903) .
(tiger)" (https://books.google.com/books?id=T37sFCl4
43. Kitchener, A. & Yamaguchi, N. (2009). "What is a Tiger? 3E8C&pg=PA825) . Walker's Mammals of the World
Biogeography, Morphology, and Taxonomy" in Tilson & (6th ed.). Baltimore: Johns Hopkins University Press.
Nyhus 2010, pp. 53–84 pp. 825–828. ISBN 978-0-8018-5789-8. Archived (http
s://web.archive.org/web/20240505145513/https://boo
44. Hasegawa, Y., Takakuwa, Y., Nenoki, K. & Kimura, T.
ks.google.com/books?id=T37sFCl43E8C&pg=PA825#v
Fossil tiger from limestone mine of Tsukumi City, Oita
=onepage&q&f=false) from the original on 5 May
Prefecture, Kyushu Island, Japan (https://www.gmnh.pr
2024. Retrieved 17 October 2020.
ef.gunma.jp/wp-content/uploads/bulletin23_1.pdf) .
Bull. Gunma Museum Nat. Hist. 23, (2019) (in 54. Sunquist, M. (2010). "What is a Tiger? Ecology and
Japanese with English abstract) Behaviour" in Tilson & Nyhus 2010, pp. 19−34

55. Thapar 2004, p. 28.

56. Sludskii 1992, pp. 99–102. 62. Xu, X.; Dong, G. X.; Schmidt-Küntzel, A.; Zhang, X. L.;
Zhuang, Y.; Fang, R.; Sun, X.; Hu, X.S.; Zhang, T. Y.; Yang,
57. Miquelle, D. "Tiger" in MacDonald 2001, pp. 18–21
H. D.; Zhang, D. L.; Marker, L.; Jiang, Z.-F.; Li, R. & Luo,
58. Allen, W. L.; Cuthill, I. C.; Scott-Samuel, N. E. & Baddeley, S.-J. (2017). "The genetics of tiger pelage color
R. (2010). "Why the leopard got its spots: relating variations" (https://www.luo-lab.org/publications/Xu17
pattern development to ecology in felids" (https://www. -CellRes-GoldenTiger.pdf) (PDF). Cell Research. 27
ncbi.nlm.nih.gov/pmc/articles/PMC3061134) . (7): 954–957. doi:10.1038/cr.2017.32 (https://doi.org/
Proceedings of the Royal Society B. 278 (1710): 1373– 10.1038%2Fcr.2017.32) . PMC 5518981 (https://ww
1380. doi:10.1098/rspb.2010.1734 (https://doi.org/10. w.ncbi.nlm.nih.gov/pmc/articles/PMC5518981) .
1098%2Frspb.2010.1734) . PMC 3061134 (https://w PMID 28281538 (https://pubmed.ncbi.nlm.nih.gov/282
ww.ncbi.nlm.nih.gov/pmc/articles/PMC3061134) . 81538) . Archived (https://web.archive.org/web/2021
PMID 20961899 (https://pubmed.ncbi.nlm.nih.gov/209 0104160444/https://www.luo-lab.org/publications/Xu1
61899) . 7-CellRes-GoldenTiger.pdf) (PDF) from the original on

59. Caro, T. (2005). "The adaptive significance of coloration 4 January 2021. Retrieved 25 August 2018.

in mammals". BioScience. 55 (2): 125–136. 63. Xavier, N. (2010). "A new conservation policy needed
doi:10.1641/0006- for reintroduction of Bengal tiger-white" (https://www.c
3568(2005)055[0125:TASOCI]2.0.CO;2 (https://doi.org/ urrentscience.ac.in/Volumes/99/07/0894.pdf) (PDF).
10.1641%2F0006-3568%282005%29055%5B0125%3A Current Science. 99 (7): 894–895. Archived (https://we
TASOCI%5D2.0.CO%3B2) . b.archive.org/web/20240407184215/https://www.curr

60. Godfrey, D.; Lythgoe, J. N. & Rumball, D. A. (1987). entscience.ac.in/Volumes/99/07/0894.pdf) (PDF)

from the original on 7 April 2024. Retrieved 29 January
"Zebra stripes and tiger stripes: the spatial frequency
distribution of the pattern compared to that of the 2024.

background is significant in display and crypsis". 64. Sagar, V.; Kaelin, C. B.; Natesh, M.; Reddy, P. A.;
Biological Journal of the Linnean Society. 32 (4): 427– Mohapatra, R. K.; Chhattani, H.; Thatte, P.;
433. doi:10.1111/j.1095-8312.1987.tb00442.x (https:// Vaidyanathan, S.; Biswas, S.; Bhatt, S. & Paul, S. (2021).
doi.org/10.1111%2Fj.1095-8312.1987.tb00442.x) . "High frequency of an otherwise rare phenotype in a

61. Fennell, J. G.; Talas, L.; Baddeley, R. J.; Cuthill, I. C. & small and isolated tiger population" (https://www.ncbi.

Scott-Samuel, N. E. (2019). "Optimizing colour for nlm.nih.gov/pmc/articles/PMC8488692) .

Proceedings of the National Academy of Sciences. 118
camouflage and visibility using deep learning: the
effects of the environment and the observer's visual (39): e2025273118. Bibcode:2021PNAS..11825273S (h

system" (https://www.ncbi.nlm.nih.gov/pmc/articles/P ttps://ui.adsabs.harvard.edu/abs/2021PNAS..1182527

MC6544896) . Journal of the Royal Society Interface. 3S) . doi:10.1073/pnas.2025273118 (https://doi.org/

16 (154): 20190183. doi:10.1098/rsif.2019.0183 (http 10.1073%2Fpnas.2025273118) . PMC 8488692 (http

s://www.ncbi.nlm.nih.gov/pmc/articles/PMC848869
s://doi.org/10.1098%2Frsif.2019.0183) .
PMC 6544896 (https://www.ncbi.nlm.nih.gov/pmc/arti 2) . PMID 34518374 (https://pubmed.ncbi.nlm.nih.go

cles/PMC6544896) . PMID 31138092 (https://pubme v/34518374) .

d.ncbi.nlm.nih.gov/31138092) .
65. Sanderson, E. W.; Miquelle, D. G.; Fisher, K.; Harihar, A.; 71. Li, X.Y.; Hu, W.Q.; Wang, H.J. & Jiang, X.L. (2023). "Tiger
Clark, C.; Moy, J.; Potapov, P.; Robinson, N.; Royte, L.; reappearance in Medog highlights the conservation
Sampson, D.; Sanderlin, J.; Yackulic, C. B.; Belecky, M.; values of the region for this apex predator" (https://ww
Breitenmoser, U.; Breitenmoser-Würsten, C.; Chanchani, w.ncbi.nlm.nih.gov/pmc/articles/PMC10415778) .
P.; Chapman, S.; Deomurari, A.; Duangchantrasiri, S.; Zoological Research. 44 (4): 747–749.
Facchini, E.; Gray, T. N. E.; Goodrich, J.; Hunter, L.; doi:10.24272/j.issn.2095-8137.2023.178 (https://doi.or
Linkie, M.; Marthy, W.; Rasphone, A.; Roy, S.; Sittibal, D.; g/10.24272%2Fj.issn.2095-8137.2023.178) .
Tempa, T.; Umponjan, M. & Wood, K. (2023). "Range- PMC 10415778 (https://www.ncbi.nlm.nih.gov/pmc/ar
wide trends in tiger conservation landscapes, 2001– ticles/PMC10415778) . PMID 37464931 (https://pub
2020" (https://doi.org/10.3389%2Ffcosc.2023.119128 med.ncbi.nlm.nih.gov/37464931) .
0) . Frontiers in Conservation Science. 4: 1191280.
72. Simcharoen, S.; Pattanavibool, A.; Karanth, K. U.;
doi:10.3389/fcosc.2023.1191280 (https://doi.org/10.3
Nichols, J. D. & Kumar, N. S. (2007). "How many tigers
389%2Ffcosc.2023.1191280) .
Panthera tigris are there in Huai Kha Khaeng Wildlife
66. Sludskii 1992, pp. 108–112. Sanctuary, Thailand? An estimate using photographic
capture-recapture sampling" (https://doi.org/10.1017%
67. Miquelle, D. G.; Smirnov, E. N.; Merrill, T. W.; Myslenkov,
2FS0030605307414107) . Oryx. 41 (4): 447–453.
A. E.; Quigley, H.; Hornocker, M. G. & Schleyer, B.
doi:10.1017/S0030605307414107 (https://doi.org/10.
(1999). "Hierarchical spatial analysis of Amur tiger
1017%2FS0030605307414107) .
relationships to habitat and prey" in Seidensticker,
Christie & Jackson 1999, pp. 71–99 73. Wibisono, H. T.; Linkie, M.; Guillera-Arroita, G.; Smith, J.
A.; Sunarto; Pusarini, W.; Asriadi; Baroto, P.; Brickle, N.;
68. Wikramanayake, E. D.; Dinerstein, E.; Robinson, J. G.;
Dinata, Y.; Gemita, E.; Gunaryadi, D.; Haidir, I. A. &
Karanth, K. U.; Rabinowitz, A.; Olson, D.; Mathew, T.;
Herwansyah (2011). "Population status of a cryptic top
Hedao, P.; Connor, M.; Hemley, G. & Bolze, D. (1999).
predator: An island-wide assessment of Tigers in
"Where can tigers live in the future? A framework for
Sumatran rainforests" (https://www.ncbi.nlm.nih.gov/p
identifying high-priority areas for the conservation of
mc/articles/PMC3206793) . PLOS ONE. 6 (11):
tigers in the wild" in Seidensticker, Christie & Jackson
e25931. Bibcode:2011PLoSO...625931W (https://ui.ads
1999, pp. 254–272
abs.harvard.edu/abs/2011PLoSO...625931W) .
69. Jigme, K. & Tharchen, L. (2012). "Camera-trap records doi:10.1371/journal.pone.0025931 (https://doi.org/10.
of tigers at high altitudes in Bhutan". Cat News (56): 1371%2Fjournal.pone.0025931) . PMC 3206793 (http
14–15. s://www.ncbi.nlm.nih.gov/pmc/articles/PMC320679

70. Adhikarimayum, A. S. & Gopi, G. V. (2018). "First 3) . PMID 22087218 (https://pubmed.ncbi.nlm.nih.go

photographic record of tiger presence at higher v/22087218) .

elevations of the Mishmi Hills in the Eastern Himalayan 74. Bisht, S.; Banerjee, S.; Qureshi, Q. & Jhala, Y. (2019).
Biodiversity Hotspot, Arunachal Pradesh, India" (http "Demography of a high-density tiger population and its
s://doi.org/10.11609%2Fjott.4381.10.13.12833-1283 implications for tiger recovery" (https://doi.org/10.111
6) . Journal of Threatened Taxa. 10 (13): 12833– 1%2F1365-2664.13410) . Journal of Applied Ecology.
12836. doi:10.11609/jott.4381.10.13.12833-12836 (htt 56 (7): 1725–1740. Bibcode:2019JApEc..56.1725B (htt
ps://doi.org/10.11609%2Fjott.4381.10.13.12833-1283 ps://ui.adsabs.harvard.edu/abs/2019JApEc..56.1725
6) . B) . doi:10.1111/1365-2664.13410 (https://doi.org/1
0.1111%2F1365-2664.13410) .
75. Lynam, A. J.; Rabinowitz, A.; Myint, T.; Maung, M.; Latt, 79. Naha, D.; Jhala, Y. V.; Qureshi, Q.; Roy, M.; Sankar, K. &
K. T. & Po, S. H. T. (2009). "Estimating abundance with Gopal, R. (2016). "Ranging, activity and habitat use by
sparse data: tigers in northern Myanmar". Population tigers in the mangrove forests of the Sundarban" (http
Ecology. 51 (1): 115–121. s://www.ncbi.nlm.nih.gov/pmc/articles/PMC482276
Bibcode:2009PopEc..51..115L (https://ui.adsabs.harva 5) . PLOS ONE. 11 (4): e0152119.
rd.edu/abs/2009PopEc..51..115L) . Bibcode:2016PLoSO..1152119N (https://ui.adsabs.har
doi:10.1007/s10144-008-0093-5 (https://doi.org/10.10 vard.edu/abs/2016PLoSO..1152119N) .
07%2Fs10144-008-0093-5) . doi:10.1371/journal.pone.0152119 (https://doi.org/10.
1371%2Fjournal.pone.0152119) . PMC 4822765 (http
76. Phumanee, W.; Steinmetz, R.; Phoonjampa, R.;
s://www.ncbi.nlm.nih.gov/pmc/articles/PMC482276
Weingdow, S.; Phokamanee, S.; Bhumpakphan, N. &
5) . PMID 27049644 (https://pubmed.ncbi.nlm.nih.go
Savini, T. (2021). "Tiger density, movements, and
v/27049644) .
immigration outside of a tiger source site in Thailand"
(https://doi.org/10.1111%2Fcsp2.560) . Conservation 80. Pokheral, C. P. & Wegge, P. (2019). "Coexisting large
Science and Practice. 3 (12): e560. carnivores: spatial relationships of tigers and leopards
Bibcode:2021ConSP...3E.560P (https://ui.adsabs.harva and their prey in a prey-rich area in lowland Nepal".
rd.edu/abs/2021ConSP...3E.560P) . Écoscience. 26 (1): 1–9. Bibcode:2019Ecosc..26....1P
doi:10.1111/csp2.560 (https://doi.org/10.1111%2Fcsp (https://ui.adsabs.harvard.edu/abs/2019Ecosc..26....1
2.560) . P) . doi:10.1080/11956860.2018.1491512 (https://do
i.org/10.1080%2F11956860.2018.1491512) .
77. Rayan, D. M. & Linkie, M. (2015). "Conserving tigers in
S2CID 92446020 (https://api.semanticscholar.org/Cor
Malaysia: A science-driven approach for eliciting
pusID:92446020) .
conservation policy change". Biological Conservation.
184: 18–26. Bibcode:2015BCons.184...18R (https://ui. 81. Yang, H.; Han, S.; Xie, B.; Mou, P.; Kou, X.; Wang, T.; Ge,
adsabs.harvard.edu/abs/2015BCons.184...18R) . J. & Feng, L. (2019). "Do prey availability, human
doi:10.1016/j.biocon.2014.12.024 (https://doi.org/10.1 disturbance and habitat structure drive the daily
016%2Fj.biocon.2014.12.024) . activity patterns of Amur tigers (Panthera tigris
altaica)?". Journal of Zoology. 307 (2): 131–140.
78. Carter, N. H.; Shrestha, B. K.; Karki, J. B.; Pradhan, N. M.
doi:10.1111/jzo.12622 (https://doi.org/10.1111%2Fjzo.
B. & Liu, J. (2012). "Coexistence between wildlife and
12622) . S2CID 92736301 (https://api.semanticschol
humans at fine spatial scales" (https://www.ncbi.nlm.ni
ar.org/CorpusID:92736301) .
h.gov/pmc/articles/PMC3458348) . Proceedings of
the National Academy of Sciences. 109 (38): 15360– 82. Thapar 2004, pp. 26, 64–66.
15365. Bibcode:2012PNAS..10915360C (https://ui.ads
83. Barlow, A. C. D.; Smith, J. L. D.; Ahmad, I. U.; Hossain, A.
abs.harvard.edu/abs/2012PNAS..10915360C) .
N. M.; Rahman, M. & Howlader, A. (2011). "Female tiger
doi:10.1073/pnas.1210490109 (https://doi.org/10.107
Panthera tigris home range size in the Bangladesh
3%2Fpnas.1210490109) . PMC 3458348 (https://ww
Sundarbans: the value of this mangrove ecosystem for
w.ncbi.nlm.nih.gov/pmc/articles/PMC3458348) .
the species' conservation" (https://doi.org/10.1017%2F
PMID 22949642 (https://pubmed.ncbi.nlm.nih.gov/229
S0030605310001456) . Oryx. 45 (1): 125–128.
49642) .
doi:10.1017/S0030605310001456 (https://doi.org/10.
1017%2FS0030605310001456) .
84. Sarkar, M.S.; Ramesh, K.; Johnson, J. A.; Sen, S.; Nigam, 91. Burger, B. V.; Viviers, M. Z.; Bekker, J. P. I.; Roux, M.;
P.; Gupta, S. K.; Murthy, R. S. & Saha, G. K. (2016). Fish, N.; Fourie, W. B. & Weibchen, G. (2008). "Chemical
"Movement and home range characteristics of characterization of territorial marking fluid of male
reintroduced tiger (Panthera tigris) population in Panna Bengal tiger, Panthera tigris" (https://citeseerx.ist.psu.e
Tiger Reserve, central India". European Journal of du/document?repid=rep1&type=pdf&doi=586948b839
Wildlife Research. 62 (5): 537–547. 6932dd13d9e5a880e77cb7618a273f) . Journal of
doi:10.1007/s10344-016-1026-9 (https://doi.org/10.10 Chemical Ecology. 34 (5): 659–671.
07%2Fs10344-016-1026-9) . S2CID 254187854 (http Bibcode:2008JCEco..34..659B (https://ui.adsabs.harva
s://api.semanticscholar.org/CorpusID:254187854) . rd.edu/abs/2008JCEco..34..659B) .
doi:10.1007/s10886-008-9462-y (https://doi.org/10.10
85. Goodrich, J. M.; Miquelle, D. G.; Smirnov, E.M.; Kerley,
07%2Fs10886-008-9462-y) . hdl:10019.1/11220 (http
L.L.; Quigley, H. B. & Hornocker, M. G. (2010). "Spatial
s://hdl.handle.net/10019.1%2F11220) .
structure of Amur (Siberian) tigers (Panthera tigris
PMID 18437496 (https://pubmed.ncbi.nlm.nih.gov/184
altaica) on Sikhote-Alin Biosphere Zapovednik, Russia"
37496) . S2CID 5558760 (https://api.semanticschola
(https://doi.org/10.1644%2F09-mamm-a-293.1) .
r.org/CorpusID:5558760) . Archived (https://web.archi
Journal of Mammalogy. 91 (3): 737–748.
ve.org/web/20230718031651/https://citeseerx.ist.psu.
doi:10.1644/09-mamm-a-293.1 (https://doi.org/10.164
edu/document?repid=rep1&type=pdf&doi=586948b839
4%2F09-mamm-a-293.1) .
6932dd13d9e5a880e77cb7618a273f) from the
86. Klevtcova, A. V.; Miquelle, D. G.; Seryodkin, I. V.; Bragina, original on 18 July 2023. Retrieved 29 June 2023.
E. V.; Soutyrina, S. V. & Goodrich, J. M. (2021). "The
92. Smith, J. L. D.; McDougal, C. & Miquelle, D. (1989).
influence of reproductive status on home range size
"Scent marking in free-ranging tigers, Panthera tigris".
and spatial dynamics of female Amur tigers". Mammal
Animal Behaviour. 37: 1–10. doi:10.1016/0003-
Research. 66: 83–94. doi:10.1007/s13364-020-00547-2
3472(89)90001-8 (https://doi.org/10.1016%2F0003-34
(https://doi.org/10.1007%2Fs13364-020-00547-2) .
72%2889%2990001-8) . S2CID 53149100 (https://api.
87. Joshi, A.; Vaidyanathan, S.; Mondol, S.; Edgaonkar, A. & semanticscholar.org/CorpusID:53149100) .
Ramakrishnan, U. (2013). "Connectivity of Tiger
93. Thapar 2004, p. 105.
(Panthera tigris) Populations in the Human-Influenced
Forest Mosaic of Central India" (https://www.ncbi.nlm. 94. Mills 2004, p. 85–86.
nih.gov/pmc/articles/PMC3819329) . PLOS ONE. 8
95. Schaller 1967, pp. 244–251.
(11): e77980. Bibcode:2013PLoSO...877980J (https://u
i.adsabs.harvard.edu/abs/2013PLoSO...877980J) . 96. Mills 2004, p. 89.

doi:10.1371/journal.pone.0077980 (https://doi.org/10. 97. Schaller 1967, pp. 262–263.

1371%2Fjournal.pone.0077980) . PMC 3819329 (http
98. Schaller 1967, p. 263.
s://www.ncbi.nlm.nih.gov/pmc/articles/PMC381932
9) . PMID 24223132 (https://pubmed.ncbi.nlm.nih.go 99. Sunquist, M. E. & Sunquist, F. (2002). "Tiger Panthera
v/24223132) . tigris (Linnaeus, 1758)" (https://books.google.com/boo
ks?id=IF8nDwAAQBAJ&pg=PA320) . Wild Cats of the
88. Thapar 2004, p. 76.
World. Chicago: University of Chicago Press. pp. 343–
89. Smith, J. L. D. (1993). "The role of dispersal in 372. ISBN 978-0-226-77999-7.
structuring the Chitwan tiger population". Behaviour.
100. Thapar 2004, p. 29.
124 (3): 165–195. doi:10.1163/156853993X00560 (htt
ps://doi.org/10.1163%2F156853993X00560) . 101. Schaller 1967, p. 256.

90. Mills 2004, pp. 54–55. 102. Thapar 2004, p. 99.

103. Schaller 1967, pp. 258–261. 111. Karanth, K. U. & Nichols, J. D. (1998). "Estimation of
tiger densities in India using photographic captures
104. Schaller 1967, p. 261.
and recaptures" (http://erepo.usiu.ac.ke/bitstream/han
105. Peters, G. & Tonkin-Leyhausen, B. A. (1999). "Evolution dle/11732/758/Estimation%20of%20tiger%20densitie
of acoustic communication signals of mammals: s%20in%20India%20using%20photographic%20capture
Friendly close-range vocalizations in Felidae s%20and%20recaptures.pdf?sequence=4&isAllowed=
(Carnivora)". Journal of Mammalian Evolution. 6 (2): y) (PDF). Ecology. 79 (8): 2852–2862.
129–159. doi:10.1023/A:1020620121416 (https://doi.o doi:10.1890/0012-
rg/10.1023%2FA%3A1020620121416) . 9658(1998)079[2852:EOTDII]2.0.CO;2 (https://doi.org/
S2CID 25252052 (https://api.semanticscholar.org/Cor 10.1890%2F0012-9658%281998%29079%5B2852%3A
pusID:25252052) . EOTDII%5D2.0.CO%3B2) . JSTOR 176521 (https://ww
w.jstor.org/stable/176521) . Archived (https://web.arc
106. Schaller 1967, pp. 257–258.
hive.org/web/20221127044620/http://erepo.usiu.ac.k
107. Schaller 1967, pp. 256–258.
e/bitstream/handle/11732/758/Estimation%20of%20ti

108. Mills 2004, p. 62. ger%20densities%20in%20India%20using%20photogra

phic%20captures%20and%20recaptures.pdf?sequence
109. Hayward, M. W.; Jędrzejewski, W. & Jędrzejewska, B.
=4&isAllowed=y) (PDF) from the original on 27
(2012). "Prey preferences of the tiger Panthera tigris".
November 2022. Retrieved 16 December 2021.
Journal of Zoology. 286 (3): 221–231.
doi:10.1111/j.1469-7998.2011.00871.x (https://doi.or 112. Perry, R. (1965). The World of the Tiger. pp. 133–134.

g/10.1111%2Fj.1469-7998.2011.00871.x) . ASIN B0007DU2IU (https://www.amazon.com/dp/B000

7DU2IU) .
110. Karanth, K. U. (2003). "Tiger ecology and conservation
in the Indian subcontinent" (http://repository.ias.ac.in/8 113. Fàbregas, M. C.; Fosgate, G. T. & Koehler, G. M. (2015).

9489/1/50p.pdf) (PDF). Journal of the Bombay "Hunting performance of captive-born South China

Natural History Society. 100 (2 & 3): 169–189. Archived tigers (Panthera tigris amoyensis) on free-ranging prey

(https://web.archive.org/web/20231007210705/http:// and implications for their reintroduction". Biological

repository.ias.ac.in/89489/1/50p.pdf) (PDF) from the Conservation. 192: 57–64.

original on 7 October 2023. Retrieved 13 April 2024. Bibcode:2015BCons.192...57F (https://ui.adsabs.harva

rd.edu/abs/2015BCons.192...57F) .
doi:10.1016/j.biocon.2015.09.007 (https://doi.org/10.1
016%2Fj.biocon.2015.09.007) . hdl:2263/50208 (http
s://hdl.handle.net/2263%2F50208) .

114. Thapar 2004, pp. 63, 111.

115. Schaller 1967, pp. 284–285.

116. Schaller 1967, p. 288.

117. Thapar 2004, p. 120.

118. Thapar 2004, pp. 119–120, 122.

119. Schaller 1967, p. 287.

120. Thapar 2004, p. 23.

121. Thapar 2004, p. 121.

122. Schaller 1967, p. 295. 133. Karanth, K. U. & Sunquist, M. E. (1995). "Prey selection
by tiger, leopard and dhole in tropical forests". Journal
123. Mills 2004, p. 24.
of Animal Ecology. 64 (4): 439–450.
124. Christiansen, P. (2007). "Canine morphology in the Bibcode:1995JAnEc..64..439K (https://ui.adsabs.harva
larger Felidae: implications for feeding ecology" (http rd.edu/abs/1995JAnEc..64..439K) . doi:10.2307/5647
s://doi.org/10.1111%2Fj.1095-8312.2007.00819.x) . (https://doi.org/10.2307%2F5647) . JSTOR 5647 (http
Biological Journal of the Linnean Society. 91 (4): 573– s://www.jstor.org/stable/5647) .
592. doi:10.1111/j.1095-8312.2007.00819.x (https://do
134. Odden, M.; Wegge, P. & Fredriksen, T. (2010). "Do tigers
i.org/10.1111%2Fj.1095-8312.2007.00819.x) .
displace leopards? If so, why?". Ecological Research. 25
125. Schaller 1967, pp. 295–296. (4): 875–881. Bibcode:2010EcoR...25..875O (https://ui.

126. Thapar 2004, p. 126. adsabs.harvard.edu/abs/2010EcoR...25..875O) .

doi:10.1007/s11284-010-0723-1 (https://doi.org/10.10
127. Schaller 1967, p. 289.
07%2Fs11284-010-0723-1) .

128. Schaller 1967, pp. 297–300.

135. Harihar, A.; Pandav, B. & Goyal, S. P. (2011). "Responses

129. Srivathsa, A.; Ramachandran, V.; Saravanan, P.; of leopard Panthera pardus to the recovery of a tiger

Sureshbabu, A.; Ganguly, D. & Ramakrishnan, U. (2023). Panthera tigris population" (https://doi.org/10.1111%2F

"Topcats and underdogs: intraguild interactions among j.1365-2664.2011.01981.x) . Journal of Applied

three apex carnivores across Asia's forestscapes". Ecology. 48 (3): 806–814.

Biological Reviews. 98 (6): 2114–2135. Bibcode:2011JApEc..48..806H (https://ui.adsabs.harva

doi:10.1111/brv.12998 (https://doi.org/10.1111%2Fbr rd.edu/abs/2011JApEc..48..806H) .

v.12998) . PMID 37449566 (https://pubmed.ncbi.nlm. doi:10.1111/j.1365-2664.2011.01981.x (https://doi.or

nih.gov/37449566) . S2CID 259903849 (https://api.se g/10.1111%2Fj.1365-2664.2011.01981.x) .

manticscholar.org/CorpusID:259903849) .
136. Habib, B.; Nigam P.; Ghaskadbi P. & Bhandari A. (2021).

130. Thapar 2004, p. 136. "Dhole pack size variation: Assessing the effect of prey
availability and apex predator" (https://www.ncbi.nlm.ni
131. Thinley, P.; Rajaratnam, R.; Lassoie, J. P.; Morreale, S. J.;
h.gov/pmc/articles/PMC8093734) . Ecology and
Curtis, P. D.; Vernes, K.; Leki Leki; Phuntsho, S.; Dorji, T.
Evolution. 11 (9): 4774–4785.
& Dorji, P. (2018). "The ecological benefit of tigers
Bibcode:2021EcoEv..11.4774B (https://ui.adsabs.harva
(Panthera tigris) to farmers in reducing crop and
rd.edu/abs/2021EcoEv..11.4774B) .
livestock losses in the eastern Himalayas: Implications
doi:10.1002/ece3.7380 (https://doi.org/10.1002%2Fec
for conservation of large apex predators" (https://doi.or
e3.7380) . PMC 8093734 (https://www.ncbi.nlm.nih.g
g/10.1016%2Fj.biocon.2018.01.015) . Biological
ov/pmc/articles/PMC8093734) . PMID 33976847 (htt
Conservation. 219: 119–125.
ps://pubmed.ncbi.nlm.nih.gov/33976847) .
doi:10.1016/j.biocon.2018.01.015 (https://doi.org/10.1
016%2Fj.biocon.2018.01.015) . 137. Steinmetz, R.; Seuaturien, N. & Chutipong, W. (2013).
"Tigers, leopards, and dholes in a half-empty forest:
132. Karanth, K. U. & Sunquist, M. E. (2000). "Behavioural
Assessing species interactions in a guild of threatened
correlates of predation by tiger (Panthera tigris),
carnivores". Biological Conservation. 163: 68–78.
leopard (Panthera pardus) and dhole (Cuon alpinus) in
doi:10.1016/j.biocon.2012.12.016 (https://doi.org/10.1
Nagarahole, India". Journal of Zoology. 250 (2): 255–
016%2Fj.biocon.2012.12.016) .
265. doi:10.1111/j.1469-7998.2000.tb01076.x (https://
doi.org/10.1111%2Fj.1469-7998.2000.tb01076.x) . 138. Mills 2004, p. 42.

139. Thapar 2004, p. 145.

140. Sankhala, K. S. (1967). "Breeding behaviour of the tiger 151. Mills 2004, pp. 61, 66–67.
Panthera tigris in Rajasthan". International Zoo
152. Thapar 2004, p. 63.
Yearbook. 7 (1): 133–147. doi:10.1111/j.1748-
1090.1967.tb00354.x (https://doi.org/10.1111%2Fj.174 153. Schaller 1967, pp. 270, 276.

8-1090.1967.tb00354.x) . 154. Pacifici, M.; Santini, L.; Di Marco, M.; Baisero, D.;

141. Thapar 2004, p. 148. Francucci, L.; Grottolo Marasini, G.; Visconti, P.;
Rondinini, C. (2013). "Generation length for mammals"
142. Thapar 2004, p. 45.
(https://doi.org/10.3897%2Fnatureconservation.5.573
143. Mills 2004, p. 50–51. 4) . Nature Conservation. 5 (5): 87–94.
doi:10.3897/natureconservation.5.5734 (https://doi.or
144. Mills 2004, p. 50.
g/10.3897%2Fnatureconservation.5.5734) .
145. Thapar 2004, p. 51.
155. Majumder, A. (2023). "Survival strategy – Temporal
146. Mills 2004, p. 51. segregation of different age and sex classes of a

147. Petrunenko, Y. K.; Seryodkin, I. V.; Bragina, E. V.; Bengal tiger (Panthera tigris tigris Linnaeus) population

Soutyrina, S. S.; Mukhacheva, A. S.; Rybin, N. N. & in Pench Tiger Reserve, Madhya Pradesh India" (http

Miquelle, D. G. (2019). "How does a tigress balance the s://doi.org/10.61186%2FJAD.2023.5.3.5) . Journal of

Animal Diversity. 5 (3): 43–54.
opposing constraints of raising cubs?". Mammal
Research. 65: 245–253. doi:10.1007/s13364-019- doi:10.61186/JAD.2023.5.3.5 (https://doi.org/10.6118

00466-x (https://doi.org/10.1007%2Fs13364-019-0046 6%2FJAD.2023.5.3.5) .

6-x) . 156. Tidière, M.; Müller, P.; Sliwa, A.; Siberchicot, A. & Douay,

148. Robinson, H. S.; Goodrich, J. M.; Miquelle, D. G.; Miller, G. (2021). "Sex-specific actuarial and reproductive
senescence in zoo-housed tiger (Panthera tigris): The
C. S. & Seryodkin, I. V. (2015). "Mortality of Amur tigers:
The more things change, the more they stay the same". importance of sub-species for conservation" (https://fi

Integrative Zoology. 10 (4): 344–353. ndresearcher.sdu.dk/ws/files/182491417/zoo.21610.p

doi:10.1111/1749-4877.12147 (https://doi.org/10.111 df) (PDF). Zoo Biology. 40 (4): 320–329.

doi:10.1002/zoo.21610 (https://doi.org/10.1002%2Fzo
1%2F1749-4877.12147) .
o.21610) . PMID 33861886 (https://pubmed.ncbi.nlm.
149. Sadhu, A.; Jayam, P. P. C.; Qureshi, Q.; Shekhawat, R. S.;
nih.gov/33861886) . Archived (https://web.archive.or
Sharma, S. & Jhala, Y. V. (2017). "Demography of a
g/web/20240430010012/https://findresearcher.sdu.d
small, isolated tiger (Panthera tigris tigris) population in
k/ws/files/182491417/zoo.21610.pdf) (PDF) from
a semi-arid region of western India" (https://doi.org/10.
the original on 30 April 2024. Retrieved 24 April 2024.
1186%2Fs40850-017-0025-y) . BMC Zoology. 2: 16.
doi:10.1186/s40850-017-0025-y (https://doi.org/10.11 157. Mills 2004, pp. 59, 89.

86%2Fs40850-017-0025-y) . 158. Thapar 2004, pp. 55–56.

150. Barlow, A. C. D.; McDougal, C.; Smith, J. L. D.; Gurung, 159. Pandey, G. (2011). "India male tiger plays doting dad to
B.; Bhatta, S. R.; Kumal, S.; Mahato, B. & Taman, D. B. orphaned cubs" (https://www.bbc.com/news/world-so
(2009). "Temporal variation in Tiger (Panthera tigris) uth-asia-13598386) . BBC News. Archived (https://we
populations and its implications for monitoring" (http b.archive.org/web/20240408231242/https://www.bbc.
s://doi.org/10.1644%2F07-MAMM-A-415.1) . Journal com/news/world-south-asia-13598386) from the
of Mammalogy. 90 (2): 472–478. doi:10.1644/07- original on 8 April 2024. Retrieved 14 February 2024.
MAMM-A-415.1 (https://doi.org/10.1644%2F07-MAM
160. Mills 2004, p. 59.
M-A-415.1) .
161. Thapar 2004, p. 66.
162. Seimon, T. A.; Miquelle, D. G.; Chang, T. Y.; Newton, A. L.; 166. Aung, S. S.; Shwe, N. M.; Frechette, J.; Grindley, M. &
Korotkova, I.; Ivanchuk, G.; Lyubchenko, E.; Tupikov, A.; Connette, G. (2017). "Surveys in southern Myanmar
Slabe, E.; McAloose, D. (2013). "Canine distemper virus: indicate global importance for tigers and biodiversity"
an emerging disease in wild endangered Amur tigers (https://doi.org/10.1017%2FS0030605316001393) .
(Panthera tigris altaica)" (https://www.ncbi.nlm.nih.gov/ Oryx. 51 (1): 13. doi:10.1017/S0030605316001393 (htt
pmc/articles/PMC3747579) . mBio. 4 (4). ps://doi.org/10.1017%2FS0030605316001393) .
doi:10.1128/mbio.00410-13 (https://doi.org/10.1128%
167. Suttidate, N.; Steinmetz, R.; Lynam, A. J.; Sukmasuang,
2Fmbio.00410-13) . PMC 3747579 (https://www.ncbi.
R.; Ngoprasert, D.; Chutipong, W.; Bateman, B. L.; Jenks,
nlm.nih.gov/pmc/articles/PMC3747579) .
K. E.; Baker-Whatton, M.; Kitamura, S. & Ziółkowska, E.
PMID 23943758 (https://pubmed.ncbi.nlm.nih.gov/239
(2021). "Habitat connectivity for endangered
43758) .
Indochinese tigers in Thailand" (https://doi.org/10.101
163. Quigley, K. S.; Evermann, J. F.; Leathers, C. W.; 6%2Fj.gecco.2021.e01718) . Global Ecology and
Armstrong, D. L.; Goodrich, J.; Duncan, N. M. & Conservation. 29: e01718.
Miquelle, D. G. (2010). "Morbillivirus infection in a wild Bibcode:2021GEcoC..2901718S (https://ui.adsabs.harv
Siberian tiger in the Russian Far East". Journal of ard.edu/abs/2021GEcoC..2901718S) .
Wildlife Diseases. 46 (4): 1252–1256. doi:10.1016/j.gecco.2021.e01718 (https://doi.org/10.1
doi:10.7589/0090-3558-46.4.1252 (https://doi.org/10.7 016%2Fj.gecco.2021.e01718) .
589%2F0090-3558-46.4.1252) . PMID 20966275 (http
168. Shevade, V. S.; Potapov, P. V.; Harris, N. L. & Loboda, T.
s://pubmed.ncbi.nlm.nih.gov/20966275) .
V. (2017). "Expansion of industrial plantations
164. McCauley, D.; Stout, V.; Gairhe, K. P.; Sadaula, A.; Dubovi, continues to threaten Malayan tiger habitat" (https://do
E.; Subedi, S. & Kaufman, G. E. (2021). "Serologic i.org/10.3390%2Frs9070747) . Remote Sensing. 9 (7):
survey of selected pathogens in free-ranging Bengal 747. Bibcode:2017RemS....9..747S (https://ui.adsabs.h
tigers (Panthera tigris tigris) in Nepal" (https://doi.org/1 arvard.edu/abs/2017RemS....9..747S) .
0.7589%2FJWD-D-20-00046) . The Journal of Wildlife doi:10.3390/rs9070747 (https://doi.org/10.3390%2Frs
Diseases. 57 (2): 393–398. doi:10.7589/JWD-D-20- 9070747) . hdl:1903/31503 (https://hdl.handle.net/19
00046 (https://doi.org/10.7589%2FJWD-D-20-0004 03%2F31503) .
6) . PMID 33822151 (https://pubmed.ncbi.nlm.nih.go
169. Debonne, N.; van Vliet, J. & Verburg, P. (2019). "Future
v/33822151) .
governance options for large-scale land acquisition in
165. Schoen, J. M.; Neelakantan, A.; Cushman, S. A.; Dutta, Cambodia: impacts on tree cover and tiger landscapes"
T.; Habib, B.; Jhala, Y. V.; Mondal, I.; Ramakrishnan, U.; (https://doi.org/10.1016%2Fj.envsci.2018.12.031) .
Reddy, P. A.; Saini, S. & Sharma, S. (2022). Environmental Science & Policy. 94: 9–19.
"Synthesizing habitat connectivity analyses of a Bibcode:2019ESPol..94....9D (https://ui.adsabs.harvar
globally important human-dominated tiger- d.edu/abs/2019ESPol..94....9D) .
conservation landscape" (https://www.ncbi.nlm.nih.go doi:10.1016/j.envsci.2018.12.031 (https://doi.org/10.1
v/pmc/articles/PMC9545158) . Conservation Biology. 016%2Fj.envsci.2018.12.031) . hdl:1871.1/1dced676-
36 (4): e13909. Bibcode:2022ConBi..36E3909S (http 560b-46fb-a7c5-e0c888c5cff1 (https://hdl.handle.net/
s://ui.adsabs.harvard.edu/abs/2022ConBi..36E3909 1871.1%2F1dced676-560b-46fb-a7c5-e0c888c5cff
S) . doi:10.1111/cobi.13909 (https://doi.org/10.111 1) .
1%2Fcobi.13909) . PMC 9545158 (https://www.ncbi.n
lm.nih.gov/pmc/articles/PMC9545158) .
PMID 35288989 (https://pubmed.ncbi.nlm.nih.gov/352
88989) .
170. Wang, D.; Smith, J. L.; Accatino, F.; Ge, J. & Wang, T. 175. Rasphone, A.; Kéry, M.; Kamler, J. F. & Macdonald, D. W.
(2023). "Addressing the impact of canine distemper (2019). "Documenting the demise of tiger and leopard,
spreading on an isolated tiger population in northeast and the status of other carnivores and prey, in Lao
Asia". Integrative Zoology. 18 (6): 994–1008. PDR's most prized protected area: Nam Et-Phou Louey"
doi:10.1111/1749-4877.12712 (https://doi.org/10.111 (https://doi.org/10.1016%2Fj.gecco.2019.e00766) .
1%2F1749-4877.12712) . PMID 36881515 (https://pu Global Ecology and Conservation. 20: e00766.
bmed.ncbi.nlm.nih.gov/36881515) . Bibcode:2019GEcoC..2000766R (https://ui.adsabs.harv
ard.edu/abs/2019GEcoC..2000766R) .
171. Tilson, R.; Defu, H.; Muntifering, J. & Nyhus, P. J. (2004).
doi:10.1016/j.gecco.2019.e00766 (https://doi.org/10.1
"Dramatic decline of wild South China tigers Panthera
016%2Fj.gecco.2019.e00766) .
tigris amoyensis: field survey of priority tiger reserves"
(https://doi.org/10.1017%2FS0030605304000079) . 176. Linkie, M.; Martyr, D.; Harihar, A.; Mardiah, S.; Hodgetts,
Oryx. 38 (1): 40–47. doi:10.1017/S0030605304000079 T.; Risdianto, D.; Subchaan, M. & Macdonald, D. (2018).
(https://doi.org/10.1017%2FS0030605304000079) . "Asia's economic growth and its impact on Indonesia's
tigers". Biological Conservation. 219: 105–109.
172. Nyhus, P. (2008). "Panthera tigris ssp. amoyensis" (http
Bibcode:2018BCons.219..105L (https://ui.adsabs.harv
s://www.iucnredlist.org/species/15965/5334628) .
ard.edu/abs/2018BCons.219..105L) .
IUCN Red List of Threatened Species. 2008:
doi:10.1016/j.biocon.2018.01.011 (https://doi.org/10.1
e.T15965A5334628.
016%2Fj.biocon.2018.01.011) .
doi:10.2305/IUCN.UK.2008.RLTS.T15965A5334628.en
(https://doi.org/10.2305%2FIUCN.UK.2008.RLTS.T1596 177. Slaght, J. C.; Milakovsky, B.; Maksimova, D.A.; Zaitsev,
5A5334628.en) . V. A.; Seryodkin, I.; Panichev, A. & Miquelle, D. (2017).
"Anthropogenic influences on the distribution of a
173. Jhala, Y.; Gopal, R.; Mathur, V.; Ghosh, P.; Negi, H. S.;
Vulnerable coniferous forest specialist: habitat
Narain, S.; Yadav, S. P.; Malik, A.; Garawad, R. & Qureshi,
selection by the Siberian musk deer Moschus
Q. (2021). "Recovery of tigers in India: Critical
moschiferus" (https://doi.org/10.1017%2FS003060531
introspection and potential lessons" (https://doi.org/1
6001617) . Oryx. 53 (1): 174–180.
0.1002%2Fpan3.10177) . People and Nature. 3 (2):
doi:10.1017/S0030605316001617 (https://doi.org/10.
281–293. Bibcode:2021PeoNa...3..281J (https://ui.ads
1017%2FS0030605316001617) .
abs.harvard.edu/abs/2021PeoNa...3..281J) .
doi:10.1002/pan3.10177 (https://doi.org/10.1002%2Fp 178. Nijman, V. & Shepherd, C. R. (2015). "Trade in tigers and
an3.10177) . other wild cats in Mong La and Tachilek, Myanmar – A
tale of two border towns" (https://doi.org/10.1016%2Fj.
174. Shwe, N. M.; Grainger, M.; Ngoprasert, D.; Aung, S. S.;
biocon.2014.10.031) . Biological Conservation. 182:
Grindley, M. & Savini, T. (2023). "Anthropogenic
1–7. Bibcode:2015BCons.182....1N (https://ui.adsabs.
pressure on large carnivores and their prey in the highly
harvard.edu/abs/2015BCons.182....1N) .
threatened forests of Tanintharyi, southern Myanmar"
doi:10.1016/j.biocon.2014.10.031 (https://doi.org/10.1
(https://doi.org/10.1017%2FS0030605321001654) .
016%2Fj.biocon.2014.10.031) .
Oryx. 57 (2): 262–271.
doi:10.1017/S0030605321001654 (https://doi.org/10.
1017%2FS0030605321001654) . hdl:11250/3040580
(https://hdl.handle.net/11250%2F3040580) .
179. Wong, R. & Krishnasamy, K. (2022). Skin and Bones: 184. Saif, S.; Rahman, H. T. & MacMillan, D. C. (2018). "Who
Tiger Trafficking Analysis from January 2000 – June is killing the tiger Panthera tigris and why?" (https://doi.
2022 (https://www.traffic.org/site/assets/files/19714/ org/10.1017%2FS0030605316000491) . Oryx. 52 (1):
skin_and_bones_tiger_trafficking_analysis_from_januar 46–54. doi:10.1017/S0030605316000491 (https://doi.
y_2000_to_june_2022_r7.pdf) (PDF). Petaling Jaya, org/10.1017%2FS0030605316000491) .
Selangor, Malaysia: TRAFFIC, Southeast Asia Regional
185. Uddin, N.; Enoch, S.; Harihar, A.; Pickles, R. S. & Hughes,
Office. Archived (https://web.archive.org/web/2024011
A. C. (2023). "Tigers at a crossroads: Shedding light on
7215040/https://www.traffic.org/site/assets/files/197
the role of Bangladesh in the illegal trade of this iconic
14/skin_and_bones_tiger_trafficking_analysis_from_jan
big cat" (https://doi.org/10.1111%2Fcsp2.12952) .
uary_2000_to_june_2022_r7.pdf) (PDF) from the
Conservation Science and Practice. 5 (7): e12952.
original on 17 January 2024. Retrieved 1 March 2024.
Bibcode:2023ConSP...5E2952U (https://ui.adsabs.harv
180. Paudel, P. K.; Acharya, K. P.; Baral, H. S.; Heinen, J. T. & ard.edu/abs/2023ConSP...5E2952U) .
Jnawali, S. R. (2020). "Trends, patterns, and networks doi:10.1111/csp2.12952 (https://doi.org/10.1111%2Fc
of illicit wildlife trade in Nepal: A national synthesis" (ht sp2.12952) .
tps://doi.org/10.1111%2Fcsp2.247) . Conservation
186. Davis, E. O.; Willemsen, M.; Dang, V.; O’Connor, D. &
Science and Practice. 2 (9): e247.
Glikman, J. A. (2020). "An updated analysis of the
Bibcode:2020ConSP...2E.247P (https://ui.adsabs.harva
consumption of tiger products in urban Vietnam" (http
rd.edu/abs/2020ConSP...2E.247P) .
s://doi.org/10.1016%2Fj.gecco.2020.e00960) . Global
doi:10.1111/csp2.247 (https://doi.org/10.1111%2Fcsp
Ecology and Conservation. 22: e00960.
2.247) .
Bibcode:2020GEcoC..2200960D (https://ui.adsabs.har
181. Nittu, G.; Shameer, T. T.; Nishanthini, N. K. & Sanil, R. vard.edu/abs/2020GEcoC..2200960D) .
(2023). "The tide of tiger poaching in India is rising! An doi:10.1016/j.gecco.2020.e00960 (https://doi.org/10.1
investigation of the intertwined facts with a focus on 016%2Fj.gecco.2020.e00960) .
conservation" (https://www.ncbi.nlm.nih.gov/pmc/artic
187. Henry, L. (2020). "5 Things Tiger King Doesn't Explain
les/PMC9005341) . GeoJournal. 88 (1): 753–766.
About Captive Tigers" (https://www.worldwildlife.org/st
doi:10.1007/s10708-022-10633-4 (https://doi.org/10.1
ories/5-things-tiger-king-doesn-t-explain-about-captive-
007%2Fs10708-022-10633-4) . PMC 9005341 (http
tigers) . World Wildlife Fund for Nature. Archived (http
s://www.ncbi.nlm.nih.gov/pmc/articles/PMC900534
s://web.archive.org/web/20240220001943/https://ww
1) . PMID 35431409 (https://pubmed.ncbi.nlm.nih.go
w.worldwildlife.org/stories/5-things-tiger-king-doesn-t-
v/35431409) .
explain-about-captive-tigers) from the original on 20
182. Khanwilkar, S.; Sosnowski, M. & Guynup, S. (2022). February 2024. Retrieved 19 February 2024.
"Patterns of illegal and legal tiger parts entering the
188. Singh, R.; Nigam, P.; Qureshi, Q.; Sankar, K.; Krausman,
United States over a decade (2003–2012)" (https://doi.
P. R.; Goyal, S. P. & Nicholoson, K. L. (2015).
org/10.1111%2Fcsp2.622) . Conservation Science and
"Characterizing human–tiger conflict in and around
Practice. 4 (3): e622. Bibcode:2022ConSP...4E.622K (ht
Ranthambhore Tiger Reserve, western India". European
tps://ui.adsabs.harvard.edu/abs/2022ConSP...4E.622
Journal of Wildlife Research. 61 (2): 255–261.
K) . doi:10.1111/csp2.622 (https://doi.org/10.1111%2
doi:10.1007/s10344-014-0895-z (https://doi.org/10.10
Fcsp2.622) .
07%2Fs10344-014-0895-z) .
183. Van Uhm, D. P. (2016). The Illegal Wildlife Trade: Inside
the World of Poachers, Smugglers and Traders (Studies
of Organized Crime). New York: Springer.
189. Chowdhurym, A. N.; Mondal, R.; Brahma, A. & Biswas, 194. Global Tiger Recovery Program (2023–34) (https://glob
M. K. (2016). "Ecopsychosocial aspects of human– altigerforum.org/global-tiger-recovery-program-2-0-202
tiger conflict: An ethnographic study of tiger widows of 3-34/) (Report). Global Tiger Forum and the Global
Sundarban Delta, India". Environmental Health Insights. Tiger Initiative Council. 2023. Archived (https://web.arc
10: 1–29. doi:10.4137/EHI.S24 (https://doi.org/10.413 hive.org/web/20240408150606/https://globaltigerforu
7%2FEHI.S24) (inactive 10 April 2024). m.org/global-tiger-recovery-program-2-0-2023-34/)
from the original on 8 April 2024. Retrieved 8 April
190. Dhungana, R.; Savini, T.; Karki, J. B.; Dhakal, M.;
2024.
Lamichhane, B. R. & Bumrungsri, S. (2018). "Living with
tigers Panthera tigris: Patterns, correlates, and contexts 195. DNPWC & DFSC (2022). Status of Tigers and Prey in
of human–tiger conflict in Chitwan National Park, Nepal 2022 (https://dnpwc.gov.np/media/files/Status_
Nepal" (https://doi.org/10.1017%2FS00306053160015 of_Tigers_Ic2ylSC.pdf) (PDF). Kathmandu, Nepal:
87) . Oryx. 52 (1): 55–65. Department of National Parks and Wildlife
doi:10.1017/S0030605316001587 (https://doi.org/10. Conservation and Department of Forests and Soil
1017%2FS0030605316001587) . hdl:1887/57668 (htt Conservation. Ministry of Forests and Environment.
ps://hdl.handle.net/1887%2F57668) . Archived (https://web.archive.org/web/202304081040
09/https://dnpwc.gov.np/media/files/Status_of_Tigers
191. Lubis, M. I.; Pusparini, W.; Prabowo, S. A.; Marthy, W.;
_Ic2ylSC.pdf) (PDF) from the original on 8 April 2023.
Tarmizi; Andayani, N. & Linkie, M. (2020). "Unraveling
Retrieved 6 August 2022.
the complexity of human–tiger conflicts in the Leuser
Ecosystem, Sumatra". Animal Conservation. 23 (6): 196. Department of Forests and Park Services (2023).
741–749. Bibcode:2020AnCon..23..741L (https://ui.ad "National Tiger Survey 2021–22". Thimphu:
sabs.harvard.edu/abs/2020AnCon..23..741L) . Government of Bhutan.
doi:10.1111/acv.12591 (https://doi.org/10.1111%2Fac
197. Sludskii 1992, p. 202.
v.12591) .
198. Tandin, T.; Penjor, U.; Tempa, T.; Dhendup, P.; Dorji, S.;
192. Neo, W. H. Y.; Lubis, M. I. & Lee, J. S. H. (2023).
Wangdi, S. & Moktan, V. (2018). Tiger Action Plan for
"Settlements and plantations are sites of human–tiger
Bhutan (2018-2023): A landscape approach to tiger
interactions in Riau, Indonesia" (https://doi.org/10.101
conservation (Report). Thimphu, Bhutan: Nature
7%2FS0030605322000667) . Oryx. 57 (4): 476–480.
Conservation Division, Department of Forests and Park
doi:10.1017/S0030605322000667 (https://doi.org/10.
Services, Ministry of Agriculture and Forests.
1017%2FS0030605322000667) . hdl:10356/165557
doi:10.13140/RG.2.2.14890.70089 (https://doi.org/10.
(https://hdl.handle.net/10356%2F165557) .
13140%2FRG.2.2.14890.70089) .
193. Qureshi, Q.; Jhala, Y. V.; Yadav, S. P. & Mallick, A. (2023).
199. Aryal, R. S. (2004). CITES Implementation in Nepal and
Status of tigers, co-predators and prey in India 2022 (htt
India. Law, Policy and Practice. Kathmandu: Bhrikuti
ps://wii.gov.in/images//images/documents/publicatio
Aademic Publications. ISBN 99933-673-4-6.
ns/statu_tiger_copredators-2022.pdf) (PDF). New
Delhi, Dehradun: National Tiger Conservation Authority
& Wildlife Institute of India. Archived (https://web.archi
ve.org/web/20240505145403/https://wii.gov.in/image
s//images/documents/publications/statu_tiger_copred
ators-2022.pdf) (PDF) from the original on 5 May
2024. Retrieved 18 February 2024.
200. Malaysian Conservation Alliance for Tigers (2006). The 205. Hossain, A. N. M.; Lynam, A. J.; Ngoprasert, D.; Barlow,
Malayan Tiger Conservation Programme (https://conse A.; Barlow, C. G. & Savini, T. (2018). "Identifying
rvewildcats.org/wp-content/uploads/sites/5/WildCats/ landscape factors affecting tiger decline in the
ProjectReports/Malaysia/MYCATFinalreport2006-07.p Bangladesh Sundarbans" (https://doi.org/10.1016%2Fj.
df) (PDF) (Report). Kuala Lumpur: Department of gecco.2018.e00382) . Global Ecology and
Wildlife and National Parks Peninsular Malaysia. Conservation. 13: e00382.
Archived (https://web.archive.org/web/202404092034 Bibcode:2018GEcoC..1300382H (https://ui.adsabs.har
38/https://conservewildcats.org/wp-content/uploads/ vard.edu/abs/2018GEcoC..1300382H) .
sites/5/WildCats/ProjectReports/Malaysia/MYCATFin doi:10.1016/j.gecco.2018.e00382 (https://doi.org/10.1
alreport2006-07.pdf) (PDF) from the original on 9 016%2Fj.gecco.2018.e00382) .
April 2024. Retrieved 9 April 2024.
206. Pisdamkham, C.; Prayurasiddhi, T.; Kanchanasaka, B.;
201. Ministry of Forestry (2007). Conservation Strategy and Maneesai, R.; Simcharoen, S.; Pattanavibool, A.;
Action Plan for the Sumatran Tiger (Panthera tigris Duangchantrasiri, S.; Simcharoen, A.; Pattanavibool, R.;
sumatrae) Indonesia 2007-2017 (https://globaltigerforu Maneerat, S.; Prayoon, U.; Cutter, P. G. & Smith, J. L. D.
m.org/wp-content/uploads/2017/04/indonesia.pdf) (2010). Thailand Tiger Action Plan 2010–2022 (https://
(PDF). Government of Indonesia. Archived (https://web. www.researchgate.net/publication/303881962) .
archive.org/web/20230923144656/https://globaltigerf Bangkok: Department of National Parks, Wildlife and
orum.org/wp-content/uploads/2017/04/indonesia.pd Plant Conservation, Ministry of Natural Resources and
f) (PDF) from the original on 23 September 2023. Environment. Archived (https://web.archive.org/web/2
Retrieved 12 April 2024. 0240505145525/https://www.researchgate.net/public
ation/303881962_Thailand_Tiger_Action_Plan_2010-2
202. Yeh, E. T. (2012). "Transnational environmentalism and
022) from the original on 5 May 2024. Retrieved
entanglements of sovereignty: The Tiger Campaign
16 April 2024.
across the Himalayas". Political Geography. 31 (7):
408–418. doi:10.1016/j.polgeo.2012.06.003 (https://d 207. Chandradewi, D. S.; Semiadi, G.; Pinondang, I.; Kheng, V.
oi.org/10.1016%2Fj.polgeo.2012.06.003) . & Bahaduri, L. D. (2019). "A decade on: The second
collaborative Sumatra-wide Tiger survey". Cat News.
203. Kampongsun, S. (2022). "The future of Panthera tigris
69: 41–42.
in Thailand and globally" (https://web.archive.org/web/
20231111015312/https://www.iucn.org/story/202208/ 208. Wibisono, H. T. (2021). An Island-wide Status of
future-panthera-tigris-thailand-and-globally) . IUCN. Sumatran Tiger (Panthera tigris sumatrae) and Principal
Archived from the original (https://www.iucn.org/story/ Prey in Sumatra, Indonesia (https://udspace.udel.edu/s
202208/future-panthera-tigris-thailand-and-globally) erver/api/core/bitstreams/db24ae3d-1681-4d30-9773-
on 11 November 2023. Retrieved 8 April 2024. 4abb62cd861d/content) (Doctor of Philosophy in
Entomology and Wildlife Ecology). Delaware: University
204. Lynam, A. J.; Khaing, S. T. & Zaw, K. M. (2006).
of Delaware. Archived (https://web.archive.org/web/20
"Developing a national tiger action plan for the Union of
240505064621/https://udspace.udel.edu/server/api/c
Myanmar". Environmental Management. 37 (1): 30–39.
ore/bitstreams/db24ae3d-1681-4d30-9773-4abb62cd8
Bibcode:2006EnMan..37...30L (https://ui.adsabs.harvar
61d/content) from the original on 5 May 2024.
d.edu/abs/2006EnMan..37...30L) .
Retrieved 14 April 2024.
doi:10.1007/s00267-004-0273-9 (https://doi.org/10.10
07%2Fs00267-004-0273-9) . PMID 16362487 (https://
pubmed.ncbi.nlm.nih.gov/16362487) .
209. Hötte, M. H.; Kolodin, I. A.; Bereznuk, S. L.; Slaght, J. C.; 213. Rahaman, M. H.; Masroor, M.; Sajjad, H. & Saha, T. K.
Kerley, L. L.; Soutyrina, S. V.; Salkina, G. P.; Zaumyslova, (2024). "Assessment of habitat suitability and potential
O. Y.; Stokes, E. J. & Miquelle, D. G. (2016). "Indicators corridors for Bengal Tiger (Panthera tigris tigris) in
of success for smart law enforcement in protected Valmiki Tiger Reserve, India, using MaxEnt model and
areas: A case study for Russian Amur tiger (Panthera Least-Cost modeling approach". Environmental
tigris altaica) reserves". Integrative Zoology. 11 (1): 2– Modeling & Assessment. 29 (2): 405–422.
15. doi:10.1111/1749-4877.12168 (https://doi.org/10.1 Bibcode:2024EMdAs..29..405R (https://ui.adsabs.harv
111%2F1749-4877.12168) . PMID 26458501 (https:// ard.edu/abs/2024EMdAs..29..405R) .
pubmed.ncbi.nlm.nih.gov/26458501) . doi:10.1007/s10666-024-09966-w (https://doi.org/10.1
007%2Fs10666-024-09966-w) .
210. Lam, W. Y.; Phung, C. C.; Mat, Z. A.; Jamaluddin, H.;
Sivayogam, C. P.; Zainal Abidin, F. A.; Sulaiman, A.; 214. Rahman, H.; Hidayat, R. H.; Nofrizal, A. Y.; Wilastra, I. &
Cheok, M. K. Y.; Osama, N. A. W.; Sabaan, S.; Abu Nasution, A. F. R. (2023). "Priority corridor zone for
Hashim, A. K.; Booton, M. D.; Harihar, A.; Clements, G. human-tiger conflict mitigation: A landscape
R. & Pickles, R. S. A. (2023). "Using a crime prevention connectivity approach in West Sumatra region,
framework to evaluate tiger counter-poaching in a Indonesia". Journal for Nature Conservation. 76:
Southeast Asian rainforest" (https://doi.org/10.3389%2 126501. Bibcode:2023JNatC..7626501R (https://ui.ads
Ffcosc.2023.1213552) . Frontiers in Conservation abs.harvard.edu/abs/2023JNatC..7626501R) .
Science. 4: 1213552. doi:10.3389/fcosc.2023.1213552 doi:10.1016/j.jnc.2023.126501 (https://doi.org/10.101
(https://doi.org/10.3389%2Ffcosc.2023.1213552) . 6%2Fj.jnc.2023.126501) .

211. Bhatt, T. R.; Castley, J. G.; Sims-Castley, R.; Bara, H. S. & 215. Paudyal, B. N. (2023). Evaluation of the project on
Chauvenet, A. L. M. (2023). "Connecting tiger (Panthera transboundary cooperation on the conservation of
tigris) populations in Nepal: Identification of corridors Amur tigers, Amur leopards and Snow leopards in
among tiger-bearing protected areas" (https://www.ncb North-East Asia (https://repository.unescap.org/bitstre
i.nlm.nih.gov/pmc/articles/PMC10227491) . Ecology am/handle/20.500.12870/6674/ESCAP-2023-RP-Trans
and Evolution. 13 (5): e10140. boundary-cooperation-conservation-Amur-tigers-Amur-l
Bibcode:2023EcoEv..1310140B (https://ui.adsabs.harv eopards-Snow-leopards-ENEA.pdf) (PDF) (Report).
ard.edu/abs/2023EcoEv..1310140B) . Bangkok, Thailand: United Nations Economic and
doi:10.1002/ece3.10140 (https://doi.org/10.1002%2Fe Social Commission for Asia and the Pacific. Archived
ce3.10140) . PMC 10227491 (https://www.ncbi.nlm.n (https://web.archive.org/web/20240407175636/http
ih.gov/pmc/articles/PMC10227491) . s://repository.unescap.org/bitstream/handle/20.500.1
PMID 37261321 (https://pubmed.ncbi.nlm.nih.gov/372 2870/6674/ESCAP-2023-RP-Transboundary-cooperatio
61321) . n-conservation-Amur-tigers-Amur-leopards-Snow-leopa
rds-ENEA.pdf) (PDF) from the original on 7 April
212. Harihar, A.; Pandav, B.; Ghosh-Harihar, M. & Goodrich, J.
2024. Retrieved 7 April 2024.
(2020). "Demographic and ecological correlates of a
recovering tiger (Panthera tigris) population: Lessons
learnt from 13-years of monitoring". Biological
Conservation. 252: 108848.
Bibcode:2020BCons.25208848H (https://ui.adsabs.har
vard.edu/abs/2020BCons.25208848H) .
doi:10.1016/j.biocon.2020.108848 (https://doi.org/10.
1016%2Fj.biocon.2020.108848) .
216. Priatna, D.; Santosa, Y.; Prasetyo, L. B. & Kartono, A. P. 220. Luo, S.-J.; Johnson, W. E.; Martenson, J.; Antunes, A.;
(2012). "Home range and movements of male Martelli, P.; Uphyrkina, O.; Traylor-Holzer, K.; Smith, J. L.
translocated problem tigers in Sumatra" (http://ajcb.in/j D.; O'Brien, S. J. (2008). "Subspecies genetic
ournals/full_papers/4_AJCB-VOL1-ISSUE1-Priatna%20 assignments of worldwide captive tigers increase
et%20al.pdf) (PDF). Asian Journal of Conservation conservation value of captive populations". Current
Biology. 1 (1): 20–30. Archived (https://web.archive.or Biology. 18 (8): 592–596.
g/web/20240207230500/http://ajcb.in/journals/full_pa doi:10.1016/j.cub.2008.03.053 (https://doi.org/10.101
pers/4_AJCB-VOL1-ISSUE1-Priatna%20et%20al.pdf) 6%2Fj.cub.2008.03.053) .
(PDF) from the original on 7 February 2024. Retrieved
221. Thapar 2004, pp. 187–200.
7 February 2024.
222. Kothari, A. S.; Chhapgar, B. S.; Chhapgar, B. F., eds.
217. Rozhnov, V. V.; Naidenko, S. V.; Hernandez–Blanco, J.
(2005). "The Manpoora Tiger (about a Tiger Hunt in
A.; Chistopolova, M. D.; Sorokin, P. A.; Yachmennikova,
Rajpootanah)". The Treasures of Indian Wildlife.
A. A.; Blidchenko, E. Yu.; Kalinin, A. Yu. & Kastrikin, V.A.
Mumbai: Bombay Natural History Society. pp. 22–27.
(2021). "Restoration of the Amur Tiger (Panthera tigris
ISBN 0195677285.
altaica) population in the Northwest of its distribution
area". Biological Bulletin. 48 (8): 1401–1423. 223. Thapar 2004, p. 193.

Bibcode:2021BioBu..48.1401R (https://ui.adsabs.harva 224. Nyhus, P. J. & Tilson, R. (2010). "Panthera tigris vs

rd.edu/abs/2021BioBu..48.1401R) . Homo sapiens: Conflict, coexistence, or extinction?" in
doi:10.1134/S1062359021080239 (https://doi.org/10. Tilson & Nyhus 2010, pp. 125–142
1134%2FS1062359021080239) .
225. Goodrich, J. M. (2010). "Human–tiger conflict: A review
218. Gray, T. N.; Rosenbaum, R.; Jiang, G.; Izquierdo, P.;
and call for comprehensive plans" (https://doi.org/10.1
Yongchao, J .I. N.; Kesaro, L.; Lyet, A.; Pasha, M. K. S.; 111%2Fj.1749-4877.2010.00218.x) . Integrative
Patterson, D. J.; Channa, P.; Jinzhe, Q. I.; Ripple, W. J.; Zoology. 5 (4): 300–312. doi:10.1111/j.1749-
Roberts, J. L.; Roy, S.; Shwe, N. M.; Wolf, C. & Chapman, 4877.2010.00218.x (https://doi.org/10.1111%2Fj.1749-
S. (2023). "Restoring Asia's roar: Opportunities for tiger
4877.2010.00218.x) . PMID 21392348 (https://pubme
recovery across the historic range" (https://doi.org/10. d.ncbi.nlm.nih.gov/21392348) .
3389%2Ffcosc.2023.1124340) . Frontiers in
226. Mills 2004, pp. 108–110.
Conservation Science. 4: 1124340.
doi:10.3389/fcosc.2023.1124340 (https://doi.org/10.3 227. Thapar 2004, p. 276.
389%2Ffcosc.2023.1124340) .
228. Green 2006, pp. 73–74.
219. Gray, T. N. E.; Crouthers, R.; Ramesh, K.; Vattakaven, J.;
229. Powell, M. A. (2016). "People in peril, environments at
Borah, J.; Pasha, M. K. S.; Lim, T.; Phan, C.; Singh, R.;
risk: coolies, tigers, and colonial Singapore's ecology of
Long, B. & Chapman, S. (2017). "A framework for
poverty". Environment and History. 22 (3): 455–482.
assessing readiness for Tiger Panthera tigris
doi:10.3197/096734016X14661540219393 (https://do
reintroduction: a case study from eastern Cambodia".
i.org/10.3197%2F096734016X14661540219393) .
Biodiversity and Conservation. 26 (10): 2383–2399.
hdl:10356/88201 (https://hdl.handle.net/10356%2F882
Bibcode:2017BiCon..26.2383G (https://ui.adsabs.harva
01) . JSTOR 24810674 (https://www.jstor.org/stable/
rd.edu/abs/2017BiCon..26.2383G) .
24810674) .
doi:10.1007/s10531-017-1365-1 (https://doi.org/10.10
07%2Fs10531-017-1365-1) . 230. Thapar 2004, p. 274.
231. Barlow, A.C.; Ahmad, I. & Smith, J.L. (2013). "Profiling 244. "June 18 Deadline for Compliance With Big Cat Public
tigers (Panthera tigris) to formulate management Safety Act" (https://www.fws.gov/press-release/2023-
responses to human-killing in the Bangladesh 04/big-cat-owners-must-register-june-18#:~:text=The%
Sundarbans". Wildlife Biology in Practice. 9 (2): 30–39. 20Big%20Cat%20Public%20Safety%20Act%20makes%
doi:10.2461/wbp.2013.9.6 (https://doi.org/10.2461%2 20it%20illegal%20to,with%20big%20cats%2C%20includ
Fwbp.2013.9.6) (inactive 19 April 2024). ing%20cubs) . U.S. Fish & Wildlife Service. 2023.
Archived (https://web.archive.org/web/202402202314
232. Mills 2004, pp. 111–113.
26/https://www.fws.gov/press-release/2023-04/big-ca
233. Manfredi, P. "The Tiger in the Ancient World" in Thapar t-owners-must-register-june-18#:~:text=The%20Big%20
2004, p. 173 Cat%20Public%20Safety%20Act%20makes%20it%20ill

234. Thapar 2004, pp. 179–180. egal%20to,with%20big%20cats%2C%20including%20cu

bs) from the original on 20 February 2024. Retrieved
235. Green 2006, pp. 126–130.
20 February 2024.

236. "EU's 'unlikely' role in global tiger trade revealed in new

245. Guidance Document on the export, re-export and intra-
WWF, TRAFFIC report" (https://tigers.panda.org/?9167
EU trade of captive-born and bred live tigers and their
41/EU-tigers-trade-WWF-report) . World Wildlife Fund
parts and derivatives (https://eur-lex.europa.eu/legal-c
for Nature. 30 September 2020. Retrieved 6 June 2023.
ontent/EN/TXT/PDF/?uri=CELEX:52023XC0418(01)#:

237. Smith, K. D.; Snider, R. J.; Dembiec, D. P.; Siegford, J. M. ~:text=While%20most%20EU%20Member%20States,an

& Ali, A. B. (2023). "Effects of a modern exhibit design d%20travelling%20exhibitions%20(9)) (Report).

on captive tiger welfare" (https://doi.org/10.1002%2Fz European Commission. 18 April 2023. Retrieved 6 June

oo.21746) . Zoo Biology. 42 (3): 371–382. 2024.

doi:10.1002/zoo.21746 (https://doi.org/10.1002%2Fzo
246. "Endangered tiger earns its stripes as the world's most
o.21746) . PMID 36478300 (https://pubmed.ncbi.nlm.
popular beast" (https://archive.today/2008012022241
nih.gov/36478300) .
6/http://findarticles.com/p/articles/mi_qn4158/is_200

238. Damasceno, J.; Genaro, G.; Quirke, T.; McCarthy, S.; 41206/ai_n12814678) . The Independent. 2004.

McKeown, S. & O'Riordan, R. (2017). "The effects of Archived from the original (http://findarticles.com/p/ar

intrinsic enrichment on captive felids". Zoo Biology. 36 ticles/mi_qn4158/is_20041206/ai_n12814678) on

(3): 186–192. doi:10.1002/zoo.21361 (https://doi.org/ 20 January 2008. Retrieved 7 March 2009.

10.1002%2Fzoo.21361) . PMID 29165868 (https://pu

247. Albert, C.; Luque, G. M. & Courchamp, F. (2018). "The
bmed.ncbi.nlm.nih.gov/29165868) .
twenty most charismatic species" (https://www.ncbi.nl

239. Thapar 2004, pp. 202–204. m.nih.gov/pmc/articles/PMC6037359) . PLoS One.

13 (7): e0199149. Bibcode:2018PLoSO..1399149A (htt
240. Green 2006, p. 140.
ps://ui.adsabs.harvard.edu/abs/2018PLoSO..1399149
241. Iossa, G.; Soulsbury, C. D. & Harris, S. (2009). "Are wild A) . doi:10.1371/journal.pone.0199149 (https://doi.or
animals suited to a travelling circus life?". Animal g/10.1371%2Fjournal.pone.0199149) . PMC 6037359
Welfare. 18 (2): 129–140. (https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6037
doi:10.1017/S0962728600000270 (https://doi.org/10. 359) . PMID 29985962 (https://pubmed.ncbi.nlm.nih.
1017%2FS0962728600000270) . S2CID 32259865 (ht gov/29985962) .
tps://api.semanticscholar.org/CorpusID:32259865) .
248. Werness, H. B. (2007). The Continuum Encyclopedia of
242. Thapar 2004, pp. 204–205. Animal Symbolism in World Art. New York: Continuum.
pp. 402–404. ISBN 978-0826419132.
243. Thapar 2004, p. 214.
249. Green 2006, pp. 39, 46.
250. Thapar, R. "In Times Past" in Thapar 2004, pp. 156, 164 252. Green 2006, pp. 60, 86–88, 96.

251. Cooper, J. C. (1992). Symbolic and Mythological 253. Green 2006, p. 96.
Animals. London: Aquarian Press. p. 227. ISBN 978-1-
254. Thapar 2004, p. 152.
85538-118-6.
255. Green 2006, pp. 72–73, 78, 125–127, 147–148.

Bibliography

Thapar, V. (2004). Tiger: The Ultimate Guide (https://archive.org/details/tigerultimategui0000thap/mode/2u

p) . New Delhi: CDS Books. ISBN 1-59315-024-5.

Green, S. (2006). Tiger. London: Reaktion Books. ISBN 978-1-86189-276-8.

MacDonald, D., ed. (2001). The Encyclopedia of Mammals (Second ed.). Oxford: Oxford University Press.
ISBN 978-0-7607-1969-5.

Tilson, R.; Nyhus, P. J., eds. (2010). Tigers of the World: The Science, Politics and Conservation of Panthera
tigris (Second ed.). London: Academic Press. ISBN 978-0-08-094751-8.

Mills, S. (2004). Tiger. Richmond Hill: Firefly Books. ISBN 1-55297-949-0.

Schaller, G. B. (1967). The Deer and the Tiger: A Study of Wildlife in India (https://archive.org/details/deertiger
studyof0000scha/page/n419/mode/2up) . Chicago: University of Chicago Press. ISBN 0-226-73631-8.

Seidensticker, J.; Christie, S.; Jackson, P., eds. (1999). Riding the Tiger: Tiger Conservation in Human-
Dominated Landscapes (https://archive.org/details/ridingtigertiger00unse) . Cambridge: Cambridge
University Press. ISBN 978-0521648356.

Sludskii, A. A. (1992). "Tiger Panthera tigris Linnaeus, 1758" (https://archive.org/stream/mammalsofsov221

992gept#page/94/mode/2up) . In Heptner, V. G.; Sludskii, A. A. (eds.). Mlekopitajuščie Sovetskogo Soiuza.
Moskva: Vysšaia Škola [Mammals of the Soviet Union]. Vol. II, Part 2. Carnivora (Hyaenas and Cats)
(Second ed.). Washington DC: Smithsonian Institution and the National Science Foundation. pp. 95–202.
ISBN 978-90-04-08876-4.

External links

Media related to Panthera tigris (category) at Wikimedia Commons

Data related to Panthera tigris at Wikispecies

Quotations related to Tigers at Wikiquote

Tigers travel guide from Wikivoyage

"Tiger Panthera tigris" (http://www.catsg.org/index.php?id=124) . IUCN/SSC Cat Specialist Group.

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