ANNALES HISTORICO-NATURALES MUSEI NATIONALIS HUNGARICI

Volume 98 Budapest, 2006 pp. 185–280.

An annotated list of Imre Frivaldszky’s publications

and the species-group and infraspecies names proposed by him
for plants and animals (Regnum Plantare and Animale)

ZS. BÁLINT1 & S. ABADJIEV2

1
Department of Zoology, Hungarian Natural History Museum
H-1088 Budapest, Baross utca 13, Hungary. E-mail: [email protected]
2
81 Lyuben Karavelov Street, 1000 Sofia, Bulgaria. E-mail: [email protected]

Abstract – A list of 13 publications of IMRE FRIVALDSZKY containing descriptions or indications to

binomens attributing the authorship to himself is presented. This is followed by a list of 296 intro-
duced names of plants (104) and animals (192). Each entry includes the species-group name, the orig-
inal combination quoted from the original publication, the type locality, and taxonomic notes. With
12 figures.

Key words – Plantare, Animale, Aves, Gastropoda, Coleoptera, Lepidoptera, bibliography, nomen-
clature, taxonomy, Hungary, Slovakia, Romania, Balkans, Asia Minor.

INTRODUCTION

It is not enough to stress again and again that the natural objects originated
from the collections of IMRE FRIVALDSZKY (1799–1870) (Fig. 1) belong to the
most important historical items of the Hungarian Natural History Museum (Buda-
pest, Hungary) (HNHM). They represent or provide clues for evidence to the stu-
dents, who want to study the period of pioneer floristic and faunistic explorations
carried out in the Carpathian Basin, in the Balkans and in the western part of
Anatolia (BÁLINT 2002).
IMRE FRIVALDSZKY was an aid custodian of “Camera Naturae et Artis Pro-
ductorum” of the National Museum of Hungary (= Magyar Nemzeti Múzeum) with
JOHANNES SALAMON PETÉNYI (1799–1855). The botanist JOHANNES SADLER
(1791–1849) was the senior custodian. When SADLER died, IMRE FRIVALDSZKY
became the appointed custodian in 1849, but only for a short term, because JÁNOS
FRIVALDSZKY (1822–1895), his relative from the side of his father, became
SADLER’s successor, and later the first general director of the Natural History Cab-
inet. Both FRIVALDSZKYs, who became ordinary members of the Hungarian Acad-
emy of Sciences (in the time of I. FRIVALDSZKY still named as “Magyar Tudós

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Társaság” = Society of Hungarian Scientists), left a grand legacy: I. FRIVALDSZKY

as botanist, zoologist and language reformer, whilst J. FRIVALDSZKY as a zoologist
in general, who focused his activities on the insect order Coleoptera. The present
paper concentrates only on the activity of the senior FRIVALDSZKY, strictly related
to his descriptive works on living organisms.
I. FRIVALDSZKY was the pioneer scientist in the floristic and faunistic explo-
ration of the Balkans and the western part of Anatolia, but from a zoological point
of view, he also pioneered in the less known invertebrate fauna of the Carpathian
Basin. FRIVALDSZKY preapred a detailed programme on how to study the Balkan
fauna and flora (FRIVADSZKY 1835a: 238–240). With this he applied the Magyar
Tudós Társaság (the subsequent Hungarian Academy of Sciences) for a financial
support. The application was unsuccessful. Consequently, he had to finance the
programme by other means, which was general in those times: FRIVALDSZKY cre-
ated a broad network of correspondence with many of the contemporary botanists
and zoologists, as the greatest natural historian of his time also did, just to mention
HENRY WALTER BATES (1825–1892), the great explorer of the Amazonian low-
land. Specimens of desiccated plants and animals thought to represent new discov-
eries were sent for exchange or for selling under names he proposed. This was a
general practice, as we know, among naturalists, causing difficulties already at that
time (cf. CHAINEY 2005: 288–289). This is also testified by the notes we have
found in The Natural History Museum (London, UK) (NHM) as item nos 46 and 52
(1846) in the “Register of Entomology (British Museum), volume 4”, pp. 104–147:
“Purchased from Dr. Frivaldszky, Bill 53, The insects from Turkey and Hungary
on this and the following pages were named by Dr Frivaldszky. The remainder of
Dr Frivaldszky’s entered under 46.52.” (no. 46) and “Brought of Dr. Frivaldszky
Named by Dr. Frivaldszky; Cont. from 46.36.” (no. 52). In this inventory there are
altogether seven entries indicating that the material was purchased from I. FRI-
VALDSZKY as follows: 1843: no. 81; 1846: nos 36, 46, 52, 54, 65; 1848: no. 118.
Accordingly, DOUBLEDAY (1847, 1848) catalogued all these material under FRI-
VALDSZKY’s authority; and indeed, we could locate many butterfly specimens
originated from I. FRIVALDSZKY in the collections of the NHM (cf. BÁLINT 1999,
BÁLINT & OLIVIER 2001).
The network of I. FRIVALDSZKY was really wide. In the Archives of the Uni-
versity Museum, Oxford (United Kingdom) one copy of the published catalogue of
his collection (FRIVALDSZKY 1834) can be found alongside with several letters he
addressed to FREDERICK WILLIAM HOPE (1797–1862) (BÁLINT, unpublished).
Many specimens of beetles are still existing in the Hope Entomological Collec-
tions (BÁLINT, unpublished; cf. SMITH 1986: 76 and 120). Similarly, in the collec-
tions of the Muséum National d’Historie Naturelle, Paris (France) we have found

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 An annotated list of Imre Frivaldszky’s plant and animal names 187

Fig. 1. IMRE FRIVALDSZKY probably in the years when he was elected as ordinary member of the
Magyar Tudós Társaság (today Hungarian Academy of Sciences) (anonymous, original in the collec-
tion of the Hungarian Natural History Museum)

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numerous butterfly specimens sent by I. FRIVALDSZKY to ACHILLE DEYROLLE

(1813–1865), a well-known trader of natural history objects of his time (BÁLINT,
unpublished). According to the correspondence of FRIVALDSZKY and many
sources of the literature, he was in even closer relationship with several German
contemporary merchants or naturalists (BÁLINT & OLIVIER 2001), so it can be pre-
sumed that important material originating from I. FRIVALDSZKY is hiding in vari-
ous German public collections. All this material belongs to the first natural history
samples from the Balkans and the western part of Anatolia, still exotic for Europe-
ans in those times.
I. FRIVALDSZKY proposed many species-group names for animals and plants,
and also some infraspecies-group names for plants in his publications. Some of
them are mere redescriptions of taxa described by classical authors, such as CARL
VON LINNÉ (1707–1778) or JACOB HÜBNER (1761–1826). However, many of
these taxa are formally described in his publications, therefore, they are available
under the authorship of I. FRIVALDSZKY. Another large portion of the names were
introduced as nomina nuda. A great percentage of these names remained undes-
cribed by him, or became available via the authorship of the contemporary bota-
nists or zoologists whom I. FRIVALDSZKY corresponded with (cf. BÁLINT &
OLIVIER 2001 for the butterflies). Consequently, syntypical specimens represent-
ing taxa described by I. FRIVALDSZKY exist unrecorded in the general collections
of numerous museums (cf. BÁLINT 1999). Similarly, numerous specimens which
are under a name proposed by I. FRIVALDSZKY, but described by someone else,
most probably are syntypical or originate from the sample of the type material;
therefore they are the best candidate for neotype designation in a taxonomic revi-
sion (BÁLINT & OLIVIER 2001).
This paper presents an annotated list of all the publications of I. FRIVALDSZKY,
which includes scientific names and lists of all the species-group and infraspecific
names proposed by him for animal and plant species. It has the following aims: (1)
to provide a brief overview of I. FRIVALDSZKY’s descriptive works and to present a
brief account of them for the wider public; (2) to enable that syntypical material of
taxa described by I. FRIVALDSZKY can be more easily located and catalogued not
only in the HNHM, but also in other public collections; (3) to help locating and cat-
aloguing in the HNHM as important “historical” specimens; of those originating
from I. FRIVALDSZKY, which served as type material of taxa described by other au-
thors under the names proposed by I. FRIVALDSZKY, and finally (4) to provide
clues for collecting sites or origin of the type material as precisely as possible, be-
cause this information is crucial for a taxonomic revision, when names proposed
by I. FRIVALDSZKY are involved.
The present paper cannot aim at precisely identifying the publication date of
his papers as the general HNHM Library is in constant move, because of the reset-
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 An annotated list of Imre Frivaldszky’s plant and animal names 189

tlement of the whole institute (cf. MATSKÁSI 2002). This makes intense and de-
tailed bibliographic studies almost impossible. However, it is hoped that it can be
completed in the near future. Similarly, detection and revision of all the species-
group names listed is beyond the purpose of the present work. This task needs con-
siderable knowledge in the taxonomy and science history of the groups involved.
However, our work is the first step for a final revision of all these names and of their
type material, which is planned to be published in a single volume under the title
“Animal and plant names proposed by IMRE FRIVALDSZKY: Their nomenclature
and taxonomic history” in coorperation of the broader staff of our institutes.
Consequently, this paper does not give a full list of IMRE FRIVALDSZKY pub-
lished work, as he also published faunistical surveys in a strict sense (FRIVALDSZKY
1854, 1863), but he was also involved in applied entomology (FRIVALDSZKY 1848),
not to mention his most well-known doctoral thesis (FRIVALDSZKY 1823) and an
extensive catalogue published for mercantile purposes (FRIVALDSZKY 1834).
Moreover, I. FRIVALDSZKY’s manuscript legacy is enormous: besides all the docu-
mentation of the expeditions he organized (accounts, diaries, letters and passes), in
the archives of the HNHM there are manuscripts of several books of high historical
value. The scientific elaboration of these documents is also beyond the scope of the
present study.

MATERIALS AND METHODS

All the publications of IMRE FRIVALDSZKY containing descriptions of animals and plants with
formal establishment of scientific names or nude usage of Latin binomens with indication to I.
FRIVALDSZKY as author are listed in a chronological order. I. FRIVALDSZKY’s names are given al-
ways according to the cited publications. The papers were published in German, Hungarian and
Latin. Therefore, the English translation of the titles are provided by us between square brackets if the
original is in Hungarian. As the works of I. FRIVALDSZKY are very rare, we present their HNHM Li-
brary inventory number or other location if the original of the published item cannot be found in Bu-
dapest. We mention here that all the papers discussed can be found in a photocopied version in the
“Frivaldszky Imre” file of the HNHM Lepidoptera reprint collection. In the closing note a brief de-
scription of the publication is given for informing the reader unfamiliar with the works of I.
FRIVALDSZKY, but interested in the animals or plants he worked with.
Species-group names proposed by IMRE FRIVALDSZKY were compiled from his published pa-
pers and books. The compiled names are given according to their original scripts and spellings. The
names are grouped in two large kingdoms: plants and animals. Therein the names are listed in alpha-
betical order according to their generic name of the original combination in the case of plants, since
all the involved taxa are anthophytous. The animals are grouped according the three phyla
Arthropoda, Mollusca and Vertebrata, also in alphabetical order. As a main entry for the spe-
cies-group names, the first published binomen is given always with reference to the family (in brack-
ets), where the taxon actually belongs.

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References to binomens are listed chronologically as they appear in the publications. The spell-
ings of the names are often different, altered by I. FRIVALDSZKY himself or they are original lapses.
After the Latin binomen we cite also the abbreviation of the author’s name, which in certain cases is
not FRIVALDSZKY. If this happens we always note it for the reader who is familiar with the group for
future revision (see the below).
I. FRIVALDSZKY was very inconsistent, and sometimes even cryptic, in abbreviating his own
name, as he indicated certain taxa described by CHRISTIAN FRIEDRICH FREYER (1794–1888) as
“Fr”, but he often used the same abbreviation for indicating himself as the author, or he put only
“Frivald.” or “Frivaldszky for taxa described jointly by I. and J. FRIVALDSZKY”.
If a scientific name occurs with a common Hungarian name, either preceding or subsequent to
it, it is also given because in many cases these names provide a clue for the identity of the taxon. All
the citations refer precisely to the page(s) of the given publication(s) and repeat verbatim the words,
which contain indications to the type locality. In these cases translation of the originally Hungarian or
Latin words are provided [in square brackets]. As a closing remark, there is an indication to the status
of the type material deduced from these sources. In many cases I. FRIVALDSZKY indicated a general-
ised locality for the material he based his descriptions on, but subsequently he often presented a more
precise locality of the given taxon. We understand most of these subsequent actions of I.
FRIVALDSZKY as type locality clarifications in the sense of the International Commission of Zoologi-
cal Nomenclature (1999) (abbreviated as ICZN), Recommendation 76A.
The geographical names used by IMRE FRIVALDSZKY are almost totally altered. He used Hun-
garian or Latin scripts of localities, which were situated in the regions of the Hungarian Kingdom (in-
cluding present-day Croatia, the western part of Romania, the northern part of Serbia–Montenegro
and Slovakia) or the Ottoman Empire (including present-day Bulgaria, Greece, Macedonia and west-
ern Turkey). Therefore, a Gazetteer is provided for all the names with their modern equivalents
emerging in connection with the names I. FRIVALDSZKY proposed.

PUBLICATIONS OF IMRE FRIVALDSZKY CONTAINING

FORMAL DESCRIPTIONS OR INDICATIONS TO BINOMENS
ATTRIBUTING THE AUTHORSHIP TO HIMSELF

FRIVALDSZKY, IMRE 1835a: Közlések a’ Balkány’ vidékén tett természettu-

dományi utazásról. [Notes on the natural history journey taken in the Balkan re-
gion.] – A’ Magyar Tudós Társaság Évkönyvei. Második kötet. Budán. Magyar
Királyi Egyetem nyomtatása. Pp. 235–276, pls I–VII. – Inventory number: Ad
1127. Written in Hungarian with extensive lists of plants and animals in binomen
introducing numerous nomina nuda. This is the first account of the Balkan expedi-
tions with the establishment of the plant genus Haberlea along with the diagnoses
in Latin and descriptions in Hungarian of four plant species (including the type
species of Haberlea) (Figs 2–4), 24 species of Coleoptera, nine Lepidoptera and
three Gastropoda.
FRIVALDSZKY VON FRIVALD, EMRICH 1835b: Antrag der im Jahre 1833 und
1834 in der europäischen Türkey, namentlich in Rumelien, auf dem Balkan und
Rhodope gesammelten wildwachsenden Pflanzen. Von Hinke und Manolesko. –

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 An annotated list of Imre Frivaldszky’s plant and animal names 191

Flora, Intelligenzblatt zur allgemeinen botanischen Zeitung, Regensburg 1: 53–59.

– Inventory number: P. 12. Written in German, and published for mercantile pur-
poses, introducing many nomina nuda and misapplications of formerly established
names (cf. HAMPE 1837).
FRIVALDSZKY, E.(mrich) 1835c: Succinctae diagnoses specierum plantarum
novarum europaeo-turcicarum in catalogo meo occurrentium. – Flora, Intelligenz-
blatt zur allgemeinen botanischen Zeitung, Regensburg 1: 331–366. – Inventory
number: P. 12. Written exclusively in Latin, introducing one generic and 24 spe-
cies-group names including the ones which have already been published.
FRIVALDSZKY, E.(mrich) 1836a: Succinctae diagnoses specierum plantarum
novarum anno 1835 in Turcia europaea collectarum. – Flora oder allgemeine bo-
tanische Zeitung, Regensburg 19(2): 433–440. – Inventory number: P. 12. Written
exclusively in Latin, introducing 30 species-group names providing formal de-
scriptions of many nomina nuda introduced in previous papers.
FRIVALDSZKY VON FRIVALD, EMRICH 1836b: Antrag getrockneter türkisch-
europäischer Pflanzen. – Flora, Intelligenzblatt zur allgemeinen botanischen Zei-
tung, Regensburg 2: 19–28. – Inventory number: P. 12. Written in German and
published for mercantile purposes with a long list of scientific binominal names,
introducing many nomina nuda and misapplications of formerly established names
(cf. A. HAMPE 1837).
FRIVALDSZKY VON FRIVALD, EMRICH 1837a: Neue Kaefer, Falter und Schen-
ken aus den balkanischen Gebirgen. – Faunus, München 1: 84–93. – Inventory
number: Ad 2. This is a compilation of the first Balkan report published in Hungar-
ian repeating faunistic lists and species diagnoses of the first Balkan report. Proba-
bly it was plainly taken from the publication itself and not seen by FRIVALDSZKY
as the main headings of the lists are Hungarian, a curious manner of a journal pub-
lished in Germany, and the reference to the source and the main body of the text it-
self is full of misspellings.
FRIVALDSZKY IMRE 1837b: Balkány vidéki természettudományi utazás.
[Natural history journey taken in Balkan region.] – A’Magyar Tudós Társaság
Évkönyvei. Harmadik kötet. Budán. A ’Magyar Királyi Egyetem’ betűivel. Pp.
156–184, pls I–VIII. – Inventory number: Ad 1127. This is the second report of the
Balkan expeditions providing a brief account of the collecting sites and the discov-
ered plants and animals introducing again nomina nuda. The paper itself contains
the diagnoses in Latin and descriptions in Hungarian of six plants, three lepido-
pterous and nine coleopterous insects and one bird, the Collared Dove Streptopelia
decaocto (FRIVALDSZKY, 1835).
FRIVALDSZKY IMRE 1840: Természettudományi utazás a’ Balkány’ vidékén
[Natural history journey in the region of the Balkans]. – A Magyar Tudós Társaság

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Fig. 2. Haberlea rhodopensis FRIVALDSZKY, 1835 (HNHM), voucher of Herbarium HAYNALD

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 An annotated list of Imre Frivaldszky’s plant and animal names 193

Fig. 3. Haberlea rhodopensis FRIVALDSZKY, 1835 (HNHM), the syntypical specimen selected as
lectotype by A. MEYER

Fig. 4. Haberlea rhodopensis FRIVALDSZKY, 1835 (HNHM), FRIVALDSZKY’s identification label in

the lower right corner of the voucher

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Évkönyvei. Negyedik kötet. Budán. A’ Magyar Királyi Egyetem’ betűivel. Pp.

195–207, pls I–XII. Inventory number: Ad 1127. – This is the third report of the
Balkan expeditions, which is dedicated to diagnose in Latin and describe in Hun-
garian twelve plant species new to science. The coloured lithographic plates be-
long to the most gorgeous ones ever produced in Hungary.
FRIVALDSZKY IMRE 1845: Rövid áttekintése egy természetrajzi utazásnak, az
európai Törökbirodalomban, egyszersmind nehány a közben újdonnat fölfedezett
állatnak leírása. [Brief overview of a natural history journey taken in the European
part of the Ottoman Empire, supplemented with the description of some newly dis-
covered animals.] – A Királyi Magyar Természettudományi Társulat évkönyvei.
Első kötet. Pesten, nyomatott Beimel Józsefnél. Pp. 163–187, pls I–III. – Inventory
number: Ad 222. This is the conclusion of all the collecting activities and reports
FRIVALDSZKY previously managed and published. It gives a brief overview of all
the expeditions listing the new taxa found in the regions, many of them still nomina
nuda or described by other scientists, but FRIVALDSZKY considered himself the au-
thor of practically all the names. In this report he establishes the lepidopterous in-
sect genus Caloptera, and provides the Latin diagnoses and Hungarian descrip-
tions of twelve coleopterous and three lepidopterous insects (including the type
species of Caloptera) (Figs 5–6).
FRIVALDSZKY IMRE 1856: Magyarország és Erdély édesvizi és földi puhá-
nyai. [The fresh water and land arthropods of Hungary and Transylvania.] – Ma-
gyar Tudományos Akadémiai Értesítő 16 (2): 1–69. – Inventory number: Ad 1470.
This is the published version of the oral presentation dated February 4, 1856. It is
entirely Hungarian providing an overview of the entire gastropod fauna of Hun-
gary and Transylvania with a brief overview of the previous activities. It specifi-
cally treats taxa considered to be endemic. FRIVALDSZKY considers himself to be
the author of four taxa. He presents proper diagnoses only to one of them (Helix
tiraria). The other three remained as nomina nuda of FRIVALDSZKY. All these taxa
had previously been described by the contemporary malacologist EMIL ADOLF ROSS-
MÄSSLER (1806–1868) in his series of “Iconographie der Land- und Süsswasser
Mollusken Berüchsichtingung der europäischen noch nich abgebildete Arten” on
the basis of specimens sent by FRIVALDSZKY as species under the authorship of
FRIVALDSZKY.
FRIVALDSZKY, EMERICH und FRIVALDSZKY, JOHANN 1857: Drei neue Grot-
ten-Käfer aus Ungarn. – Verhandlungen des zoologischen-botanischen Vereins in
Wien 7: 44–46. – Inventory number: Eg 1257. This is the only joint publication of
the two FRIVALDSZKYs providing descriptions of three coleopterous insects living
in caves. The language of the paper is German with Latin diagnoses for the taxa
they proposed.

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 An annotated list of Imre Frivaldszky’s plant and animal names 195

Figs 5–9. 5–6 = Caloptera ocellata FRIVALDSZKY, 1845 (HNHM): 5 = male (left) and female (right)
syntypical specimens in dorsal view, 6 = corresponding labels of the male syntype (upper) and the fe-
male (lower) syntype. 7–9 = Haplolophus neglectus FRIVALDSZKY, 1865 (HNHM): 7 = HNHM
syntype specimens in dorsal view, 8 = the syntype erroneously labelled as “Holotype” in lateral view,
9 = corresponding labels of the syntype labelled as “Holotype”

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Fig. 10. The manuscript “Die Schmetterlinge Ungarn’s eine freie Uebersetzung aus dem unga-
rischen” (HNHM: Ec 61), title page

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 An annotated list of Imre Frivaldszky’s plant and animal names 197

FRIVALDSZKY IMRE 1859: Hazánk faunájára vonatkozó adatok, és a Puszta-

peszéri erdő [Data on the fauna of our motherland, and the forest of Puszta-peszér.]
– A Magyar Tudós Társaság Évkönyvei, Buda 9: 19–28. – Inventory number: Ad
1127. This is the published form of an oral presentation given in the Hungarian
Academy of Sciences. It is entirely written in Hungarian, but again some nomina
nuda are introduced in binominal Latin. The paper is a faunistic survey of the leg-
endary hunting place of Melanargia suwarovius formerly known to occur in scat-
tered colonies on the plain around Pest. The forest of Pusztapeszér was one of the
most famous ones (FOUNTAINE 1980: 106–109). The aim of the presentation was
to describe meticulously the early stages and life history of “Autophila Pannonica,
Friv.”, a noctuid moth species described by FREYER from “Unterungarn” as “Noct.
Pannonica (Frivaldszky)” on the basis of material sent to him by FRIVALDSZKY
(FREYER 1842: 67, pl. 330, figs 3–4). FRIVALDSZKY considered himself to be the
author of the name.
FRIVALDSZKY IMRE 1865: Jellemző adatok Magyarország faunájához [Char-
acteristic data of the fauna of Hungary.] – A Magyar Tudományos Akadémiai
Évkönyvei, Pest 9 (4): 1–274 pp., 1–13 pls. – Inventory numbers: B 501, B 502, Eg
574. This is the main work of FRIVALDSZKY dealing with the whole fauna of all the
countries belonging at that time under the crown of Hungary. Thus, it does not deal
with present-day Croatia and Transylvania, but with the faunas of present-day up-
per Serbia (Vojvodina) and Slovakia, as well as the western-border region of Ro-
mania (Banat and Partium). The body of the text is entirely in Hungarian and con-
tains two parts devoted to faunistics and systematics. The latter part is actually a
more detailed description of the animal species considered by FRIVALDSZKY to be
characteristic for the Hungarian fauna. It provides the formal descriptions of nu-
merous coleopterans including the genera Haplolophus (Figs 7–9) and Myceto-
mychus and their type species, plus one lepidopteran (Orgyia ericae intermedia).
In the special Lepidoptera Library of the HNHM under the entry Ec. 61 there
is a manuscript with the title “Die Schmetterlinge Ungarn’s eine freie Ueberset-
zung aus dem ungarischen” dated as “Ofen, 25tn/4, 1868” which is the German
translation of the published Hungarian text specifically dealing with certain butter-
fly and moth species. This was probably done by FRIVALDSZKY himself, as the
style of the handwriting is similar to his own, and every page is signed with his ini-
tial “f” (Figs 10–12).
Interestingly, an extensive summary of this publication was published in Ger-
man, which cannot be considered to be authoritative as it is written “nach Fri-
valdszky” (based on FRIVALDSZKY) (ANONYMOUS 1868). It was most probably
written by LUDWIG AIGNER (alias ABAFI-AIGNER LAJOS), a book-handler, a con-
temporary of FRIVALDSZKY, but subsequently became a prominent lepidopterist,

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Fig. 11. The manuscript “Die Schmetterlinge Ungarn’s eine freie Uebersetzung aus dem unga-
rischen” (HNHM: Ec 61), page I

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 An annotated list of Imre Frivaldszky’s plant and animal names 199

who was actually the publisher. He published a series of entomological observa-

tions taken verbatim from the manuscript diaries of FRIVALDSZKY possessed by
the HNHM (ABAFI-AIGNER 1898), and from the diaries of the ANKER brothers he
purchased (AIGNER 1900). FRIVALDSZKY’s manuscript diaries and letters, ac-
counts, etc. and the reports of the collecting trips sent back by ANDRÁS FÜLE and
JOSEPH HINKE are well preserved and catalogued in the HNHM archives, but the
legacy of the ANKER brothers has disappeared.

Fig. 12. The manuscript “Die Schmetterlinge Ungarn’s eine freie Uebersetzung aus dem unga-
rischen” (HNHM: Ec 61), last (xx) page

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SPECIES-GROUP AND INFRASPECIFIC NAMES

PROPOSED BY IMRE FRIVALDSZKY

REGNUM PLANTARE

Achillea buglossis (Asteraceae)

Achillea buglossis FRIVALDSZKY – FRIVALDSZKY 1836a: 433, “Hab. in subalpinis ad Carlova” [In-
habits the subalpine region near Karlovo] (description).

Remarks – The species was described from on the basis of an unstated number
of syntypes from the subalpine region near Karlovo.

Achillea chrysocoma (Asteraceae)

Achillea chrysocoma FRIVALDSZK. – FRIVALDSZKY 1835b: 58, “Rumelia” (no description).

Remarks – Nomen nudum.

Achillea grandiflora (Asteraceae)

Achillea grandiflora FRIVALDSZKY – FRIVALDSZKY 1836a: 433, “Hab, in mont. Calophier. Julio”
[Inhabits Kaloferska Planina, July] (description).
Terepélylevelű cziczikóró (Achilea grandiflora) – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Re-
gion of Karlovo] (redescription).
Terebélylevelü cziczkóró (Achillea grandifolia FR.) –FRIVALDSZKY 1845: 168, “Karlova vidéke”
[Region of Karlovo].

Remarks – The species was described from an unstated number of syntypical

specimens collected in the Kaloferska Planina. It was subsequently redescribed by
FRIVALDSZKY in more details from the same region.

Acinos acuminatus (Lamiaceae)

Acinos acuminatus FRIVALDSZ. – FRIVALDSZKY 1835b: 54, “Rumelia” (no description).

Acynos acuminatus FRIVALDSZKY – FRIVALDSZKY 1835c: 332, “Habit. In Rumelia” [Inhabits
Rumelia] (description).
Acynos acuminatus FRIVALDSZKY – FRIVALDSZKY 1836b: 18, “Rumelia”.

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 An annotated list of Imre Frivaldszky’s plant and animal names 201

Hegyeslevelű csobor. Acynos acuminatus. FRIV. – FRIVALDSZKY 1840: 166, “Philipopoly vidéke”;
195, “Terem Karlova ‘s a’ Rhodope hegység’ vidékein Rumeliában” [Grows in the regions of
Karlovo and Rhodope Mountains, in Rumelia], pl. I. (redescription).

Remarks – The name was introduced as nomen nudum, but the species was
subsequently described on the basis of an unstated number of syntypical speci-
mens originating from “Rumelia”. It was redescribed and illustrated, restricting the
type locality as “regions of Karlovo”.

Acynos erectus (Lamiaceae)

Acynos erectus FRIVALDSZKY – FRIVALDSZKY 1835b: 55, “Rumelia” (no description).

Acynos erectus FRIVALDSK. – FRIVALDSZKY 1835c: 333, “Hab. In Rumelia” [Inhabits Rumelia] (de-
scription).
Acynos erectus FRIVALDSZKY – FRIVALDSZKY 1836b: 18, “Rumelia”.
Merevény csobor. Acynos erectus. FRIV. – FRIVALDSZKY 1840: 196, “Virágzik Karlova’ ‘s Filipopol
vidékin, juliusban” [Blossoming in July, in the regions of Karlovo and Plovdiv], pl. II.
(redescription).
Merevény csobor (Acynos erectus FRIV.) – FRIVALDSZKY 1845: 166, “Philipopoly vidéke” [Region
of Plovdiv].

Remarks – The species was described from an unstated number of syntypical

specimens originating from “Rumelia”. It was redescribed and illustrated subse-
quently, providing more precise type locality.

Ajuga salicifolia (Lamiaceae)

Fűzlevelü kacskanyak. Ajuga salicifolia. SCHREBER –FRIVALDSZKY 1837b: 169, “Balkány’

hegyeinek aljasában, junius és juliusban.” [At the lower regions of Stara Planina, in June and
July], pl. V. (description).
Fűzlevelű kacskanyak (Ajuga salicifolia SCHREBER) –FRIVALDSZKY 1845: 165, “Szlivno
(Selimno)… Haemus déli s éjszaki vidékei” [Northern and southern parts of Sliven and the
Stara Planina] (type locality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens from the lower regions of the Stara Planina. Subsequently the type lo-
cality was restricted to the mountain “Haemus” [Stara Planina]. FRIVALDSZKY in-
dicated the botanist JOHANN CHRISTOPH DANIEL VON SCHREBER (1739–1810) as
the author of the taxon.

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Aira triflora (Poaceae)

Aira triflora FRIVALDSZKY – FRIVALDSZKY 1836a: 433, “Hab. in subalpinis ad Carlova” [Inhabits
the subalpine regions near Karlovo] (description).

Remarks – The species was described from an unstated number of syntype

specimens originating from the subalpine regions near Karlovo.

Arctium atrorubra (Asteraceae)

Arctium Lappa L. var. atrorubra FRIV. – FRIVALDSZKY 1836b: 23, “Rumelia” (no description).

Remarks – Nomen nudum.

Arenaria saxifraga (Caryophyllaceae)

Arenaria saxifraga FRIVALDSZKY – FRIVALDSZKY 1836a: 434, “Hab. in Rumelia” [Inhabits

Rumelia] (description).
Arenaria saxifraga Friv. – FRIVALDSZKY 1836b: 23, “Rumelia”.
Kőtör homokhúr (Arenaria saxifraga), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke”
[Region of Karlovo] (type locality restriction).
Kötör homokhúr. Arenaria saxifraga FRIV. – FRIVALDSZKY 1840: 199, “Terem Karlova’ havasain.
Virágzik junius és juliusban” [Grows in the high mountains of Karlovo, blossoming in June
and July], pl. V. (redescription).
Kőtör homokhúr (Arenaria saxifraga FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo].

Remarks – The species was described from an unstated number of syntypical

specimens from “Rumelia”. It was redescribed and illustrated subsequently, plus
the type locality was restricted as the high mountains of “Karlova” [Karlovo].

Asperula tubiflora (Rubiaceae)

Csővirági müge. Friv. Asperula tubiflora. FRIV. – FRIVALDSZKY 1837b: 163, “Hortiát” (no descrip-
tion).
Csővirágú müge (Asperula tubiflora Fr.) – FRIVALDSZKY 1845: 169, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

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Astrantia elatior (Apiaceae)

Astrantia elatior FRIVALDSZKY – FRIVALDSZKY 1836a: 434, “Hab. in Rumelia” [Inhabits Rumelia]
(description).
Astrantia elatior FRIVALDSZKY – FRIVALDSZKY 1836b: 23, “Rumelia”.
Nyúlánk zápócza (Astrantia elatior FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Regions of
Karlovo] (type locality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens from “Rumelia”. Subsequently the type locality was restricted as “re-
gions of Karlovo”.

Authemis contorta (Asteraceae)

Tekercs montika. FRIV. Authemis contorta. FRIV. – FRIVALDSZKY 1837: 163, “Hortiát” (no descrip-
tion).
Tekercs montika (Anthemis contorta FR.) – FRIVALDSZKY 1845: 169, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Avena glabrescens (Poaceae)

Sima zab. FRIV. Avena glabrescens. FRIV. – FRIVALDSZKY 1837b: 163, “Hortiát” (no description).
Sima zab (Avena glabrescens FR.) –FRIVALDSZKY 1845: 169, “Macedonia hegyei, s bérczei között”
[In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Berberis microphylla (Berberidaceae)

Aprólevelű borboja. Berberis mycrophilla. FRIVALDSZKY – FRIVALDSZKY 1837b: 170, pl. VI. (no
patria) (description).
Aprólevelű borbója (Berberis microphylla ROCHL.) – FRIVALDSZKY 1845: 165, “Szlivno (Selim-
no)… Haemus déli s éjszaki vidékei” [Northern and southern parts of Sliven and the Stara
Planina] (redescription, type locality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens originating from unknown place(s). Subsequently the type locality was
restricted as “Haemus” [Stara Planina], in the vicinity of “Szlivno” [Sliven]. Sub-

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sequently FRIVALDSZKY attributed the authorship of the species-group name to the

botanist ANTON ROCHEL (1770–1847).

Berteroa luteola (Brassicaceae)

Sárgálló Berteroa. Frivaldszky Berteroa luteola. FRIVALDSZKY – FRIVALDSZKY 1835a: 163, “Hortiát”
(no description).
Sárgálló berteroa (Berteroa luteola FR.) – FRIVALDSZKY 1845: 169, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).
Remarks – Nomen nudum.

Bulbocodium hastulatum (Colchicaceae)

Bulbocodium hastulatum FRIVALDSZKY – FRIVALDSZKY 1836a: 434, “Hab. in Rumelia” [Inhabits

Rumelia] (description).
Bulbocodium hastulatum FRIV. – FRIVALDSZKY 1836b: 23, “Rumelia”.
Dárdácskás földike. Bulbocodium hastulatum. FRIVALDSZKY – FRIVALDSZKY 1837b: 161, “Karlova
vidéke” [Region of Karlovo], 166, “Terem a’ Balkány’ közép hegyeinek völgyeiben.” [Grows in
the valleys of the central Stara Planina], pl. II. (redescription).
Dárdácskás földike (Bulbocodium hastulatum FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke”
[Region of Karlovo] (type locality restriction).
Remarks – The species was described from an unstated number of syntypical
specimens from “Rumelia”. Subsequently it was redescribed and illustrated, plus
the type locality was restricted as “region of Karlovo”.

Bromus villosis (Poaceae)

Bromus squarrosus var. spiculis villosis FRIVALDSZKY – FRIVALDSZKY 1836b: 18, “Rumelia”
(nomen nudum).

Remarks – Nomen nudum.

Bupleurum apiculatum (Apiaceae)

Bupleurum apiculatum FRIVALDSZK. – FRIVALDSZKY 1835b: 56, “Rumelia” (no description).

Bupleurum apiculatum FRIVALDSZK. – FRIVALDSZKY 1835c: 335, “Hab. in Rumelia” [Inhabits
Rumelia] (description).
Bupleurum apiculatum FRIVALDSZK. – FRIVALDSZKY 1836b: 18, “Rumelia”.

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Szálkás szingallér. Bupleurum apiculatum FRIV. – FRIVALDSZKY 1840: 200, “Karlova’ vidékin,
juniusban” [Region of Karlovo, in June], pl. VI (redescription, type locality restriction).
Szálkás szingallér (Bupleurum apiculatum FR.) – FRIVALDSZKY 1845: 166, “Philipopoly vidéke”
[Region of Plovdiv].

Remarks – Introduced as nomen nudum, subsequently it was described from

an unstated number of syntypical specimens from “Rumelia”. It was redescribed
and illustrated, plus the type locality was restricted as “Region of Karlovo”.

Campanula pusilla (Campanulaceae)

Campanula alpina var. pusilla FRIVALDSZKY – FRIVALDSZKY 1836b: 18, “Rumelia” (no descrip-
tion).

Remarks – Nomen nudum.

Campanula expansa (Campanulaceae)

Campanula expansa FRIVALDSZKY – FRIVALDSZKY 1836a: 434, “Hab. in mont. Calophier. Floret
Julio” [Inhabits Kaloferska Planina. Blossoming in July] (description).
Terjedt csengetyűke (Campanula expansa), FRIV. – FRIVALDSZKY 1837b: 161, “Karlova vidéke”
[Region of Karlovo].
Terjedt csengetyűke (Campanula expansata FRIV.) – FRIVALDSZKY 1845: 168, “Karlova vidéke”
[Region of Karlovo].

Remarks – The species was described from an unstated number of syntypical

specimens originating from the Kaloferska Planina.

Campanula lanata (Campanulaceae)

Campanula lanata FRIVALDSZKY – FRIVALDSZKY 1836a: 434, “Hab. in Rumelia” [Inhabits

Rumelia] (description).
Szöszös csengetyűke (Campanula lanata), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke”
[Region of Karlovo], 165, “Terem a’ Balkánynak Calophir hegye’ vidékén” [Grows in the
Kaloferska Planina of the Stara Planina], pl. I. (redescription).
Szöszös csengetyűke (Campanula lanata) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo].

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Remarks – The species was described from an unstated number of syntypical

specimens from “Rumelia”. Subsequently it was redescribed and illustrated, plus
the type locality was restricted to the Kaloferska Planina.

Campanula sparsa (Campanulaceae)

Zilált csengetyűke. Campanula sparsa FRIV. – FRIVALDSZKY 1840: 201, “Nő Karlova hegység’
árnyas völgyeiben” [Grows in the sheltered valleys of the mountain near Karlovo], pl. VII. (de-
scription).

Remarks – The species was described from an unstated number of syntypical

specimens originating from the valleys of the mountain near Karlovo.

Centaurea affinis (Asteraceae)

Centaurea affinis FRIVALDSZKY – FRIVALDSZKY 1836a: 435, “Hab. in Rumelia” [Inhabits Rumelia]
(description).
Centaurea affinis FRIVALDSZKY – FRIVALDSZKY 1836b: 24, “Rumelia”.

Remarks – The species was described from an unstated number of syntypical

specimens originating from “Rumelia”.

Centaurea albiflora (Asteraceae)

Centaurea virgata WILLD. var. albiflora FRIV. – FRIVALDSZKY 1836b: 24, “Rumelia” (no descrip-
tion).

Remarks – Nomen nudum.

Centaurea albotomentosa (Asteraceae)

Centaurea ciliata var. albo-tomentosa FRIVALDSZK. – FRIVALDSZKY 1835b: 55, “Rumelia” (no de-
scription).
Centaurea ciliata var. albo-tomentosa FRIVALDSZK. – FRIVALDSZKY 1835c: 334, “Hab. prope
Philippopolim” [Occurs near Plovdiv] (description).
Centaurea ciliata var. albotomentosa FRIV. – FRIVALDSZKY 1836b: 19, “Rumelia”.

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Remarks – Introduced as nomen nudum, but it was subsequently described

from an unstated number of syntypical specimens originating from the vicinity of
Plovdiv.

Centaurea langii (Asteraceae)

Láng csüküllő. Friv. Centaurea Langii. FRIV. – FRIVALDSZKY 1837: 163, “Hortiát” (no description).

Remarks – Nomen nudum.

Centaurea carlovensis (Asteraceae)

Centaurea Karlovensis FRIV. – FRIVALDSZKY 1836b: 24, “Rumelia” (no description).

Karlovai csüküllő (Centaurea carlovensis FR.) – FRIVALDSZKY 1845: “Karlova vidéke” [Regions of
Karlovo] (no description).

Remarks – Nomen nudum.

Centaurea napulifera (Asteraceae)

Répás csüküllő. Centaurea napulifera. ROCH. – FRIVALDSZKY 1835a: 250, “Találtatik a’ Balkány’
hegyének közép ormain, Junius és Juliusban” [Can be found on the central cliffs of the Stara
Planina, in June and July], pl. III (description).
Centaurea napulifera ROCHL. – FRIVALDSZKY 1835b: 58, “Balkan”.
Centaurea napulifera ROCHL. – FRIVALDSZKY 1835c: 356, “Hab. in Montib. Balkan” [Inhabits
Stara Planina] (redescription).
Centaurea napulifera ROCHL. – FRIVALDSZKY 1836b: 19, “Rumelia”.
Répás csüküllő (Centaurea napulifera ROCHL.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)…
Haemus déli s éjszaki vidékei” [Sliven, southern and northern regions of Stara Planina] (type
locality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens from the central cliffs of Stara Planina collected in June and July. Sub-
sequently the type locality was restricted to the vicinity of Sliven. FRIVALDSZKY
attributed the authorship of the species group name to the botanist ANTON
ROCHEL.

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Centaurea pallida (Asteraceae)

Centaurea pallida FRIVALDSK. – FRIVALDSZKY 1835b: 55, “Rumelia” (no description).

Centaurea pallida FRIVALDSZK. – FRIVALDSZKY 1835c: 333, “Hab. in Rumelia” [Inhabits Rumelia]
(description).
Centaurea ciliata pallida FRIVALDSZKY – FRIVALDSZKY 1836b: 19, “Rumelia”.

Remarks – Introduced as nomen nudum, subsequently the species was described

from an unstated number of syntypical specimens originating from “Rumelia”.

Centaurea thessalonica (Asteraceae)

Thessaliai csüküllő (Centaurea thessalonica Fr.) – FRIVALDSZKY 1845: 169, “Macedonia hegyei, s
bérczei között” [In the mountains and cliffs of Macedonia] (nomen nudum).

Remarks – Nomen nudum.

Cerastium moesiacum (Caryophyllaceae)

Cerastium moesiacum FRIVALDSZKY – FRIVALDSZKY 1836a: 435, “Hab. in Rumelia” [Inhabits

Rumelia] (description).
Cerastcum moesiacum FRIV. – FRIVALDSZKY 1836b: 24, “Rumelia”.
Moesiai madárhúr (Cerastium moesiacum), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke”
[Region of Karlovo] (type locality restriction).
Moesiai madárhúr (Cerastium moesiacum FRIV.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Re-
gion of Karlovo].

Remarks – The species was described from an unstated number of syntypical

specimens from “Rumelia”. Subsequently the type locality was restricted as “re-
gion of Karlovo”.

Cerasticum rectum (Caryophyllaceae)

Cerasticum rectum FRIVALDSZKY – FRIVALDSZKY 1836a: 435, “Hab. in Rumelia” [Inhabits

Rumelia] (description).
Cerasticum rectum FRIVALDSZKY – FRIVALDSZKY 1836b: 24, “Rumelia”.
Súgár madárhúr (Cerastium rectum), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Re-
gion od Karlovo] (type locality restriction).
Sugár madárhúr (Cerastium rectum FRIV.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo].

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Remarks – The species was described from an unstated number of syntypical

specimens from “Rumelia”. Subsequently the type locality was restricted as region
of Karlovo.

Chondrilla intybacea (Asteraceae)

Chondrilla intybacea FRIVALDSZK. – FRIVALDSZKY 1835b: 56, “Rumelia” (no description).

Chondrilla intybacea FRIVALDSZK. – FRIVALDSZKY 1835c: 335, “Hab. ad Philippopolim” [Occurs
near Plovdiv] (description).
Katáng kakics (Chondrilla intybacea FR.) – FRIVALDSZKY 1845: 166, “Philipopoly vidéke” [Region
of Plovdiv].

Remarks – Mentioned first as nomen nudum, but subsequently the species

was described from an unstated number of syntypical specimens originating from
the vicinity of Plovdiv.

Chondrilla prenanthoides (Asteraceae)

Csakkör kakics (Chondrilla prenanthoides) – FRIVALDSZKY 1845: 169, “Macedonia hegyei, s

bérczei között” [In the mountains and cliffs of Macedonia], (no description).

Remarks – Nomen nudum.

Cynosorus viridis (Poaceae)

Cynosorus echiantus WILLD. variet. Viridis FRIV. – FRIVALDSZKY 1836b: 19, “Rumelia” (no de-
scription).

Remarks – Nomen nudum.

Crocus hybernus (Iridaceae)

Crocus hybernus FRIVALDSZKY – FRIVALDSZKY 1836b: 24: “Rumelia” (no description).

Telelő sáfrány (Crocus hybernus FRIV.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo] (no description).

Remarks – Nomen nudum.

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Delphinium cinerascens (Ranunculaceae)

Delphinium junceum DC. var. cinerascens FRIVALDSZK. – FRIVALDSZKY 1835b: 56, “Rumelia” (no
description).
Delphinium junceum DC. var. cinerascens FRIVALDSZK. – FRIVALDSZKY 1835c: 335, “Hab. in
Rumelia” [Inhabits Rumelia].
Delphinium junceum var. cinerascens FRIV. – FRIVALDSZKY 1836b: 19, “Rumelia”.

Remarks – Introduced as nomen nudum, but subsequently the species was de-
scribed from an unstated number of syntypical specimens originating originating
from “Rumelia”.

Dianthus brevifolius (Caryophyllaceae)

Rövid levelű szegfű Dianthus brevifolius FRIVALDSZK. – FRIVALDSZKY 1835b: 55, “Rumelia” (no
description).
Dianthus brevifolius FRIVALDSZK. – FRIVALDSZKY 1835c: 334, “Hab. in Rumelia” [Inhabits
Rumelia] (description).
Dianthus brevifolius FRIVALDSZKY – FRIVALDSZKY 1836b: 19, “Rumelia”.
Rövid levelű szegfű. Dianthus brevifolius. FRIV. – FRIVALDSZKY 1840: 202, “Karlova havas’ alján,
juliusban” [At the foothills of the mountain near Karlovo, in July], pl. VIII. (redescription, type
locality restriction).
Rövidlevelű szegfű (Dianthus brevifolius FR.) –FRIVALDSZKY 1845: 166, “Philipopoly vidéke” [Re-
gion of Plovdiv].

Remarks – Introduced as nomen nudum, subsequently the species was de-

scribed on the basis of unstated number of syntypical specimens originating from
“Rumelia”. Later FRIVALDSZKY presented a redescription with figures. Subse-
quently the type locality was restricted as the foothill region of the mountain near
Karlovo.

Dianthus grandiflora (Caryophyllaceae)

Dianthus Armeria var. grandiflora FRN. – FRIVALDSZKY 1836b: 24, “Rumelia” (no description).

Remarks – Nomen nudum.

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Dianthus longisquamis (Caryophyllaceae)

Dianthus petraeus W. K. var longisquamis FR. – FRIVALDSZKY 1836b: 24, “Rumelia” (no descrip-
tion).

Remarks – Nomen nudum.

Dianthus minor (Caryophyllaceae)

Dianthus capitulus DC. var. minor FRIV. – FRIVALDSZKY 1836b: 24, “Rumelia” (no description).

Remarks – Nomen nudum.

Equisetum zonatum (Equisetaceae)

Equisetum zonatum FRIV. – FRIVALDSZKY 1846b: 24, “Rumelia” (no description).

Öves zsurló (Equisetum zonatum FR.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke” [Region of
Plovdiv] (no description).

Remarks – Nomen nudum.

Euphorbia pubifera (Euphorbiaceae)

Leplekes fütéj. FRIV. Euphorbia pubifera. FRIV. – FRIVALDSZKY 1837b: 163, “Hortiát” (no descrip-
tion).
Leplekes fűtej (Euphorbia pubigera FR.) – FRIVALDSZKY 1845: 169, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Euphorbia rupestris (Euphorbiaceae)

Euphorbia rupestris FRIVALDSZKY – FRIVALDSZKY 1836a: 435, “Hab. In Rumelia” [Inhabits

Rumelia] (descripition).
Euphorbia rupestris FRIVALDSZKY – FRIVALDSZKY 1836b: 24, “Rumelia”.
Szirtifűtej (Euphorbia rupestris), FRIV. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Region of
Karlovo].

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Szirti fűtej. Euphorbia rupestris. FRIV. – FRIVALDSZKY 1840: 203, pl. IX. “Virágzik májusban,
Banjo’ granithegye’ ormain” [Blossoming in May, on the cliffs of the granite mountain Banjo]
(redescription; type locality restriction).
Szirti fűtej (Euphorbia rupestris FRIV.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo].

Remarks – The species was described from an unstated number of syntypical

specimens originating from “Rumelia”. Subsequently the taxon was more pre-
cisely described and the type locality restricted as mountain “Banjo”, in the region
of “Karlovo”.

Ferula hispida (Apiaceae)

Ferula hispida FRIVALDSZK. – FRIVALDSZKY 1835b: 55, “Rumelia” (no description).

Ferula hispida FRIVALDSZK. – FRIVALDSZKY 1835c: 334, “Hab. in Rumelia” [Inhabits Rumelia]
(description).
Ferula hispida FRIVALDSZKY – FRIVALDSZKY 1836b: 20, “Rumelia”.
Érdes büdör. Ferula hispida. FRIVALDSZKY – FRIVALDSZKY 1837b: 168, “Terem Philippopol’
vidékén.” [Grows in the region of Plovdiv.], pl. IV (redescription; type locality restriction).
Érdes büdör (Ferula hispida FR.) – FRIVALDSZKY 1845: 166, “Philipopoly vidéke” [Region of Plovdiv].

Remarks – Introduced as nomen nudum, but subsequently it was described on

the basis of an unstated number of syntypical specimens originating from Rumelia.
Later the species was redescribed, and the type locality became restricted to the re-
gion of Plovdiv.

Galeopsis hispidor (Lamiaceae)

Galeopsis versicolor SCHMIDT. var. hispidor FRIV. – FRIVALDSZKY 1836b: 20, “Rumelia” (no de-
scription).

Remarks – Nomen nudum.

Genista incerta (Fabaceae)

Kétes rekettye. FRIV. Genista incerta. FRIV. – FRIVALDSZKY 1837: 163, “Hortiát” (no description).
Kétes rekettye (Genista incerta FR.) – FRIVALDSZKY 1845: 169, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Annls hist.-nat. Mus. natn. hung. 98, 2006

 An annotated list of Imre Frivaldszky’s plant and animal names 213

Geum sadleri (Rosaceae)

Geum Sadleri FRIVALDSZKY – FRIVALDSZKY 1835b: 54, “Rumelia” (no description).

Geum Sadleri FRIVALDSZK. – FRIVALDSZKY 1835c: 332, “Hab. In Rumelia” [Inhabits Rumelia] (de-
scription).
Geum Sadleri FRIVALDSZKY – FRIVALDSZKY 1836b: 20, “Rumelia”.
Sadler-féle czikász (Geum Sadleri FR.) – FRIVALDSZKY 1845: 166, “Philipopoly vidéke” [Region of
Plovdiv] (type locality restriction).

Remarks – Introduced as nomen nudum, but subsequently it was described on

the basis of an unstated number of syntypical specimens from “Rumelia”. Subse-
quently the species was redescribed, and the type locality restricted as region of
Plovdiv.

Haberlea rhodopensis (Gesneriaceae)

(Figs 2–4)

Rhodopei Haberlea. Haberlea rhodopensis. FRIVALDSZKY – FRIVALDSZKY 1835a: 249, “Rumeliá-

ban a’ Rodope’ közép magosságú hegyein virágzik.” [Blossoming in Rumelia, in Rhodope
Mountains on moderate elevations.] pl. I. (description).
Haberlea rhodopensis FRIVALDSZKY – FRIVALDSZKY 1835b: 54, “Rhodop.” [Rodopi].
Haberlea rhodopensis FRIVALDSZKY – FRIVALDSZKY 1835c: 331, “Habitat in montibus Rhodope
Rumeliae, floret Junio” [Inhabits the Rhodope Mountains of Rumelia; blossoming in June] (re-
description).
Haberlea rhodopensis FRIVALDSZKY – FRIVALDSZKY 1836b: 20, “Rumelia”.

Remarks – The species was described from an unstated number of syntypical

specimens originating from the Rhodope Mountains, collected at moderate eleva-
tions.

Heliotropium turcicum (Boraginaceae)

Heliotropium turcicum FRIVALDSZKY – FRIVALDSZKY 1836b: 436, “Hab. in Rumelia” [Inhabits

Rumelia] (description).
Török kunkor (Heliotropium turcicum), FRIV. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Re-
gion of Karlovo] (type locality restriction).
Török kunkor (Heliotropium turcicum FRIV.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region
of Karlovo].

Remarks – The species was described from an unstated number of syntypical

specimens originating from Rumelia. Subsequently the type locality was restricted
to the region of Karlovo.

Annls hist.-nat. Mus. natn. hung. 98, 2006

214 Zs. Bálint & S. Abadjiev

Hieracium cernuum (Asteraceae)

Konyuló hölgymál. Hieracium cernuum. FRIV. – FRIVALDSZKY 1840: 204, “Dervent’ havasain,
septemberben” [On the high mountains of Dervent, in September], pl. X. (description).
Konyuló holgyomál (Hieracium cernuum FR.) – FRIVALDSZKY 1845: 166, “Philipopoly vidéke”
[Region of Plovdiv].

Remarks – The species was described from an unstated number of syntypical

specimens originating from the high mountains of “Dervent, region of Plovdiv”.

Hieracium leucophylla (Asteraceae)

Hieracium Pilosella var. leucophylla FRIV. – FRIVALDSZKY 1836b: 20, “Rumelia” (no description).

Remarks – Nomen nudum.

Hieracium petraeum (Asteraceae)

Hieracium petraeum FRIVALDSZKY – FRIVALDSZKY 1836a: 436, “Hab. in Rumelia” [Inhabits

Rumelia] (description).
Hieracium petraeum FRIVALDSZKY – FRIVALDSZKY 1836b: 25, “Rumelia”.

Remarks – The species was described from an unstated number of syntypical

specimens originating from “Rumelia”.

Hieracium pilosissimum (Asteraceae)

Hieracium pilosissimum FRIVALDSZKY – FRIVALDSZKY 1836a: 436, “Hab. in Rumelia” [Inhabits

Rumelia], (description).
Legszőrösebb holgyomál (Hieracium pilosissimum), FRIV. – FRIVALDSZKY 1837b: 161, “Karlova
vidéke” [Region of Karlovo] (type locality restriction).
Legszőrösebb holgyomál (Hieracium pilosissimum) – FRIVALDSZKY 1845: 168, “Karlova vidéke”
[Region of Karlovo].

Remarks – The species was described from an unstated number of syntypical

specimens originating from “Rumelia”. Subsequently the type locality was re-
stricted as region of Karlovo.

Annls hist.-nat. Mus. natn. hung. 98, 2006

 An annotated list of Imre Frivaldszky’s plant and animal names 215

Hieracium rhodopeum (Asteraceae)

Hieracium rhodopeum FRIV. – FRIVALDSZKY 1836b: 25, “Rumelia” (no description).

Remarks – Nomen nudum.

Hieracium sessiliflorum (Asteraceae)

Kocsánytalan hölgymál. Hieracium sessiliflorum. FRIV. – FRIVALDSZKY 1840: 205, “Terem Calo-
phir’ havasain, augustusban” [Grows in the highest parts of Kaloferska Planina, in August.], pl.
XI. (description).
Kocsántalan holgyomál (Hieracium sessiliflorum FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke”
[Region of Karlovo].

Remarks – The species was described from an unstated number of syntypical

specimens originating from the highest part of Kaloferska Planina, region of
Karlovo.

Hieracium sparsum (Asteraceae)

Hieracium sparsum FRIVALDSZKY – FRIVALDSZKY 1836b: 436, “Hab, in Rumelia” [Inhabits

Rumelia] (description).

Remarks – The species was described from an unstated number of syntypical

specimens originating from “Rumelia”.

Hypericum rhodoppeum (Hyperiaceae)

Hypericum rhodoppeum FRIVALDSZKY – FRIVALDSZKY 1836a: 436, “Hab. in montib. Rhodoppe”

[Inhabits the Rhodope Mountains] (description).
Hypericum rhodopeum FRIV. – FRIVALDSZKY 1836b: 25, “Rumelia”.
Rhodope linka (Hypericum rhodopeum), FRIV. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Re-
gion of Karlovo].
Rodopei linka (Hypericum rodopaeum FR.) – FRIVALDSZKY 1845: 166, “Philipopoly vidéke” [Re-
gion of Plovdiv].

Remarks – The species was described from an unstated number of syntypical

specimens originating from various places in the Rhodope Mountains and the re-
gion of Karlovo.

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216 Zs. Bálint & S. Abadjiev

Juncus melanocephalus (Juncaceae)

Juncus melanocephalus FRIVALDSZKY – FRIVALDSZKY 1836a: 437, “Hab. in Rumelia” [Inhabits

Rumelia] (no description).
Juncus melanocephalus FRIV. – FRIVALDSZKY 1836b: 25, “Rumelia” (description).
Fekete füzérű szittyó (Juncus melanocephalus FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke”
[Region of Karlovo] (type locality restriction).

Remarks – Introduced as nomen nudum, but subsequently the species was de-
scribed from an unstated number of syntypical specimens originating from Ru-
melia. The type locality was later restricted as region of Karlovo.

Lactuca ramosissima (Asteraceae)

Ágas saláta. FRIV. Lactuca ramosissima. FRIV. – FRIVALDSZKY 1837b: 163, “Hortiát” (no descrip-
tion).

Remarks – Nomen nudum.

Lathyrus inermis (Fabaceae)

Kacstalan bükkön. Lathyrus inermis. ROCHEL. – FRIVALDSZKY 1835a: 250, “Virágzik Majus’ és
Juniusban a’ Balkány’ hegyeinek déli oldalában” [Blossoming in May and June, in the south-
ern side of Stara Planina.], pl. II. (description).
Kacsatlan bükköny (Lathyrus inermis ROCHL.) – FRIVALDSZKY 1845: 165, “Szlivno, Haemus déli s
éjszaki vidékei” [Sliven, southern and northern regions of Stara Planina] (type locality descrip-
tion).

Remarks – The species was described from an unstated number of syntypical

specimens from the southern side of Stara Planina. Subsequently the type locality
was restricted to Sliven. FRIVALDSZKY consequently attributed the authorship of
the species-group name to the botanist ANTON ROCHEL.

Lathyrus villosus (Fabaceae)

Lathyrus villosus FRIVALDSZKY – FRIVALDSZKY 1836a: 437, “Hab. in Rumelia” (no description).
Lathyrus villosus FRIV. – FRIVALDSZKY 1836b: 25, “Rumelia” (description).
Gyapjas bükköny (Lathyrus villosus F.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo] (type locality restriction).

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 An annotated list of Imre Frivaldszky’s plant and animal names 217

Remarks – Introduced as nomen nudum, but subsequently the species was de-
scribed from an unstated number of syntypical specimens originating from Ru-
melia. Subsequently the type locality was restricted as region of Karlovo.

Linum rigidum (Linaceae)

Könnyentörő len. FRIV. Linum rigidum. FRIV. – FRIVALDSZKY 1837: 163, “Hortiát” (no description).

Remarks – Nomen nudum.

Lithospermum rochelii (Boraginaceae)

Lithospermum Rochelii FRIVALDSZK – FRIVALDSZKY 1835b: 58, “Rumelia” (no description).

Lithospermum Rochelii FRIVALDSZK. – FRIVALDSZKY 1835c: 58, “Hab in Rumelia” [Inhabits
Rumelia] (description).
Lithospermum Rochelii FRIV. – FRIVALDSZKY 1836b: 20, “Rumelia”.
Rochel-féle kőmag (Lithospermum Rochelii FR.) – FRIVALDSZKY 1845: 166, “Philipopoly vidéke”
[Region of Plovdiv] (type locality restriction).

Remarks – Introduced as nomen nudum, but later it was described from an un-
stated number of syntypical specimens originating from Rumelia. Subsequently
the type locality was restricted as region of Plovdiv.

Mentha microphylla (Lamiaceae)

Mentha pulegium var. microphylla FRIVALDSZK. – FRIVALDSZKY 1835b: 57, “Rumelia” (no de-
scription).
Mentha pulegium var. microphylla FRIVALDSZK. – FRIVALDSZKY 1835c: 356, “Hab in Rumelia”
[Inhabits Rumelia].
Mentha Pulegium var. microphylla FRIV. – FRIVALDSZKY 1836b: 21, “Rumelia” (no description).

Remarks – Introduced as nomen nudum, but subsequently the species was de-
scribed from an unstated number of syntypical specimens originating from
“Rumelia”.

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218 Zs. Bálint & S. Abadjiev

Ostrya cylindrica (Betulaceae)

Hengerded gyertyán. FRIV. Ostrya cylindrica. FRIV. – FRIVALDSZKY 1837b: 163, “Hortiát” (no de-
scription).

Remarks – Nomen nudum.

Poa tenuis (Poaceae)

Poa pilosa var. tenuis FRIVALDSZK. – FRIVALDSZKY 1835b: 55, “Rumelia” (no description).
Poa pilosa var. tenuis FRIVALDSZK. – FRIVALDSZKY 1836b: 21, “Rumelia” (no description).

Remarks – Nomen nudum.

Potentilla beniczkyi (Rosaceae)

Potentilla Beniczkyi FRIVALDSZKY – FRIVALDSZKY 1836a: 437, “Hab. in Rumelia” [Inhabits

Rumelia] (description).
Potentilla Benyitzkyi FRIVALDSZKY – FRIVALDSZKY 1836b: 26, “Rumelia”.
Benyitzki pimpó (Poteutilla Benyitzkyi), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke”
[Region of Karlovo] (type locality restriction).
Benyiczky-féle pimpó (Potentilla Benyitzkyi F.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Re-
gion of Karlovo].

Remarks – The species was described from an unstated number of syntypical

specimens originating from Rumelia. Subsequently the type locality was restricted
as region of Karlovo.

Potentilla parva (Rosaceae)

Potentilla parva FRIV. – FRIVALDSZKY 1836b: 26, “Rumelia” (no description).

Remarks – Nomen nudum.

Potentilla tenuis (Rosaceae)

Potentilla rupestris var. gracilis FRIV. – FRIVALDSZKY 1836b: 21, “Rumelia” (no description).

Remarks – Nomen nudum.

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 An annotated list of Imre Frivaldszky’s plant and animal names 219

Primula turcica (Primulaceae)

Primula farinosa L. var. turcica FRIV. – FRIVALDSZKY 1836b: 26, “Rumelia” (no description).

Remarks – Nomen nudum.

Ranunculus nendtwichii (Ranunculaceae)

Ranunculus Nendtwichii FRIVALDSZKY – FRIVALDSZKY 1836a: 437, “Hab. in Rumelia” [Inhabits

Rumelia] (description).
Ranunculus Nendtvichii FRIVALDSZKY – FRIVALDSZKY 1836b: 26, “Rumelia”.
Nendtvich-szironták. Ranunculus Nentvichii FRIV. – FRIVALDSZKY 1840: 197, “Karlovai havasok’
tövében, juliusban” [At the foothills of the high mountains near Karlovo, in July], pl. III.
(redescription; type locality restriction).
Nendtvich-féle szironták (Ranunculus Nendtvichii) – FRIVALDSZKY 1845: 168, “Karlova vidéke”
[Region of Karlovo].

Remarks – The species was described from an unstated number of syntypical

specimens originating from Rumelia. Subsequently it was redescribed and the type
locality was restricted as the foothill region of the high mountains in the region of
Karlovo.

Reseda pseudovirens (Resedaceae)

Reseda pseudovirens FRIVALDSZKY – FRIVALDSZKY 1836b: 26, “Rumelia” (no description).

Remarks – Nomen nudum.

Rhamnus rumeliacus (Rhamnaceae)

Ruméliai benge. Rhamnus rumeliacus FRIVALDSZK. – FRIVALDSZKY 1835b: 54, “Rumelia” (no de-
scription).
Rhamnus rumeliaceus FRIV. – FRIVALDSZKY 1835c: 332, “Hab. In mont. Rodope” [Inhabits the
Rhodope Mountains] (description).
Rhamnus rumelianus FRIV. – FRIVALDSZKY 1836b: 21, “Rumelia”.
Rumeliai benge (Rhamnus rumeliacus FR.) – FRIVALDSZKY 1845: 166, “Philipopoly vidéke” [Re-
gion of Plovdiv] (type locality restriction).

Annls hist.-nat. Mus. natn. hung. 98, 2006

220 Zs. Bálint & S. Abadjiev

Remarks – Introduced as nomen nudum; later the species was described from
an unstated number of syntypical specimens from the Rhodope Mountains. Subse-
quently the type locality was restricted as region of Plovdiv.

Rubus sanguineus (Rosaceae)

Rubus sanguineus FRIVALDSZK. – FRIVALDSZKY 1835b: 55, “Rumelia” (no description).

Rubus sanguineus FRIVALDSZK. – FRIVALDSZKY 1835c: 334, “Hab. in montib. Rhodope” [Inhabits
the Rhodope Mountains] (description).
Rubus sanguineus FRIVALDSZKY – FRIVALDSZKY 1836b: 21, “Rumelia”.

Remarks – Introduced as nomen nudum, but subsequently the species was de-
scribed from an unstated number of syntypical specimens from the Rhodope
Mountains.

Rottboellia sandorii (Poaceae)

Rottboellia Sandorii FRIVALDSZK. – FRIVALDSZKY 1835b: 56, “Rumelia” (no description).

Rottboellia Sandorii FRIVALDSZK. – FRIVALDSZKY 1835c: 335, “Hab. in planit. Rumeliae” [Inhabits
the plains of Rumelia] (description).
Rottboellia Sandorii FRIV. – FRIVALDSZKY 1836b: 21, “Rumelia”.
Sándor törfű. Rottboellia Sándorii. FRIV. – FRIVALDSZKY 1840: 206, “Terem Filipopol és Karlova’
vidékin.” [Grows in the regions of Plovdiv and Karlovo], pl. XII. (redescription; type locality
restriction).
Sándor-féle törfű (Rotboelliea Sándorii FR.) – FRIVALDSZKY 1845: 166, “Philipopoly vidéke” [Re-
gion of Plovdiv].

Remarks – Introduced as nomen nudum, but later the species was described
from an unstated number of syntypical specimens originating from the plains of
Rumelia. Subsequently it was redescribed and the type locality was restricted as re-
gions of Plovdiv and Karlovo.

Satureja approximata (Lamiaceae)

Satureja approximata FRIVALDSZKY – FRIVALDSZKY 1836a: 438, “Hab. in Rumelia” (description).

Remarks – The species was described from an unstated number of syntypical

specimens originating from “Rumelia”.

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 An annotated list of Imre Frivaldszky’s plant and animal names 221

Satureja fasciculata (Lamiaceae)

Satureja montana, var. fasciculata FRIV. – FRIVALDSZKY 1836b: 26, “Rumelia” (no description).

Remarks – Nomen nudum.

Scabiosa ambigua (Dipsacaceae)

Scabiosa ambigua FRIVALDSZKY – FRIVALDSZKY 1836a: 438, “Hab. in Rumelia” (description).

Scabiosa ambigua FRIVALDSZKY – FRIVALDSZKY 1836b: 26, “Rumelia”.
Kétes sikkantyú (Scabiosa ambigua), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Re-
gion of Karlovo] (type locality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens originating from “Rumelia”. Subsequently the type locality was re-
stricted as region of Karlovo.

Scabiosa triniaefolia (Dipsacaceae)

Scabiosa triniaefolia FR. – FRIVALDSZKY 1835b: 55, “Rumelia” (no description).

Scabiosa triniaefolia FRIVALDSZK. – FRIVALDSZKY 1835c: 333, “Hab. in Rumelia” [Inhabits
Rumelia] (description).
Scabiosa triniaefolia FRIVALDSZK. – FRIVALDSZKY 1836b: 21, “Rumelia”.
Sallangos sikkantyú (Scabiosa triniaefolia FR.) – FRIVALDSZKY 1845: 166, “Philipopoly vidéke”
[Regions of Plovdiv] (type locality restriction).

Remarks – Introduced as nomen nudum, but subsequently the species was de-
scribed from an unstated number of syntypical specimens originating from “Ru-
melia”. Later the type locality was restricted as regions of Plovdiv.

Scrofularia majalis (Scrophulariaceae)

Május tákajak (Scrofularia majalis FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” (no descrip-
tion).

Remarks – Nomen nudum.

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222 Zs. Bálint & S. Abadjiev

Scutellaria pinnatifida (Lamiaceae)

Keleti csucsóka. Scutellaria orientalis. LIN. b. hasadt levelű. b. pinnatifida. ROCHL. – FRIVALDSZKY
1835a: 251, “Találtatik a’ Balkány’ hegyeinek dél-oldalaiban, virágzik Májusban” [It is found
on the southern sides of Stara Planina; flowering in May], pl. IV. (description).

Remarks – The species was described from an unstated number of syntypical

specimens originating from the southern slopes of Stara Planina. FRIVALDSZKY at-
tributed the authorship to the botanist ANTON ROCHEL.

Sedum ferrugineum (Crassulaceae)

Vörhenyeges szaka. Friv. Sedum ferrugineum. FRIV. – FRIVALDSZKY 1837b: 163, “Hortiát” (no de-
scription).
Vörhenyeges szaka (Sedum ferrugineum FR.) – FRIVALDSZKY 1845: 169, “Macedonia hegyei, s
bérczei között” [Grows among the mountains and cliffs of Macedonia.] (no description).

Remarks – Nomen nudum.

Sesleria coerulans (Poaceae)

Sesleria coerulans FRIVALDSZKY – FRIVALDSZKY 1836a: 438, “Hab. in Rumelia” [Inhabits

Rumelia] (description).
Sesleria coerulans FRIVALDSZKY – FRIVALDSZKY 1836b: 27, “Rumelia”.

Remarks – The species was described from an unstated number of syntypical

specimens.

Silene colorata (Caryophyllaceae)

Silene colorata FRIVALDSZK. – FRIVALDSZKY 1835b: 56, “Rumelia” (no description).

Silene colorata FRIVALDSZK. – FRIVALDSZKY 1835c: 334, “Hab. in Rumelia” [Inhabits Rumelia]
(description).
Silene colorata FRIVALDSZKY – FRIVALDSZKY 1836b: 22, “Rumelia”.

Remarks – Introduced as a nomen nudum, but subsequently the species was

described from an unstated number of specimens originating from “Rumelia”.

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Silene exaltata (Caryophyllaceae)

Silene exaltata FRIVALDSZ. – FRIVALDSZKY 1835b: 55, “Rumelia” (no description).

Silene exaltata FRIVALDSZK. – FRIVALDSZKY 1835c: 333, “Hab in Rumelia” [Inhabits Rumelia]
(description).
Magas sziléne (Silene exaltata FR.) – FRIVALDSZKY 1845: 166, “Philipopoly vidéke” [Region of
Plovdiv] (type locality restriction).

Remarks – Introduced as nomen nudum, subsequently the species was de-

scribed on the basis of an unstated number of syntypical specimens originating
from “Rumelia”. Later the type locality was restricted to the region of Plovdiv.

Silene römeri (Caryophyllaceae)

Silene Römeri FRIVALDSZKY – FRIVALDSZKY 1836a: 439, “Hab. in Rumelia” [Inhabits Rumelia]
(description).
Silene Römeri FRIVALDSZKY – FRIVALDSZKY 1836b: 27, “Rumelia”.
Römer siléne (Silene Römeri), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Region of
Karlovo] (type locality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens originating from “Rumelia”. Subsequently the type locality was re-
stricted as region of Karlovo.

Silene subconica (Caryophyllaceae)

Silene subconica FRIVALDSZK. – FRIVALDSZKY 1835b: 56, “Rumelia” (no description).

Silene subconica FRIVALDSZK. – FRIVALDSZKY 1835c: 334, “Hab. in Rumelia”[Inhabits Rumelia],
(description).
Silene subconica FRIVALDSZK. – FRIVALDSZKY 1836b: 22, “Rumelia”.

Remarks – Introduced as nomen nudum, but subsequently the species was de-
scribed from an unstated number of syntypical specimens originating from “Rumelia”.

Sisymbrium elatior (Brassicaceae)

Sisymbrium lipitzense var. elatior FRIV. – FRIVALDSZKY 1836b: 27, “Rumelia” (no description).

Remarks – Nomen nudum.

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224 Zs. Bálint & S. Abadjiev

Solanum assimile (Solanaceae)

Solanum assimile FRIVALDSZKY – FRIVALDSZKY 1836a: 439, “Hab. in Rumelia” [Inhabits Ru-
melia].

Remarks – The species was described from an unstated number of syntypical

specimens originating from “Rumelia”.

Stachys angustifolia (Lamiaceae)

Stachys recta JAQ. var. angustifolia FRIVALDSZK. – FRIVALDSZKY 1835b: 56, “Rumelia” (no de-
scription).
Stachys recta var. angustifolia FRIVALDSZK. – FRIVALDSZKY 1836b: 22, “Rumelia” (no description).

Remarks – Nomen nudum.

Stachys atelulata (Lamiaceae)

Csillagcsás hunyász. FRIV. Stachys atelulata. FRIV. – FRIVALDSZKY 1837b: 163, “Hortiát” (no de-
scription).
Csillagcsács hunnyász (Stachys stelulata FR.) – FRIVALDSZKY 1845: 169, “Macedonia hegyei, s
bérczei között” [Grows among the mountains and cliffs of Macedonia.] (no description).

Remarks – Nomen nudum.

Stachys villosa (Lamiaceae)

Stachy palustris var. villosa FRIVALDSZKY – FRIVALDSZKY 1836b: 27, “Rumelia” (no description).

Remarks – Nomen nudum.

Stellaria tenuifol[ia] (Caryophyllaceae)

Stellaria glauca var. tenuifol. FRIVALDSZKY – FRIVALDSZKY 1836b: 27, “Rumelia” (no descrip-
tion).

Remarks – Nomen nudum.

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 An annotated list of Imre Frivaldszky’s plant and animal names 225

Symphytum ottomanum (Boraginaceae)

Symphytum ottomanum FRIVALDSZKY – FRIVALDSZKY 1836a: 439, “Hab. in Rumelia” [Inhabits

Rumelia] (description).
Symphytum Ottomanum FRIVALDSZKY – FRIVALDSZKY 1836b: 27, “Rumelia”.
Ozmán nadálytő (Symphitum ottomanum), FRIV. – FRIVALDSZKY 1837b: 161, “Karlova vidéke”
[Region of Karlovo]; Symphytum ottomanum: 167, “Terem Carlova’ és Salonik’ vidé-
kén”[Grows in the region of Karlovo and Thessaloniki], pl. III. (redescription, type locality re-
striction).
Ozmán nadálytő (Symphytum ottomanum FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Re-
gion of Karlovo].

Remarks – The species was described from an unstated number of syntypical

specimens originating from “Rumelia”. In the redescription the type locality was
restricted as region of Karlovo and Thessaloniki.

Thymus comptus (Lamiaceae)

Thymus comptus FRIVALDSZKY – FRIVALDSZKY 1836a: 439, “Hab. in Rumelia” [Inhabits Rumelia]
(description).
Thymus comptus FRIVALDSZKY – FRIVALDSZKY 1836b: 27, “Rumelia”.
Szép démutka (Thymus comptus), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Region
of Karlovo] (type locality restriction).
Ékes démutka (Thymus comptus FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” (Region of
Karlovo).

Remarks – The species was described from an unstated number of syntypical

specimens originating from “Rumelia”. Subsequently the type locality was re-
stricted as region of Karlovo.

Thymus glaucus (Lamiaceae)

Zöldelő démutka FRIV. Thymus glaucus. FRIV. – FRIVALDSZKY 1837b: 163, “Hortiát” (no descrip-
tion).
Zöldellő démutka (Thymus glaucus FR.) – FRIVALDSZKY 1845: 169, “Macedonia hegyei, s bérczei
között” (no description).

Remarks – Nomen nudum.

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Tribulus albidus (Zygophyllaceae)

Tribulus terrestis L. var. albidus FRIVALDSZK. – FRIVALDSZKY 1835b: 57, “Rumelia” (no description).
Tribulus terrestis L. var. albidus FRIVALDSZK. – FRIVALDSZKY 1835b: 356, “Hab. in planit. Rume-
liae” [Occurs in the plain of Rumelia] (description).
Tribulus terrestis var. albid. FRIV. – FRIVALDSZKY 1836b: 22, “Rumelia”.
Fehérlő szurdancs. FRIV. Tribulus albidus. FRIV. – FRIVALDSZKY 1837b: 163, “Hortiát” (redescrip-
tion; type locality restriction).

Remarks – Introduced as a nomen nudum; subsequently the species was de-

scribed on the basis of an unstated number of syntypical specimens originating
from the plain of “Rumelia”. Later the type locality was restricted as “Hortiát” via
the redescription of the taxon.

Trifolium orientalis (Fabaceae)

Trifolium repens var. orientalis FRIV. – FRIVALDSZKY 1836b: 22, “Rumelia” (no description).

Remarks – Nomen nudum.

Verbascum glabratum (Scrophulariaceae)

Verbascum glabratum FRIVALDSZKY – FRIVALDSZKY 1836a: 440, “Hab. Rumelia” [Inhabits

Rumelia] (description).

Remarks – The species was described from an unstated number of syntypical

specimens originating from “Rumelia”.

Verbascum hinkei (Scrophulariaceae)

Verbascum Hinkei FRIVALDSZKY – FRIVALDSZKY 1936a: 440, “Hab. in Rumelia” [Inhabits

Rumelia] (description).
Hinke farkkóró (Verbascum Hinkei), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Re-
gion of Karlovo] (type locality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens from “Rumelia”. Subsequently the type locality was restricted as region
of Karlovo.

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 An annotated list of Imre Frivaldszky’s plant and animal names 227

Veronica microphylla (Scrophulariaceae)

Veronica hederaefolia var. microphylla FR. – FRIVALDSZKY 1836b: 28, “Rumelia” (no description).

Remarks – Nomen nudum.

Ziziphora compacta (Lamiaceae)

Ziziphora compacta FRIVALDSZK. – FRIVALDSZKY 1835b: 57, “Rumelia” (no description).

Ziziphora compacta FRIVALDSZK. – FRIVALDSZKY 1835c: 356, “Hab. in Rumelia” [Inhabits
Rumelia] (description).
Ziziphora compacta FRIVALDSZKY – FRIVALDSZKY 1836b: 22, “Rumelia”.
Tömött zizifű. Ziziphora compacta FRIV. – FRIVALDSZKY 1840: 198, “Találtatott Guklina’ vidékin
Rumeliában, virágzik juliusban” [Found in the region of Guklina, Rumelia; blossoming in
June], pl. IV (redescription).
Tömött zizifű (Ziziphora compacta FR.) – FRIVALDSZKY 1845: 166, “Philipopoly vidéke” [Region
of Plovdiv].

Remarks – Introduced as nomen nudum, but subsequently the species was de-
scribed from an unstated number of syntypical specimens from “Rumelia”. The
type locality was restricted via redescription as “Guklina”.

GASTROPODA

Bulimus athensis (Bulinidae)

Szenthegyi ehő (Bulimus Athensis FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Clausilia bicristata (Clausiliidae)

Kéttaréjú zárász (Clausilia bicristata FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

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Clausilia clathrata (Clausiliidae)

Oromos Z.(árasz ), Cl. (ausilia) clathrata FRIV. – FRIVALDSZKY 1856: 68, “Torna megyében a
szadellői völgy sziklafalain” [On the rocky walls of the Szádelő Gorge in County Torna] (no
description).

Remarks – Nomen nudum.

Clausilia dacica (Clausiliidae)

Dáciai Z.(árasz ), Cl. (ausilia) dacica FRIV. – FRIVALDSZKY 1856: 67, “a Duna szorosnál Kazán és
Islas vidékén a meredek mészkősziklák üregeiben s hasadékaiban” [In the regions of Kazán
and Islas at the Danube Gorge, in holes and openings of steep calcareous cliffs] (no descrip-
tion).

Remarks – Nomen nudum

Clausilia fritillaria (Clausiliidae)

Koczkás zárász. Clausilia fritillaria. FRIVALDSZKY – FRIVALDSZKY 1835a: 275, pl. VII, fig. 12.
“Találtatik a’ Balkány’ éjszak-oldali erdeiben” [Can be found in the northern side forests of the
Balkan] (description).
Clausilia fritilaria, FRIV. – FRIVALDSZKY 1837a: 92, “Habitat in Balcani montosis” [Inhabits Stara
Planina].
Kockás zárász (Clausilia fritillaria) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a Haemus
déli s éjszaki vidékei” [Sliven... the northern and southern regions of Stara Planina] (type local-
ity restriction).

Remarks – The species was described from an unstated number of syntypical

specimens from the northern side forests of the Balkan. Subsequently the type lo-
cality was restricted as Stara Planina.

Clausilia silacea (Clausiliidae)

Kovásd zárász. Clausilia silacea. FRIVALDSZKY – FRIVALDSZKY 1835a: 275, pl. VII, fig. 13.
“Találtatott a’ Balkány’ hegyeinek éjszak-oldai erdeiben.” [Found in the northern side forests
of the Balkan] (description).
C.(lauasilia) silacea, FRIV. – FRIVALDSZKY 1837a: 92, “Habitat in Balcani montosis” [Inhabits
Stara Planina].

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 An annotated list of Imre Frivaldszky’s plant and animal names 229

Kovásd zárász (Clausilia silecea FR.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a Haemus
déli s éjszaki vidékei” [Sliven… the southern and northern regions of Stara Planina] (type lo-
cality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens from the “northern side forests of the Balkans”. Subsequently the type
locality was restricted as Stara Planina.

Clausilia thessalonica (Clausiliidae)

Thessaloniki zárász (Clausilia thessalonica FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s
bérczei között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Helix dibotrion (Helicidae)

Két dudorú B.(iga), H. (elix) triaria, FRIV. – FRIVALDSZKY 1856: 67, “A Kárpát hegyláncolatának
lakója” [Inhabitant of the chain of the Carpathians] (no description).

Remarks – Nomen nudum.

Helix gorva (Helicidae)

Csempe biga. Helix gorva. FRIVALDSZKY – FRIVALDSZKY 1835a: 274, pl. VII, fig. 11. “Gyüjtetett a’
Balkány’ hegyeinek aljasabb domboldalaiban.” [Collected in the lower foothills of Stara
Planina] (description).
Helix girva, FRIV. – FRIVALDSZKY 1837a: 92, “Habitat in Balcani montosis.” [Inhabits Stara Planina].
Csempe biga (Helix girva FR.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a Haemus déli s
éjszaki vidékei” [Sliven… the southern and northern regions of Stara Planina] (type locality re-
striction).

Remarks – The species was described from an unstated number of syntypical

specimens from the lower foothills of Stara Planina. Subsequently the type locality
was restricted as the southern and northern regions of Stara Planina.

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230 Zs. Bálint & S. Abadjiev

Helix philibensis (Helicidae)

Philipopolyi bige (Helix Philibensis FR.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke” [Region
of Plovdiv] (no description).

Remarks – Nomen nudum.

Helix triaria (Helicidae)

Hármas B.(iga), H. (elix) triaria, FRIV. – FRIVALDSZKY 1856: 67, “Mehádia vidékén a Csorna folyó
völgyében” [In the valley of Csorna river, in region of Mehadia] (description).

Remarks – The species was described from an unstated number of syntypical

specimens collected in the valley of Cserna river, near Mehadia.

COLEOPTERA

Akis alutacea (Tenebrionidae)

Mákos tüsköny (Akis alutacea) FRIV. – FRIVALDSZKY 1837b: 163, “Hortiát” (no description).
Irhaféle tüsköny (Akis alutacea FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei
között” [Amongst the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Amphycoma bicolor (Glaphyridae)

Kétszínű bozont (Amphycoma bicolor) FRIVALD. – FRIVALDSZKY 1837b: 163, “Hortiát” (no de-
scription).
Kétszínű bozont (Amphycoma bicolor) FR. – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Anchomenus banaticus (Carabidae)

Anchomenus banaticus FRIV. (Bánsági Fojta). – FRIVALDSZKY 1865: 30, “… néhány példányban a
Korniarevai havason” [… a few specimen in the high mountains of Korniareva]; 176, “A

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Bánátban a Cornia-revai havason találtatott” [… found in the Banat, in the high mountains of
Cornia-Reva], pl. IX, fig. 6 (description).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex from the Banat, high mountains of Korniareva.

Ancylocheira dermestoides (Buprestidae)

Porvaféle pompály. (Buprestis Ancylocheira dermestoides FRIV.) – FRIVALDSZKY 1845: 180,

“Hazája Smyrna és Brussa vidéke, virágokon” [its habitat is Izmir and region of Bursa, found
on flowers], pl. 1, fig. 5. (description).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex from Izmir and the region of Bursa.

Amphycome cyanipennis (Glaphyridae)

Zöldkék bozont (Amphycome cyanipennis) FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke”
[Region of Karlovo] (no description).

Remarks – Nomen nudum.

Amphycoma martes (Glaphyridae)

Nyuszt bozont (Amphycoma martes FR.) – FRIVALDSZKY 1845: 177, “bythiniai Olympus” [Olympus
at Bythinia]; 182 “Találtatott Constantinápoly és Brussa környékén” [found in the region of Is-
tanbul and Bursa], pl. 2, fig. 2. (description).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sexes from the mount “Olympus” at “Bythinia”, and from
the regions of Istanbul and Bursa.

Anisoplia rumeliaca (Scarabeidae)

Rumeliai szipoly. Anisoplia rumeliaca. FRIVALD. – FRIVALDSZKY 1835a: 260, pl. V, fig. 12. “A
Balkány’ hegyeinek magasabb mezein” [In the higher pastures of the Stara Planina] (descrip-
tion).
Anisoplia rumeliaca, FRIV. – FRIVALDSZKY 1837a: 88, “Habitat in Balcani montosis” [Inhabits
Stara Planina] (redescription).

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232 Zs. Bálint & S. Abadjiev

Rumeliai szipoly (Anisoplia rumeliaca FR.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a
Haemus déli s éjszaki vidékei” [northern and southerns regions of Stara Planina near Sliven]
(type locality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex from higher regions of the Stara Planina. Subsequently
the type locality was restricted as the northern and southern slopes of Stara Planina
near Sliven.

Anophthalmus milleri (Carabidae)

Miller vaklája. Anophthalmus Milleri FRIV. J. (Miller Vaklandja). – FRIVALDSZKY 1865: 65, “… a
szokolávczi barlang” [the szokolávczi cave] (no description).
Anophthalmus Milleri FRIVALD. (Miller Vaklája). – FRIVALDSZKY 1865: 182, “Él a Krassó megyei
Szokolovácz völgyi barlangban” [It lives in the cave of the Szokolovácz valley, County
Krassó], pl. IX. fig. 15 (description).

Remarks – Introduced as a nomen nudum under the authorship of JÁNOS

FRIVALDSZKY, but subsequently in the same work the species was described by
FRIVALDSZKY on the basis of an unstated number of syntypical specimens of un-
stated sex from the cave in the “Szokolovácz” valley, County “Krassó”.

Anophthalmus redtenbacheri (Carabidae)

Anophthalmus Redtenbacheri nobis. – FRIVALDSZKY & FRIVALDSZKY 1857: 44, “… zuerst unter
Steinen in den tieferen Verzweigungen der Igritzer Grotte, später auch in anderen grösseren
Höhlen des Biharer Komitates” [First found under stones in the deeper laying branches of the
cave Igritz, but later was found other larger holes in County Bihar] (description).
Anophthalmus Redtenbacheri FRIV. (Redtenbacher Vaklandja). – FRIVALDSZKY 1865: 65, “Lelhe-
lyei az igiriczi és fericsei barlangok” (Collecting sites are the caves of Igricz and Fericse] (type
locality restriction); 182, “Találtatott a délbihar-megyei Igricz nevü barlangban.” [Found in the
cave named Igricz, in the southern part of County Bihar], pl. IX, fig. 14. (redescription).

Remarks – The species was described by JÁNOS and IMRE FRIVALDSZKY

from an unstated number of syntypical specimens of unstated sex originating from
the caves of County “Bihar”. Subsequently the type locality was restricted as the
caves of “Igricz” and “Fericse”.

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 An annotated list of Imre Frivaldszky’s plant and animal names 233

Apate aurita (Bostrichidae)

Füles görgöcz. Apate aurita. FRIVALD. – FRIVALDSZKY 1835a: 267, “Ritka a’ Balkány – oldali dűlt
fákban” [Rare in the fallen trees in the forests of the Balkans], pl. VI, fig. 10. “Ritka a’ Balkány
– oldali dűlt fákban.” (description).
Apate aurita, FRIV. – FRIVALDSZKY 1837a: 90, “Habitat in Balcani montosis” [Inhabits the Stara
Planina] (redescription, type locality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens from unstated sexes originating from the Balkans. Subsequently the
type locality was restricted as Stara Planina.

Apsis virens (Curculionidae)

Zöldellő gömbör (Apsis virens FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegylánczolata”
[Crete… the mountain range of Sphakia] (no description).

Remarks – Nomen nudum.

Armideus fossor (Geotrupidae)

Ásó fegyész (Armideus fossor) FRIV. – FRIVALDSZKY 1837a: 163, “Hortiát” (no description);
FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei között” [Among the cliffs and moun-
tains of Macedonia] (no description).

Remarks – Nomen nudum.

Asida banatica (Tenebrionidae)

Bánsági azád. Asida banatica FRIVALD. (bánsági Azád). – FRIVALDSZKY 1865: 46, “a Herkules-
fürdők közelében…” [In the vicinity of Herculean baths] (no description), 200 “Lelhelye
Mehádia vidéke” [It is found in the region of Mehadia], pl. XI, fig. 3. (description).

Remarks – Introduced as a nomen nudum from the region of “Herkules-

fürdő”, but subsequently in the same work the species was described from an un-
stated number of syntypical specimens unstated sex from the region of Mehadia.

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234 Zs. Bálint & S. Abadjiev

Boletophagus squarrosus (Tenebrionidae)

Berzedt taplász (Boletophagus squarrosus FR.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke”
[Region of Plovdiv] (no description).

Remarks – Nomen nudum.

Buprestis bivittis (Buprestidae)

Kétcsikú pompály (Buprestis bivittis FR.) – FRIVALDSZKY 1845: 177, “bythiniai Olympus” [Olym-
pus near Bythinia] (no description).

Remarks – Nomen nudum.

Buprestis decorata (Buprestidae)

Ékes pompály (Buprestis decorata FR.) – FRIVALDSZKY 1845: 177, “bythiniai Olympus” [Olympus
near Bythinia] (no description).

Remarks – Nomen nudum.

Buprestis flavostriata (Buprestidae)

Sárgavonatú pompály (Buprestis flavostriata FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia
hegylánczolata” [Crete… mountain ridge of Sphakia] (no description); Sárga vonatú pompály.
(Buprestis Ancylocheira flavostrigata Friv.): 179 “Hazája Kréta szigete, találtatott juniusban”
[Inhabits Crete, found in June], pl. 1, fig. 3. (description).

Remarks – Introduced as a nomen nudum from the mountain ridge of Spha-

kia, Crete. Subsequently in the same work the species was described on the basis of
an unstated number of syntypical specimens of unstated sex originating from Crete
collected in June.

Buprestis hyacinthina (Buprestidae)

Jáczint pompály (Buprestis hyacinthina FR.) – FRIVALDSZKY 1845: 177, “bythiniai Olympus”
[Olympus in Bythinia] (no description).
Jáczint pompály. (Buprestis Ancylocheira hyacinthina FRIV.) – FRIVALDSZKY 1845: 179, pl. 1, fig. 4.
“Hazája Brussa és Smyrna vidéke.” [Its habitat is the region of Bursa and Izmir] (description).

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 An annotated list of Imre Frivaldszky’s plant and animal names 235

Remarks – Introduced as a nomen nudum from the mount “Olympus” in

“Bythinia”, but subsequently in the same work the species was described on the ba-
sis of an unstated number of syntypical specimens of unstated sex originating from
the region of Bursa and Izmir.

Buprestis pilosellae (Buprestidae)

Holgyomáli pompály. Buprestis pilosellae. BONEL. – FRIVALDSZKY 1835a: 255, pl. V, fig. 5. “Talál-
tatott Szlivnó, és Philippopolis’ vidékin” [Found in the region of Sliven and Plovdiv] (descrip-
tion).
Buprestis pilosellae, FRIVALDSKY – FRIVALDSZKY 1837a: 87, “Habitat in Balcani montosis” [In-
habits the mountains of the Balkans] (redescription).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex originating from the region of Sliven and Plovdiv.

Buprestis olympica (Buprestidae)

Olympi pompály (Buprestis olympica FR.) – FRIVALDSZKY 1845: 177, “bythiniai Olympus” [Olym-
pus in Bythinia] (no description).

Remarks – Nomen nudum.

Buprestis ottomana (Buprestidae)

Ozmán pompály (Buprestis ottomana), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke”
[Region of Karlovo] (nomen nudum), 175, “Philippopol’ és Bicáncz’ vidékén találtatott”
[Found in the region of Plovdiv and Istanbul], pl.VII, fig. 5. (description).
Ozmán pompály (Buprestis ottomana FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo]; 172, “Constantinápoly kies vidéke” [Surroundings of Istanbul].

Remarks – Introduced as a nomen nudum for the region of Karlovo, but sub-
sequently in the same work the species was described on the basis of an unstated
number of syntypical specimens of unstated sex originating form the region of
Plovdiv and Istanbul. Later, in an other paper, FRIVALDSZKY repeated again that
the species occurred in the region of Karlovo and Istanbul.

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Buprestis scorzonerae (Buprestidae)

Pozdor pompály (Buprestis scorzonerae), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke”
[Region of Karlovo] (no description), 174, “Rumeliában és Macedoniában az együttnősző
növények’ virágain ritka.” [In Rumelia and Macedonia, on the flowers of monodyctelous
plants, rare] pl. VII, fig. 4. (description).
Pozdor pompály (Buprestis scorzonerae FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region
of Karlovo] (type locality restriction).

Remarks – Introduced as a nomen nudum from the region of Karlovo, but sub-
sequently it was described on the basis of an unstated number of syntypical speci-
mens originating from “Rumelia” and “Macedonia”. Later the type locality was re-
stricted as “Karlovo” in accordance to the first published record of the species.

Buprestis signaticollis (Buprestidae)

Jegyeltnyakú pompaly. Buprestis signaticollis. FRIVALD. – FRIVALDSZKY 1835a: 246. (no descrip-
tion)
Bélyegezett pompály. Buprestis stigmatica. SCHÖNH. – FRIVALDSZKY 1835a: 254, “Találtatott egy
példányban a’ Balkány’ déli oldalában” [A single specimen was found in the southern side of
Stara Planina], pl. V, fig. 4. (description).
Buprestis stigmatica, FRIVALDSKY. – FRIVALDSZKY 1837a: 86, “Habitat in Balcani montosis” [In-
habits Stara Planina] (redescription).

Remarks – Introduced as a nomen nudum from an unstated locality, but sub-

sequently the species was described on the basis of a single specimen (= holotype)
collected in the southern side of Stara Planina. In this publication FRIVALDSZKY
attributed the authorship of the name to the entomologist CARL JOHANN SCHÖNHERR
(1772–1848). Later FRIVALDSZKY “took” the authorship and recorded the species
in accordance to the original description.

Buprestis velox (Buprestidae)

Fürge pompály (Buprestis velox FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

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 An annotated list of Imre Frivaldszky’s plant and animal names 237

Byrhus sphaeroides (Byrrhidae)

Körded tekély (Byrhus sphaeroides FR.) – FRIVALDSZKY 1845: 177, “bythiniai Olympus” [Olympus
in Bythinia] (no description).
Remarks – Nomen nudum.

Calosoma rapax (Carabidae)

Ragadozó kecsély (Calosoma rapax FR.) – FRIVALDSZKY 1845: 176, “Cordylion, Smyrnátul a
tengeröblön át egy órányira” [Cordylion, one hour via the bay from Izmir] (no description).

Remarks – Nomen nudum.

Callidium insigne (Cerambycidae)

Jeles famász (Callidium insigne FR.) – FRIVALDSZKY 1845: 172, “Constantinápoly kies vidéke”
[Surroundings of Istanbul] (no description).

Remarks – Nomen nudum.

Callidium moesiacum (Cerambycidae)

Moesiai odváncz. Callidium moesiacum. FRIVALDSZKY – FRIVALDSZKY 1837b: 177, “Kazánlik’

erdeiben egy példányban találtatott” [A single specimen was found in the forests of Kazanlak],
pl. VII, fig. 7. (description).
Moesiai famász (Callidium moesiacum FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s
bérczei között” [In the mountains and cliffs of Macedonia].

Remarks – The species was described on the basis of a single specimen (=

holotype) of unstated sex from the forests of Kazanlak. Subsequently FRIVALDSZKY
reported a wider distribution for the species.

Callidium nogeli (Cerambycidae)

Nogel-féle famász (Callidium Nogeli) – FRIVALDSZKY 1845: 172, “Constantinápoly kies vidéke”
[Surroundings of Istanbul] (no description).
Nogel-féle famász. (Callidium Nogeli FRIV.) – FRIVALDSZKY 1845: 183, “Találtatott egy
példányban Constantinápoly vidékén” [A single specimen was found in the region of Istanbul],
pl. 2, fig. 4. (description).

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238 Zs. Bálint & S. Abadjiev

Remarks – Introduced as a nomen nudum, but subsequently in the same work

the species was described on the basis of a single specimen (= holotype) originat-
ing from the region of Istanbul.

Campylus orientalis (Elateridae)

Keleti görbő (Campylus orientalis FR.) – FRIVALDSZKY 1845: 177, “bythiniai Olympus” [Olympus
in Bythinia] (no description).

Remarks – Nomen nudum.

Cantharis pupillata (Cantharidae)

Látkás puhar (Cantharis pupillata FR.) – FRIVALDSZKY 1845: 172, “Constantinápoly kies vidéke”
[Surroundings of Istanbul] (no description).

Remarks – Nomen nudum.

Carabus salax (Carabidae)

Kaczér futoncz (Carabus salax FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Carabus thorosus (Carabidae)

Zömök futrinka. Carabus thorosus. FRIVALDSZKY – FRIVALDSZKY 1835a: 252, pl. V, fig. 2. “Lakik
a’ Balkánynak déli oldalában a’ szőlőhegyek között” [It lives in vineyards, in the southern foot-
hills of Stara Planina Mountain] (description).
Carabus thorosus FRIVALDSKY – FRIVALDSZKY 1837a: 86, “Habitat in montosis Balcani” [Inhabits
Stara Planina].
Zömök futoncz (Carabus thorosus, FRIV.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… Haemus
déli s éjszaki vidékei” [Sliven… southern and northern regions of Stara Planina] (type locality
restriction).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex originating from the vineyards cultivated at the foothills

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 An annotated list of Imre Frivaldszky’s plant and animal names 239

of Stara Planina. Subsequently FRIVALDSZKY restricted the type locality as vicin-

ity of Sliven.

Carabus versicolor (Carabidae)

Változékony futrinka. Carabus versicolor. FRIVALDSZKY – FRIVALDSZKY 1835a: 253 [no locality],
pl. V, fig. 3. (description).
Carabus versicolor FRIVALDSKY – FRIVALDSZKY 1837a: 86, “Habitat in montosis Balcani” [In-
habits the Stara Planina] (redescription).
Változékony futoncz (Carabus versicolor) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… Haemus
déli s éjszaki vidékei” [Sliven... northern and southern regions of Stara Planina] (type locality
restriction].

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex from unknown locality. The type locality was subse-
quently restricted as the northern and southern regions of Stara Planina Mountain
near Sliven.

Cardiophorus rotundicollis (Elateridae)

Körtorú szüvért (Cardiophorus rotundicollis FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia
hegylánczolata” [Crete… mountain range of Sphakia] (no description).
Körtorú szüvért (Cardiophorus rotundicollis FRIV.) – FRIVALDSZKY 1845: 181, pl. 1, fig. 6. “Hazája
Kréta szigete” [Inhabits the Island of Crete] (description).

Remarks – Introduced as a nomen nudum from the mountain range “Sphakia”

in Crete Island, but subsequently the species was described in the same work on the
basis of an unstated number of syntypical specimens of unstated sex originating
from “Crete”. In accordance with FRIVALDSZKY’s first record of the species we re-
strict the type locality of the nominal taxon as Sphakia mountains, Crete.

Cassida testudo (Chrysomelidae)

Teknős paizsna (Cassida testudo FR.) – FRIVALDSZKY 1845: 172, “Constantinápoly kies vidéke”
[Surroundings of Istanbul] (no description).

Remarks – Nomen nudum.

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Cerocoma kunzei (Meloidae)

Kunczei tülökfürt. Cerocoma Kunzei. FRIVALD. – FRIVALDSZKY 1835a: 265, “A’ balkány’ aljasabb
vidékein, Májusban a’ virágokon” [In the lower regions of Stara Planina; on flowers in May]),
pl. VI, fig. 7. (description).
Cerocoma Kunczei, FRIV. – FRIVALDSZKY 1837a: 89, “Habitat in Balcani, montis Majo” [Inhabits
Stara Planina] (redescription).
Kuncze-féle tülökfürt (Cerocoma Kunczei FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Re-
gion of Karlovo] (type locality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex from the lower regions of Stara Planina. Subsequently
the type locality was restricted as region of Karlovo.

Cetonia atrovirens (Scarabaeidae)

Sötét-zöldes diszély (Cetonia atrovirens FR.) – FRIVALDSZKY 1845: 176, “Cordylion, Smyrnátul a
tengeröblön át egy órányira” [Cordylion, one hour via the bay from Izmir] (no description).

Remarks – Nomen nudum.

Cetonia atrorubens (Scarabaeidae)

Setétvörös diszély. Cetonia atrorubens. FRIVALD. – FRIVALDSZKY 1835a: 262, “Rumeliában a’

csüküllön, Juniusban.” [In Rumelia, on cornflower, in June], pl. VI, fig. 3. (description).
Cetonia atrorubens, FRIVALDS. – FRIVALDSZKY 1837a: 88, “Habitat in Rumelia, Junio mense” [In-
habits Rumelia, in the month of June] (redescription).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex from Rumelia.

Ceuthorhynchus dimidiatus (Curculionidae)

Ceuthorhynchus dimidiatus FRIV. (féligfehér Sarla). – FRIVALDSZKY 1865: 124, “Buda-Pest vidéke”
[Region of Buda-Pest] (nomen nudum), 217 “Buda-Pest vidéke” [Region of Buda-Pest] (de-
scription) pl. XII, fig. 8. (description).

Remarks – Introduced as nomen nudum, but subsequently in the same work

the species was described from an unstated number of syntypical specimens of un-
stated sex from the region of “Buda-Pest”.

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 An annotated list of Imre Frivaldszky’s plant and animal names 241

Chlaenius exultus (Carabidae)

Meztelen biborna (Chlaenius exutus FRI.) – FRIVALDSZKY 1845: 176, “Cordylion, Smyrnátul a
tengeröblön át egy órányira” [Cordylion, one hour via the bay from Izmir] (nomen nudum).
Mezetlen biborna (Chlaenius exultus FRIVALD.) – FRIVALDSZKY 1845: 178 “Hazája Smyrna vidéke,
találtatik a folyóvizek közelében kövek alatt” [Its habitat is the region of Izmir; found under
stones near the rivers] (description), pl. 1, fig. 1. (description).

Remarks – Introduced as a nomen nudum from the mountain “Cordylion”, but

subsequently it was described on the basis of an unstated number of syntypical
specimens of unstated sex originating from the vicinity of Izmir. The “exutus” spe-
cies-group name of the original nomen nudum spelling was a lapsus, FRIVALDSZ-
KY emended it as “exultus”.

Cicindela despitensis (Carabidae)

Despiti czingolány (Cicindela despitensis FR.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke”
[Region of Plovdiv] (no description).

Remarks – Nomen nudum.

Cicindela rubens (Carabidae)

Vereslő czingolány. Cicindela rubens. FRIVALDSZKY – FRIVALDSZKY 1835a: 251, “Találtatott

homokos helyeken Balkány’ aljában a’ Tundzsa vize’ folytában” [Found in sandy places along
the flowing waters of Tundzha at the foothills of Stara Planina], pl. V, fig. 1. (description).
Cicindela rubens, FRIVALDSKY – FRIVALDSZKY 1837a: 86. “Habitat in montosis balcanis” [Inhabits
Stara Planina] (redescription).
Vermes futrinka (Cicindela rubens), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Re-
gion of Karlovo] (type locality restriction); 181, “A Kalophir havason csak gyéren találtatott”
[It was rarely found in the high mountains of Kalofer], pl. VII, fig. 11. (redescription, type lo-
cality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex originating from the valley of Tundzha river. Subse-
quently the type locality was restricted as region of Karlovo, and the species was
also recorded from higher elevations.

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242 Zs. Bálint & S. Abadjiev

Clerus concinnus (Cleridae)

Hason tátalék (Clerus concinnus FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Clerus thoracicus (Cleridae)

Toros tálték (Clerus thoracicus), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Region
of Karlovo] (no description).

Remarks – Nomen nudum.

Cratonychus saliens (Elateridae)

Szökő vivoncz (Cratonychus saliens FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegy-
lánczolata” [Crete… the mountain range of Sphakia] (no description).

Remarks – Nomen nudum.

Crypticus ferrugineus (Tenebrionidae)

Vörhönyös rejtőcz (Crypticus ferrugineus FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s
bérczei között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Cryptophagus frivaldszkyi (Cryptophagidae)

(Figs 7–9)

Cryptophagus Frivaldszkyi DEJEAN. – FRIVALDSZKY 1837b: 138 (nomen nudum).

Haplolophus neglectus FRIV. (elhanyagolt Gencsér). – FRIVALDSZKY 1865: 108, “… a budai he-
gyeken…” [in the mountains of Buda] (nomen nudum); 191 “… csak a budai hegységben…”
[only in the mountains of Buda], pl. X, fig. 9. (description).

Remarks – Introduced as a species dedicated to FRIVALDSZKY by the French

entomologist PIERRE FRANÇOIS MARIE AUGUSTE DEJEAN (1780–1845). Subse-

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 An annotated list of Imre Frivaldszky’s plant and animal names 243

quently the species was described under the authorship of FRIVALDSZKY on the ba-
sis of an unstated number of syntypical specimens originating from the mountains
of Buda. The name “neglectus” probably indicated that DEJEAN wanted to describe
the species, but he failed to do it because of an unknown reason.

Cymindis sublineata (Carabidae)

Vonalló gyorsod (Cymindis sublineata FR.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke” [Re-
gion of Plovdiv] (no description).

Remarks – Nomen nudum.

Dasytes cuneatus (Melyridae)

Ékded szőreg (Dasytes cuneatus FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegylánczolata”
[Crete… the mountain range of Sphakia] (no description).

Remarks – Nomen nudum.

Dichoma foraminosa (Tenebrionidae)

Likadékus duzma (Dichoma foraminosa FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia
hegylánczolata” [Crete… the mountain of Sphakia] (no description).

Remarks – Nomen nudum.

Dorcadion byzantinum (Cerambycidae)

Byzanczi izmócz (Dorcadion Byzantinum FR.) – FRIVALDSZKY 1845: 172, “Constantinápoly kies
vidéke” [Surroundings of Istanbul] (nomen nudum).

Remarks – Nomen nudum.

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244 Zs. Bálint & S. Abadjiev

Dorcadion exornatum (Cerambycidae)

Czifra izmócz. Dorcadion exornatum. FRIVALDSZKY – FRIVALDSZKY 1835a: 268, “A’ Balkány’
aljasabb vidékein ritkán” [Seldomly in the lower regions of Stara Planina], pl. VI, fig. 11. (de-
scription).
Dorcadion exornatum, FRIVALDS. – FRIVALDSZKY 1837a: 90, “Habitat in Balcani montosis” [In-
habits Stara Planina] (redescription).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex from the foothills of Stara Planina.

Dorcadion fallax (Cerambycidae)

Ál izmocz (Dorcadion fallax) FRIVALD. – FRIVALDSZKY 1837b: 163, “Hortiát” (no description).

Remarks – Nomen nudum.

Dorcadion inclusum (Cerambycidae)

Szigetes izmócz (Dorcadion inclusum KLUG.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke”
[Region of Plovdiv] (no description).

Remarks – Nomen nudum.

Dorcadion sturmii (Cerambycidae)

Sturm izmócz (Dorcadion Sturmii), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Re-
gion of Karlovo] (nomen nudum); 179, “Philippopol’ és Karlova’ kopár hegyein gyakor” [Com-
mon on the barren hills of Plovdiv and Karlovo], pl. VII, fig. 9. (description).
Sturm-féle izmócz (Dorcadion Sturmii FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region
of Karlovo].

Remarks – The species was described from an unstated number of syntypical

specimens originating from the barren hills in the region of Plovdiv and Karlovo.

Dorcadion ocskayi (Cerambycidae)

Ocskay-féle izmócz (Dorcadion Ocskayi FR.) – FRIVALDSZKY 1845: 177, “bythiniai Olympus”
[Olympus in Bythinia] (no description).

Remarks – Nomen nudum.

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 An annotated list of Imre Frivaldszky’s plant and animal names 245

Dorcadion triste (Cerambycidae)

Bús izmócz (Dorcadion triste FR.) – FRIVALDSZKY 1845: 176, “Cordylion, Smyrnátul a tengeröblön
át egy órányira” [Cordylion, one hour via the bay from Izmir] (nomen nudum); 184, “Hazája
Smyrna vidéke, a szántóföldeken” [Its habitat is Izmir, on the fields], pl. 2, fig. 5 (male); 185
“Találtatott az előbbeni társaságában” [It was found in the company of the previous], pl. 2, fig.
6. (female) (description).

Remarks – Introduced as a nomen nudum from the mountain Cordylion, but

in the same work subsequently the species was described on the basis of an un-
stated number of syntypical specimens of male and female sexes originating from
the region of Izmir. FRIVALDSZKY described both of the sexes in details.

Drimeotus kraatzii (Leiodidae)

Drimeotus Kraatzii nobis. – FRIVALDSZKY és FRIVALDSZKY 1857: 45, “… im südlichen Biharer

Comitate dem Orte Tericse naheliegende Grotte” [In the caves situated close to the settlement
Tericse, southern part of County Bihar] (description)
Kraatz Barlangárja (Drimeotus Kraatzi FRIV.) – FRIVALDSZKY 1865: 65, “… lelhelye a fericsei
barlang” [It was collected in the Fericse cave]; 185 “Találtatott a délbihar-megyei Fericse falu
melletti barlangban” [It was found in the cave near to the village Fericse, southern part of
County Bihar] , pl. X, fig. 4. (redescription).

Remarks – The species was described jointly by IMRE and JÁNOS FRI-
VALDSZKY on the basis of an unstated number of syntypical specimens of unstated
sex originating from the cave near the settlement “Fericse”. In a detailed redescription
and subsequent documentation the authorship of the species-group name was at-
tributed to himself by IMRE FRIVALDSZKY.

Elater hinkei (Elateridae)

Hinke szökcsér (Elater Hinkei), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Region
of Karlovo] (nomen nudum); 176, “Karlova’ vidékén” [In the region of Karlovo], pl. VII, fig. 6.
(description).
Hinke-féle szökcsér (Elater Hinkei FR. másik küldöttem emlékére nevezve) – FRIVALDSZKY 1845:
168, “Karlova vidéke” [Region of Karlovo].

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246 Zs. Bálint & S. Abadjiev

Remarks – Introduced as nomen nudum, but subsequently in the same work

the species was described from an unstated number of syntypical specimens from
unstated sex originating from the region of Karlovo. Later FRIVALDSZKY pre-
sented an etymology “named to the memory of my another delegate”. The first del-
egate ANDRÁS FÜLE of Hungarian descent disappeared during the first Balkan ex-
pedition. The second one JOSEPH HINKE of Zippser descent (an ethnic German from
Upper Hungary, now Slovakia) also disappeared in the third Balkan expedition.
Their assistant RICHARD MANULESCU of Roumanian descent survived all the trou-
bles, and brought back successfully the material collected during the expeditions.

Elater orientalis (Elateridae)

Keleti szökcsér (Elater orientalis FR.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke” [Region of
Plovdiv] (no description).

Remarks – Nomen nudum.

Epomis paueri (Carabidae)

Pauer merve (Epomis Paueri) FRIV. – FRIVALDSZKY 1837b: 163, “Hortiát” (no description).

Remarks – Nomen nudum.

Glaresis ruffa (Glaresidae)

Vörhenyes csorvány (Glaresis ruffa FRIV.) – FRIVALDSZKY 1859: 23, “Rákos sivatag” [Desert of
Rákos] (no description).

Remarks – Nomen nudum.

Haliplus venustulus (Haliplidae)

Kellemes úszka (Haliplus venustulus FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegy-
lánczolata” [Crete… mountain range of Sphakia] (no description).

Remarks – Nomen nudum.

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 An annotated list of Imre Frivaldszky’s plant and animal names 247

Helops ambiguus (Tenebrionidae)

Kétes bújnok (Helops ambiguus FR.) – FRIVALDSZKY 1845: 177, “bythiniai Olympus” [Olympus in
Bythinia] (no description).

Remarks – Nomen nudum.

Helops attenuatus (Tenebrionidae)

Vékony bujnok (Helops attenuatus FR.) – FRIVALDSZKY 1845: 176, “Cordylion, Smyrnátul a
tengeröblön át egy órányira” [Cordylion, one hour via the bay from Izmir] (no description).

Remarks – Nomen nudum.

Helops coelestinus (Tenebrionidae)

Mennyei bujnok (Helops coelestinus) – FRIVALDSZKY 1845: 176, “Cordylion, Smyrnátul a

tengeröblön át egy órányira” [Cordylion, one hour via the bay from Izmir] (no description).

Remarks – Nomen nudum.

Helops fuelei (Tenebrionidae)

Füle-féle bújnok (Helops Fülei FR. egyik küldöttem emlékére nevezve) – FRIVALDSZKY 1845: 167,
“Philipopoly vidéke” [Region of Plovdiv] (no description).

Remarks – Nomen nudum. FRIVALDSZKY presented the etymology: “named

in memory to one of my delegate” (see Elater hinkei).

Helops maurus (Tenebrionidae)

Szerecsen bújnok (Helops maurus FR.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke” [Region of
Plovdiv] (no description).

Remarks – Nomen nudum.

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248 Zs. Bálint & S. Abadjiev

Helops obesus (Tenebrionidae)

Potrohos bújnok. Helops obesus FRIV. – FRIVALDSZKY 1835a: 248 (nomen nudum); 262, “Tavasz-
kor a’ Balkány’ hegyeinek éjszak-oldalaiban” [In spring; in the northern side of Stara Planina],
pl. VI, fig. 4. (description).
Helops obesus, FRIV. – FRIVALDSZKY 1837a: 89, “Habitat in Balcani montosis” (Habitat in the
mountains of the Balkans] (redescription).

Remarks – Introduced as a nomen nudum, but subsequently the species was

described in the same work on the basis of an unstated number of syntypical speci-
mens of unstated sex originating from the northern side of Stara Planina.

Helops terreni (Tenebrionidae)

Terren-féle bújnok (Helops Terreni FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegylánczo-
lata” [Crete… mountain range of Sphakia] (nomen nudum); 183, “hazája Kréta szigete,
találtatott cziprusfákon” [Inhabits Crete; it was found on cypress trees], pl. 2, fig. 3. (descrip-
tion).

Remarks – Introduced as nomen nudum, but subsequently in the same work

the species was described from an unstated number of syntypical specimens of un-
stated sex originating from Crete. In the first mentioning of the species-group
name FRIVALDSZKY gave a more restricted collecting site, which can be accepted
as type locality according to the ICZN (1999) Art. 76.1.

Hoplia flavipes (Scarabaeidae)

Sárgalábú karmócz. Hoplia flavipes. DEJ. – FRIVALDSZKY 1835a: 261 “A’ Balkány’ alján a’ fészkes
virágokon ritkábban.” [Rarely on flowers of daisy family in the foothills of Stara Planina] (de-
scription).
Hoplia flavipes, FRIVALD. – FRIVALDSZKY 1837a: 88, “Habitat in Balcani montosis” [Inhabits Stara
Planina] (redescription).
Sárgalábú izgoncz (Lytta flavipes) – FRIVALDSZKY 1845: 177, “bythiniai Olympus” [Olympus in
Bythinia] (type locality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex collected on flowers of daisy family and originating
from the mountains of the Balkans. Originally he considered the French entomolo-
gist P. F. M. A. DEJEAN to be the author of the species-group name. Subsequently

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 An annotated list of Imre Frivaldszky’s plant and animal names 249

the species-group name was transferred from Hoplia to Lytta and the type locality
was restricted as “Olympus in Bythinia” by FRIVALDSZKY. Thus the authorship of
the name was regarded to FRIVALDSZKY himself.

Laena pilosa (Tenebrionidae)

Szőrős bujka (Laena pilosa FR.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke” [Region of Plov-
div] (no description).

Remarks – Nomen nudum.

Laena pygmaea (Tenebrionidae)

Piczi bujka (Laena pygmaea FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei között”
[In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Leptonotus redtenbahii (Leiodidae)

Redtenbach-féle sugárcz (Leptonotus Redtenbahii FR.) – FRIVALDSZKY 1845: 177, “új nem!” [new
genus!] (no description).

Remarks – Nomen nudum. FRIVALDSZKY introduced two nomina nuda, one

on genus-group, and another one on species-group level.

Leptopus nigroflabellatus (Scarabaeidae)

Feketecsápú lábasugár (Leptopus nigroflabellatus FR.) – FRIVALDSZKY 1845: 170, “Macedonia

hegyei, s bérczei között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

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Liparus intermedius (Curculionidae)

Közi kente (Liparus intermedius FR.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke” [Region of
Plovdiv] (no description).

Remarks – Nomen nudum.

Lucanus crenulatus (Lucanidae)

Ajócskás agancsár (Lucanus crenulatus FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegy-
lánczolata” [Crete… mountain range of Sphakia] (no description).

Remarks – Nomen nudum.

Lytta phalerata (Meloidae)

Csótáros izgoncz (Lytta phalerata) FRIVALD. – FRIVALDSZKY 1837b: 163, “Hortiát” (no descrip-
tion), 182 “Szalonik’ vidékén a’ kőrösfán gyakran találtatott.” [It was often found on ash trees
in the region of Thessaloniki], pl. VII, fig. 12. (description).
Csótáros izgoncz (Lytta phalerata FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia].

Remarks – Introduced as nomen nudum, but in the same work subsequently

the species was described from an unstated number of syntypical specimens of un-
stated sex collected on ash trees in the region of Thessaloniki, which should be the
mountain “Hortiát” according to the first mention of the taxon. Therefore the type
locality determined here as “Hortiát”, near Thessaloniki. Later FRIVALDSZKY indi-
cated a wider distribution for the species in Greek Macedonia.

Malachius flabellatus (Melyridae)

Legyezős hólagány. Malachius flabellatus. FRIVALDSZKY – FRIVALDSZKY 1835a: 255, pl. V, fig. 6.
“Virágokon a’ Balkány’ alján” [On flowers in the foothills of the Balkans] (description).
Malachius flabellatus, FRIVALDS. – FRIVALDSZKY 1837a: 87, “Habitat in Balcani montosis” [In-
habits Stara Planina] (redescription).
Legyezős holagány (Malachius flabellatus FR.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)…
Haemus déli s éjszaki vidékei” [The northern and southern regions of Stara Planina, Sliven]
(type locality restriction).

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 An annotated list of Imre Frivaldszky’s plant and animal names 251

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex collected on flowers in the foothills of Stara Planina.
Subsequently the type locality was restricted as northern and southern sides of
Stara Planina, near Sliven.

Malachius coccineus (Melyridae)

Karmazsin holagány (Malachius coccineus FR.) – FRIVALDSZKY 1845: 172, “Constantinápoly kies
vidéke” [Surroundings of Istanbul] (no description).

Remarks – Nomen nudum.

Malacosoma thoracica (Chrysomelidae)

Toros lágyacz (Malacosoma thoracica FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s
bérczei között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Meleus lutosus (Curculionidae)

Sáros ripók. Meleus lutosus. FRIVALD. – FRIVALDSZKY 1835a: 266, pl. VI, fig. 9. “Egy példányban
találtatott Szlivnó vidékén, Juniusban” [A single specimen was found in the region of Sliven, in
June] (description).
Meleus lutosus, FRIV. – FRIVALDSZKY 1837a: 89, “Habitat in Balcani montosis, Junio” [Inhabits
Stara Planina] (redescription).

Remarks – The species was described on the basis of a single specimen (=

holotype) of unstated sex collected in June, in regions of Sliven.

Meloe aeruginosa (Meloidae)

Rezecs-zöld nünüke (Meloe aeruginosa FR.) – FRIVALDSZKY 1845: 176, “Cordylion, Smyrnátul a
tengeröblön át egy órányira” [Cordylion, one hour via the bay from Izmir] (no description).

Remarks – Nomen nudum.

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Melolontha albida (Scarabaeidae)

Deres cserebűly. Melolontha albida. DEJEAN. – FRIVALDSZKY 1835a: 258, pl. V, fig. 10. “Rume-
liában ’s a’ Jónai szigeteken estve repdes” [It flies in the evenings, in the Ionian Islands and in
Rumelia] (description).
Melolontha albida, FRIVALDS. – FRIVALDSZKY 1837a: 87, “Habitat in Balcani montosis” [Inhabits
Stara Planina] (redescription).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex collected, taken in the Ionian Islands attributing the au-
thorship first to the French entomologist P. F. M. A. DEJEAN. Subsequently
FRIVALDSZKY considered himself as the sole author.

Melolontha tenebrionides (Scarabaeidae)

Rejtényded cserebűly. Melolontha tenebrionides. PALLAS. – FRIVALDSZKY 1835a: 259, “Rumeliá-

nak laposabb vidékein” [Lower regions of Rumelia], pl. V, fig. 11. (description).
Melolontha tenebrionides, FRIVALD. – FRIVALDSZKY 1837a: 88, “Habitat in Balcani montosis” [In-
habits Stara Planina] (redescription).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex originating from the lower regions of “Rumelia”. The
authorship of the species group name was attributed to the entomologist PETER SI-
MON PALLAS (1741–1811) by FRIVALDSZKY, but in the subsequent recording of
the taxon he attributed the name to himself and gave a distribution for the species
indicating as a mountain species contradicting the remark of the previous state-
ment that it was collected in flatland.

Molops dilatatus (Carabidae)

Széleske holdarcz (Molops dilatatus FR.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke” [Region
of Plovdiv] (no description).

Remarks – Nomen nudum.

Mordella sacheri (Mordellidae)

Mordella Sacheri FRIV. (Sacher Marókája). – FRIVALDSZKY 1865: 47, “Mehádia vidékén és
Szlavóniában” [In the region of “Mehádia” and in “Szlavónia”] (no description), 202, “Me-

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 An annotated list of Imre Frivaldszky’s plant and animal names 253

hádia vidékén és Szlavóniában” [In the region of “Mehádia” and in “Szlavónia”], pl. XI, pl. 6.
(description).

Remarks – Introduced as a nomen nudum, but subsequently the species was

described in the same work on the basis of an unstated number of syntypical speci-
mens of unstated sex originating from the region of “Mehádia” and from “Szla-
vónia”.

Mylabris decora (Meloidae)

Ékes osztrák. Mylabris decora. FRIVALD. – FRIVALDSZKY 1835a: 264 “A’ Balkány’ aljában a’
virágokon junius elején gyakor” [Rather common in the foothills of the Balkans on flowers in
the beginning of June], pl. VI, fig. 6. (description).
Mylabris decora, FRIV. – FRIVALDSZKY 1837a: 89, “Habitat in Balcani montosis, Junio” [Inhabits
Stara Planina] (redescription).
Ékes nyüzse (Mylabris decora FR.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a Haemus déli
s éjszaki vidékei” [In the northern and southern regions of Stara Planina, Sliven] (type locality
restriction).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex collected on flowers in the beginning of June at the foot-
hills of the Balkans. Subsequently the type locality was restricted as northern and
southern sides of Stara Planina, near Sliven.

Mylabris turcica (Meloidae)

Török nyüzse (Mylabris turcica FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Myniops carinatus (Curculionidae)

Ormos érdöcz (Myniops carinatus FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo] (no description).

Remarks – Nomen nudum.

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Myniops obsoletus (Curculionidae)

Avult érdöcz (Myniops obsoletus) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of Kar-
lovo] (no description).

Remarks – Nomen nudum.

Oedemera gracilis (Oedemeridae)

Vékony czombony (Oedemera gracilis FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegy-
lánczolata” [Mountain ridge of Sphakia, Crete] (no description).

Remarks – Nomen nudum.

Omaloplia erythroptera (Scarabaeidae)

Vörhenyeges bársonyka. Omaloplia erythroptera. DEJ. – FRIVALDSZKY 1835a: 260, “Repül a’

Balkány’ aljasaiban Juniusban.” [Flies in June, at the foothills of the Balkans], pl. VI, fig. 1.
(description).
Omaloplia erythroptera, FRIVALD. – FRIVALDSZKY 1837a: 88, “Habitat in Balcanio montosis,
Junio” [Inhabits Stara Planina, June.] (redescription).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex collected in June at the foothills of the Balkans. First
FRIVALDSZKY attributed the authorship of the species-group name to the French
entomologist P. F. M. A. DEJEAN, but in a subsequent publication he took the au-
thorship.

Omaseus creticus (Carabidae)

Krétai csatang, (Omaseus creticus FRIV.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegylán-
czolata” [Mountain range of Sphakia, Crete] (no description), 179, “Hazája Kréta szigete,
Canea vidéke” [Inhabits region of Canea, Crete Island], pl. 1, fig. 2. (description).

Remarks – Introduced as a nomen nudum, but subsequently in the same work

the species was described from an unstated number of syntypical specimens of un-
stated sex originating from the region of “Canea”, Crete Island.

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 An annotated list of Imre Frivaldszky’s plant and animal names 255

Omias hanakii (Curculionidae)

Omias Hanákii (Hanák Segé-je). – FRIVALDSZKY 1865: 17, “M. Sziget vidékein” [In the region of
Máramaros] (no description).
Omias Hanakii FRIV. (Hanák Sergéje). – FRIVALDSZKY 1865: 205, “Máramaros megyében” [In
County Máramaros], pl. XI, fig. 7. (description).

Remarks – Introduced as nomen nudum, but subsequently in the same work

the species was described from an unstated number of syntypical specimens of un-
stated sex originating from County “Máramaros”.

Omias leuconotus (Curculionidae)

Fehérpontú serge (Omias leuconotus FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Opatrum granulosum (Tenebrionidae)

Magcsás földicze (Opatrum granulosum FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegy-
lánczolata” [Mountain ridge of Sphakia, Crete] (no description).

Remarks – Nomen nudum.

Othyorhinchus bicostatus (Curculionidae)

Kétbordás fogor (Othyorhinchus bicostatus FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s
bérczei között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Othyorhinchus metalliger (Curculionidae)

Érces fogor (Othyorhinchus metalliger FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s
bérczei között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

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Othyorinchus obesus (Curculionidae)

Potrohos fogor (Othyorinchus obesus FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegylán-
czolata” [Mountain range of Sphakia, Crete] (no description).

Remarks – Nomen nudum.

Othyorinchus rotundatus (Curculionidae)

Gömbölyü fogor (Othyorinchus rotundatus FR.) – FRIVALDSZKY 1845: 172, “Constantinápoly kies
vidéke” [Surroundings of Istanbul] (no description).

Remarks – Nomen nudum.

Pandarus moesiacus (Tenebrionidae)

Moesiai adongo (Pandarus moesiacis), FRIVALDSZKY – FRIVALDSZKY 1837b: 161, “Karlova vidé-
ke” [Region of Karlovo] (no description).

Remarks – Nomen nudum.

Pausus turcicus (Carabidae)

Török csápiker. Pausus turcicus. FRIVALDSZKY – FRIVALDSZKY 1835a: 263, “Találtatott egy
példányban Selimno’ vidékin” [A single specimen was found in the region of Sliven] pl. VI,
fig. 5. (description).
Pausus turcicus, FRIV. – FRIVALDSZKY 1837a: 89, “Habitat in Balcani montosis; Selimno” [Inhabits
Stara Planina; Sliven] (redescription).
Török csápiker (Pausus turcicus FR.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a Haemus
déli s éjszaki vidékei” [Southern and northern regions of Stara Planina, Sliven] (type locality
restriction).

Remarks – The species was described on the basis of a single specimen (=

holotype) of unstated sex collected in the region of Sliven. Subsequently the type
locality was restricted as southern and northern side of Stara Planina, near Sliven.

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 An annotated list of Imre Frivaldszky’s plant and animal names 257

Pedinus macedonicus (Tenebrionidae)

Macedóniai mezöny (Pedinus macedonicus FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s
bérczei között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Platymra depressa (Carabidae)

Lapított lapócsa (Platymra depressa FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomina nuda. Neither the species-group nor the genus-group

names were ever used subsequently.

Procrustes vicinus (Carabidae)

Szomszéd nyútovány (Procrustes vicinus), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidé-
ke” [Region of Karlovo] (no description).
Rokon nyújtovány (Procrustes vicinus FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo] (no description).

Remarks – Nomen nudum. At the second time FRIVALDSZKY named differ-

ently in Hungarian the species.

Psallidium fossulatum (Curculionidae)

Likacsos nyirócza (Psallidium fossulatum FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s
bérczei között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

Psalidium vittatum (Curculionidae)

Szalagos nyirócza (Psalidium vittatum), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke”
[Region of Karlovo] (no description). FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo] (no description).

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258 Zs. Bálint & S. Abadjiev

Remarks – Nomen nudum. FRIVALDSZKY used consequently the binomial

name two times.

Pholeuon leptodirum (Leiodidae)

Pholeuon Leptodirum nobis. – FRIVALDSZKY és FRIVALDSZKY 1857: 44, “… bewohnt im südlichen

Biharer Comitate… Funaczaer Grotte” [Inhabits the Funacza cave, southern part of County
Bihar] (description).
Pholeuon leptoderum Friv. (Hosszucsápu Odorász). – FRIVALDSZKY 1865: 65, “Lelhelye a fonáczai
barlang” [its collecting site is the cave of Fonácza] ; 183 “Találtatott a délbihar-megyei
hegyekben Rézbánya közelében létező Fonáczai barlangban.” [It was found in the cave of
Fonácza, close to Rézbánya in the southern mountains of county Bihar], pl. X, fig. 2.
(redescription).

Remarks – The species was described from an unstated number of syntypical

specimens of both sexes collected in the cave “Funacza”, in the southern part of
county Bihar. Subsequently FRIVALDSZKY gave more precise indication for the
type locality.

Platynus coerulescens (Carabidae)

Kékes lapócsa (Platynus coerulescens FR.) – FRIVALDSZKY 1845: 177, “bythiniai Olympus” [Olym-
pus in Bythinia] (no description).

Remarks – Nomen nudum.

Platyscelis hungaricus (Tenebrionidae)

Magyar gyászár (Platyscelis hungaricus) – FRIVALDSZKY 1859: 23, “Csabacsüdi puszta, Szarvas”
[Steppe of Csabacsüd, Szarvas] (no description).
Platyscelis hungaricus FRIVALDSZKY (magyarhoni Gyászár). – FRIVALDSZKY 1865: 115, “…
legelőször Békés megyében, a csabacsűdi pusztán… későbben az alsó-hevesi poói pusztán”
(no description) [First in the steppe of Csabacsűd; later in the steppe of Poó in Alsó-Heves];
200, “Kizárólagos hazája a magyar Alföld” [Its exclusively home is the Hungarian Plain], pl.
XI, fig. 2. (description).

Remarks – Introduced as nomen nudum, but subsequently the species was de-
scribed from an unstated number of syntypical specimens of male and female
sexes collected exclusively on the Hungarian Plain. The precise type locality was
indicated as the steppes of “Csabacsűd” and “Poó”, thus all FRIVALDSZKY speci-

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mens originating from these sites can be considered as syntypical as well as

FRIVALDSZKY specimens with the generalised data “Hungaria”.

Pterostichus fulgens (Carabidae)

Csillámló röpér (Pterostichus fulgens), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke”
[Region of Karlovo] (no description); FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo] (no description).

Remarks – Nomen nudum.

Pterostichus sacheri (Carabidae)

Pterostichus Sacheri FRIV. J. (Sacher Röpére). – FRIVALDSZKY 1865: 67, “Bohodej havas magas-
lata” [The peak of the high mountain Bohodej] (no description); 176, “A bihari Bohodej nevü
havas tetején” [On the top of the high mountain Bohodei in Bihar], pl. IX, fig. 7. (description).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex originating from the peak of the high mountain Bohodei,
in county Bihar. The species-group name was attributed to JÁNOS FRIVALDSZKY.

Purpuricenus desfontainii (Cerambycidae)

Desfontain-féle biboros (Purpuricenus Desfontainii FR.) – FRIVALDSZKY 1845: 172, “Constanti-

nápoly kies vidéke” [Surroundings of Istanbul] (no description).

Remarks – Nomen nudum.

Rhizotragus costulatus (Scarabaeidae)

Bordás csalya. Rhizotragus costulatus. FRIVALDSZKY – FRIVALDSZKY 1835a: 257, “Május elején,
a’ Balkány aljas vidékein, társaságosan” [Gregariously, at the beginning of May in the lower
parts of the Balkans], pl. V, fig. 9. (description).
Rhizotrogus costulatus, FRIV. – FRIVALDSZKY 1837a: 87, “Habitat in Balcani montosis” [Inhabits
Stara Planina] (redescription).
Bordás csalya (Rhizotragus costulatus FR.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a
Haemus déli s éjszaki vidékei” [Southern and northern regions of Stara Planina, Sliven] (type
locality restriction).

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Remarks – The species was described from an unstated number of syntypical

specimens of unstated number collected in series at the lower parts of the Balkans,
in May. Subsequently the type locality was restricted as southern and northern side
of Stara Planina, near Sliven.

Rhizotrogus torulosus (Scarabaeidae)

Göröngyös csalya (Rhizotrogus torulosus FR.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke”
[Region of Plovdiv] (no description).

Remarks – Nomen nudum.

Rhynchites gigas (Rhynchitidae)

Óriás eszelény (Rhynchites gigas FR.) – FRIVALDSZKY 1845: 176, “Cordylion, Smyrnátul a tenger-
öblön át egy órányira” [Cordylion, one hour via the bay from Izmir] (no description).

Remarks – Nomen nudum.

Saperda atomaria (Cerambycidae)

Parányozott zenész (Saperda atomaria), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke”
[Region of Karlovo]; 178, “Karlova’ vidékén” (In the Region of Karlovo], pl. VII, fig. 8. (de-
scription).

Remarks – Described on the basis of a single specimen (= holotype) of un-

stated sex collected in the region of Karlovo.

Saperda balcanica (Cerambycidae)

Balkáni zenész. Saperda balcanica. FRIVALD. – FRIVALDSZKY 1835a: 268, pl. VI, fig. 12. “Találta-
tott egy példányban Szlivnó’ vidékén” [A single specimen was found in the region of Sliven]
(description).
Saperda balcanica, FRIVALDS. – FRIVALDSZKY 1837a: 90, “Habitat in Balcani montosis” [Inhabits
Stara Planina] (redescription).
Balkáni zenész (Saperda balcanica FR.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a
Haemus déli s éjszaki vidékei” [Southern and northern side of Stara Planina, Sliven] (type lo-
cality restriction), 172, “Constantinápoly kies vidéke” [Surroundings of Istanbul].

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 An annotated list of Imre Frivaldszky’s plant and animal names 261

Remarks – The species was described on the basis of a single specimen (=

holotype) of unstated sex collected in the vicinity of Sliven. Subsequently the type
locality was restricted to the southern and northern sides of Stara Planina, near
Sliven.

Saperda egregia (Cerambycidae)

Jeles zenész (Saperda egregia FR.) – FRIVALDSZKY 1845: 176, “Cordylion, Smyrnátul a tengeröblön
át egy órányira” [Cordylion, one hour via the bay from Izmir] (no description).

Remarks – Nomen nudum.

Saperda pilosa (Cerambycidae)

Szőrös zenész (Saperda pilosa FR.) – FRIVALDSZKY 1845: 177, “bythiniai Olympus” [Olympus in
Bythinia] (no description).

Remarks – Nomen nudum.

Saperda zavadzkyi (Cerambycidae)

Zavadszky-féle zenész (Saperda Zavadzkyi FR.) – FRIVALDSZKY 1845: 172, “Constantinápoly kies
vidéke” [Surroundings of Istanbul] (no description).

Remarks – Nomen nudum.

Scarabaeus fulgens (Scarabaeidae)

Ragyogó csajva (Scarabaeus fulgens FR.) – FRIVALDSZKY 1845: 172, “Constantinápoly kies vi-
déke” [Surroundings of Istanbul] (no description).

Remarks – Nomen nudum.

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Scarites cyclops (Carabidae)

Cziklops túrász (Scarites cyclops FR.) – FRIVALDSZKY 1845: 176, “Cordylion, Smyrnátul a tenger-
öblön át egy órányira” [Cordylion, one hour via the bay from Izmir] (no description).

Remarks – Nomen nudum.

Silpha multipunctata (Silphidae)

Sokpontú dögész (Silpha multipunctata FR.) – FRIVALDSZKY 1845: 177, “bythiniai Olympus”
[Olympus in Bythinia] (no description); 182, “Hazája Constantinápoly és Smyrna vidéke” [In-
habits the region of Istanbul and Izmir], pl. 2, fig. 1. (description).

Remarks – Introduced as a nomen nudum, but subsequently the species was

described in the same work on the basis of an unstated number of syntypical speci-
mens of unstated sex originating from the region of Istanbul and Izmir.

Sitaris spectabilis (Meloidae)

Tünetes félmezű. Sitaris spectabilis. FRIVALD. – FRIVALDSZKY 1835a: 265, pl. VI, fig. 8. “Csak egy
példányban találtatott a’ Balkány’ déli oldalain” [Only a single specimen was found in the
southern sides of Stara Planina] (description).
Sitaris spectabilis, FRIV. – FRIVALDSZKY 1837a: 89, “Habitat in Balcani montosis” [Inhabits Stara
Planina] (redescription).
Tünetes félmezű (Sitaris spectabilis FR.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a
Haemus déli s éjszaki vidékei” [Southern and northern regions of Stara Planina, Sliven] (type
locality restriction).

Remarks – The species was described on the basis of a single specimen (=

holotype) of unstated sex collected in the southern sides of Stara Planina. Subse-
quently the type locality was restricted as southern and northern parts of Stara
Planina, near Sliven.

Stenostoma incana (Oedemeridae)

Deres karcs (Stenostoma incana FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo] (no description).

Remarks – Nomen nudum.

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Tanymechus orientalis (Curculionidae)

Keleti nyulár (Tanymechus orientalis FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo] (no description).

Remarks – Nomen nudum.

Trichodes obliquatus (Cleridae)

Ferde méhész. Trichodes obliquatus. BRÜLLÉ. – FRIVALDSZKY 1835a: 257, “Gyakori Rumeliában
a’ virágokon, Május ’s Juniusban” [Common on flowers in May and June in Rumelia], pl. V,
fig. 8. (description).
Trichodes obliquatus, FRIVALDS. – FRIVALDSZKY 1837a: 87, “Habitat in Balcani montosis” [In-
habits the mountains of the Balkans] (redescription).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex originating from Rumelia, collected in flowers in May
and June. The authorship of the name was attributed to the French entomologist
GASPARD AUGUSTE BRULLÉ (1809–1873).

Xsyletinus thoracicus (Anobiidae)

Toros rágoly. Xsyletinus thoracicus. FRIVALDSZKY – FRIVALDSZKY 1835a: 256 “Igen ritka a’
Balkány” hegyeinek déli mezein” [Very rare in the southern meadows of Stara Planina], pl. V,
fig. 7. (description).
Xyletinus thoracicus, FRIV. – FRIVALDSZKY 1837a: 87, “Habitat in Balcani montosis” [Inhabits
Stara Planina] (redescription).
Toros rágoly (Xyletinus thoracicus FR.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a Haemus
déli s éjszaki vidékei” [Southern and northern regions of Stara Planina, Sliven] (redescription).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex collected in the meadows of the southern Stara Planina.
Subsequently the type locality was restricted as southern and northern side of Stara
Planina, near Sliven.

Xylosteus spinolae (Cerambycidae)

Spinola-fakér. Xylosteus Spinolae. FRIVALDSZKY – FRIVALDSZKY 1837b: 180, “Rumeliában egy

példányban, sőt hazánk’ bánsági hegyeiben is találtatott” [A single specimen was found in Ru-
melia, furthermore also in the Banatian mountains of our country], pl. VII, fig. 10. (description).

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264 Zs. Bálint & S. Abadjiev

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex, which originated from “Rumelia” (a single syntype)
and from the “Banat” (several syntypes).

Zophosis pusila (Tenebrionidae)

Parányi sötény (Zophosis pusila FR.) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description).

Remarks – Nomen nudum.

LEPIDOPTERA

Acontia urania (Noctuidae)

Uránia sugárász. Acontia Urania. FRIVALDSZKY – FRIVALDSZKY 1835a: 274, “Röpül a’ Balkány’
dombjainak déli oldalain, Junius és Juliusban.” [It flies on the southern downs of the Balkans,
in June and July], pl. VII, fig. 10. (description).
Acontia Urania, FRIVALD. – FRIVALDSZKY 1837a: 92, “Habitat in Balcani montosis, Junio et Julio”
[Inhabits Stara Planina, in June and July] (redescription).
Uránia sugarász (Acontia urania FR.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a Haemus
déli s éjszaki vidékei” [Southern and northern regions of Stara Planina, Sliven] (type locality
restriction).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex collected in the southern downs of Stara Planina. Subse-
quently the type locality was restricted as southern and northern side of Stara
Planina, near Sliven.

Anthocharis gruneri (Pieridae)

Gruner-féle virma (Anthocharis Gruneri) – FRIVALDSZKY 1845: 170, “Macedonia hegyei, s bérczei
között” [In the mountains and cliffs of Macedonia] (no description); p. 178, “bythiniai Olym-
pus” [Olympus in Bythinia] (no description).

Remarks – Nomen nudum.

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 An annotated list of Imre Frivaldszky’s plant and animal names 265

Autophila pannonica (Noctuidae)

Pannóniai Virász. Autophila Pannonica, FRIV. – FRIVALDSZKY 1859: 21 (no description), 26,
“Puszta-peszéri erdő” [Forest of Pusztapeszér] (description).

Remarks – FRIVALDSZKY attributed the authorship of the name to himself and

presented a detailed description of not the imagines only, but also the early stages
and their life history.

Caloptera ocellata (Geometridae)

(Figs 5–6)

Bogláros röpék (Caloptera ocellata FR.) – FRIVALDSZKY 1845: 174, új nem! [new genus!] (no de-
scription), “Kréta…Sphakia hegylánczolata” [Mountain range of Sphakia, Crete]; 185,
“Hazája Kréta szigete, és Smyrna vidéke juniusban” [Inhabits Crete Island, and region of Izmir
in June], “Cordylion, Smyrnátul a tengeröblön át egy órányira” [Cordylion, one hour via the
bay from Izmir], pl. 3, fig 1. (description).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex originating from the “Sphakia” mountains of Crete, and
from the mountain Cordylion, near Izmir. The taxon is the type species of the ge-
nus Caloptera FRIVALDSZKY, 1845 by monotypy.

Catocala langvida (Noctuidae)

Halvány csallang (Catocala langvida FR.) – FRIVALDSZKY 1845: 172, “Constantinápoly kies vidé-
ke” [Surroundings of Istanbul] (no description).

Remarks – Nomen nudum.

Cleophana superba (Noctuidae)

Pompás sávoncz (Cleophana superba FR.) – FRIVALDSZKY 1845: 178, “bythiniai Olympus” [Olym-
pus in Bythinia] (no description).

Remarks – Nomen nudum.

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266 Zs. Bálint & S. Abadjiev

Dianthoecia seiuncta (Noctuidae)

Különváló Magtokász. Dianthoecia seiuncta HER. SCHEFF. (FRIV. in litt.) – FRIVALDSZKY 1865: 54,
“küldötteim előbb Törökországból hozták, későbben itt Ulma vidékén egy példányban talál-
ták” [my delegates first brought back it from Turkey, then they found a single specimen in the
region of Ulma) (no description).
Különváló Magtokász. Dianthoecia seiuncta HER. SCHEF. (FRIV. in litt.) – FRIVALDSZKY 1865: 152,
“E fajt eddigelé csak a Bánátból s Törökországból kaptuk” [Hithero we have found this species
only in the Banat and in Turkey], pl. V, fig. 6. (description).

Remarks – Introduced as a nomen nudum, but subsequently in the same work

the species was described from an unstated number of syntypical specimens of un-
stated sex, which originated from “Ulma”, in the “Banat” (a single syntype) and
from “Turkey” (several syntypes).

Erstria sphakiota (Noctuidae)

Sphakiai ledérke (Erstria Sphakiota FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegylánczo-
lata” [Mountain range of Sphakia, Crete] (no description).

Remarks – Nomen nudum.

Euprepia melana (Arctiidae)

Gyászos színdísz (Euprepia melana), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Re-
gion of Karlovo] (no description).
Gyászos színdísz. Euprepia melana. FRIVALDSZKY – FRIVALDSZKY 1837b: 171, “Calophir’ havas
aljában csak egy példányban találtatott” [At the foothills of Kaloferska Planina found only in a
single specimen], pl. VII, fig 1. (description).
Gyászos színdísz (Euprepia melana FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo].

Remarks – Introduced as nomen nudum from the region of Karlovo, but sub-
sequently in the same work the species was described on the basis of a single speci-
men (holotype) of unstated sex collected at the foothills of Kaloferska Planina.

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 An annotated list of Imre Frivaldszky’s plant and animal names 267

Euprepia placida (Arctiidae)

Tetszős színdísz. Euprepia placida. FRIVALDSZKY – FRIVALDSZKY 1835a: 271, “Találtatott Május
elején, a’ Balkány’ és Philippopolis’ vidékein” [It was found in the regions of the Balkans and
Plovdiv at the beginning of May], pl. VII, fig. 5. (description).
Euprepia placida, FRIVALDS. – FRIVALDSZKY 1837a: 91, “Habitat in Balcani montosis, Majo” [In-
habits Stara Planina, in May] (redescription).
Tetszős színdísz (Euprepia placida FR.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a
Haemus déli s éjszaki vidékei” [Southern and northern sides of Stara Planina, Sliven].

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex collected at the beginning of May in the Balkan regions
and in the vicinity of Plovdiv. Subsequently FRIVALDSZKY indicated the vicinity
of Stara Planina, near Sliven as collecting site of the species in slight contradiction
with the original description, which indicates a wider distribution.

Gasteropacha balcanica (Lasiocampidae)

Balkáni pohók (Gasteropacha balcanica FR.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke” [Re-
gion of Plovdiv] (no description).

Remarks – Nomen nudum.

Hadaena leuconota (Noctuidae)

Hópettyü émszép (Hadaena leuconota) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegylánczo-
lata” [Mountain range of Sphakia, Crete] (no description).

Remarks – Nomen nudum.

Hadena retina (Noctuidae)

Recze émszép (Hadena retina. FRIV.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegylánczo-
lata” [Mountain range of Sphakia, Crete] (no description).

Remarks – Nomen nudum.

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268 Zs. Bálint & S. Abadjiev

Heliotis decorata (Noctuidae)

Díszített naplász (Heliotis decorata FR.) – FRIVALDSZKY 1845: 177, “bythiniai Olympus” [Olympus
in Bythinia]; p. 186, “Brussa vidékén egy példányban találtatott” [A single specimen was
found in the region of Bursa]; pl. 3, figs 2a, 2b. (description).

Remarks – Introduced as nomen nudum, but subsequently in the same work

the species was described on the basis of a single specimen (= holotype) of un-
stated sex from the region of Bursa, most probably from the mountain “Olympus”
as the first mentioning of the taxon suggests.

Heliotis frivaldszkyi (Noctuidae)

Frivaldszky-naplász. Heliotis Frivaldszkyi, TREITSCHKE. – FRIVALDSZKY 1837b: 173, “a’ Balkány’

ormain” [On the cliffs of the Balkan], pl. VII, fig. 3. (description).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex from the “Balkan”. FRIVALDSZKY attributed the spe-
cies-group name to the contemporary Austrian lepidopterist GEORG FRIEDRICH
TREITSCHKE (1776–1842).

Heliothis incarnata (Noctuidae)

Vereslő naplász (Heliothis incarnata FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo] (no description).

Remarks – Nomen nudum.

Heliotis taurica (Noctuidae)

Tauriai naplász (Heliotis taurica TREITS.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a Haemus
déli s éjszaki vidékei” [Southern and northern regions of Stara Planina, Sliven] (no description).

Remarks – Nomen nudum.

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Heliothis treitschkii (Noctuidae)

Treitschke naplász. Heliothis Treitschkii. FRIVALDSZKY – FRIVALDSZKY 1835a: 273, “Találtatott

egy példányban Szlivnó lapos mezein, Juniusban” [A single specimen was found in the plain
region of Sliven, in June], pl. VII, fig. 8. (description).
Heliotis Treitschkii, FRIV. – FRIVALDSZKY 1837a: 91, “Habitat in Balcani montosis” [Inhabits Stara
Planina] (redescription).

Remarks – The species was described on the basis of a single specimen (=

holotype) of unstated sex collected on the plain near Sliven. The redescription indi-
cates a generalized locality, which contradicts the data accompaying the first de-
scription.

Hesperia phlomidis (Hesperiidae)

Bundi busók (Hesperia phlomidis FR.) – FRIVALDSZKY 1845: 178, “bythiniai Olympus” [Olympus
in Bythinia] (no description).

Remarks – Nomen nudum.

Hesperia tesselum (Hesperiidae)

Kockás busók (Hesperia tesselum) FRIV. – FRIVALDSZKY 1837b: 163, “Hortiát” (no description).

Remarks – Nomen nudum.

Hipparchia amalthea (Nymphalidae)

Amalthea pilleng (Hipparchia Amalthea FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia
hegylánczolata” [Mountain range of Sphakia, Crete] (no description), p. 186; “Hazája Kréta
szigete, az Ida és Sphakia hegység ormain” [Inhabits Crete Island, the cliffs of the Ida and
Sphakia mountains]; pl. 3, fig. 3 (male), 187, pl. 3, fig. 4. (female).

Remarks – Introduced as nomen nudum, but subsequently in the same work

the species was described from an unstated number of syntypical specimens of both
sexes originating from Crete, collected in the mountains of “Ida” and “Sphakia”.

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270 Zs. Bálint & S. Abadjiev

Hipparchia beroe (Nymphalidae)

Beroe pilleng (Hipparchia Beroe) – FRIVALDSZKY 1845: 178, “bythiniai Olympus” [Olympus in
Bythinia] (no description).

Remarks – Nomen nudum.

Hipparchia fatua (Nymphalidae)

Együgyü pilleng (Hipparchia fatua FRIV.) – FRIVALDSZKY 1845: 178, “bythiniai Olympus” [Olym-
pus in Bythinia] (no description).

Remarks – Nomen nudum.

Hipparchia thyrsis (Nymphalidae)

Thyrsis pilleng (Hipparchia Thyrsis FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegylánczo-
lata” [Mountain range of Sphakia, Crete] (no description).

Remarks – Nomen nudum.

Liparis terebinthii (Lymantriidae)

Terpentin csalyág (Liparis terebinthii FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo]; p. 172, “Constantinápoly kies vidéke” [Surroundings of Istanbul] (no description).

Remarks – Nomen nudum.

Lipparis undulata (Lymantriidae)

Habos csalyág (Lipparis undulata), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Re-
gion of Karlovo] (no description).

Remarks – Nomen nudum.

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Lycaena bellis (Lycaenidae)

Deli toportyán (Lycaena bellis FR.) – FRIVALDSZKY 1845: 172, “Constantinápoly kies vidéke” [Sur-
roundings of Istanbul] (no description).

Remarks – Nomen nudum.

Lycaena dardanus (Lycaenidae)

Dardán toportyán (Lycaena Dardanus) – FRIVALDSZKY 1845: 178, “bythiniai Olympus” [Olympus
in Bythinia] (no description).

Remarks – Nomen nudum.

Lycaena eroides (Lycaenidae)

Nőszded boglárka. Lycaena eroides. FRIVALD. –FRIVALDSZKY 1835a: 270 “Röpül Junius’ elején, a’
Balkány’ aljasabb mezein” [It flies in the lower meadows of Stara Planina], pl. VII, fig 3. (male
dorsum, ventrum), (description).
Lycaena éroides, FRIV. – FRIVALDSZKY 1837a: 91, “Habitat in Balcani montosis, Junio” [Inhabits
Stara Planina, in June].
Nőszded toportyán (Lycaena eroides FR.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke” [Region
of Plovdiv], “Constantinápoly kies vidéke” [Surroundings of Istanbul] (type locality restric-
tion).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex from the lower meadows of Stara Planina. Subsequently
the type locality was restricted as the regions of Plovdiv and Istanbul.

Lycaena hyacinthus (Lycaenidae)

Jáczint toportyán (Lycaena hyacinthus FR.) – FRIVALDSZKY 1845: 178, “bythiniai Olympus”
[Olympus in Bythinia] (no description).

Remarks – Nomen nudum.

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272 Zs. Bálint & S. Abadjiev

Lycaena iphigenia (Lycaenidae)

Iphigenia toportyán (Lycaena Iphigenia) – FRIVALDSZKY 1845: 178, “bythiniai Olympus” [Olym-
pus in Bythinia] (no description).

Remarks – Nomen nudum.

Lycaena ottomanus (Lycaenidae)

Ozmán toportyán (Lycaena ottomanus) – FRIVALDSZKY 1845: 172, “Constantinápoly kies vidéke”
[Surroundings of Istanbul] (no description).

Remarks – Nomen nudum.

Lycaena psittacus (Lycaenidae)

Kajdács toportyán (Lycaena psittacus FR.) – FRIVALDSZKY 1845: 176, “Cordylion, Smyrnátul a
tengeröblön át egy órányira” [Cordylion, one hour via the bay from Izmir] (no description).

Remarks – Nomen nudum.

Lycaena psylorita (Lycaenidae)

Psyloriti toportyán (Lycaena psylorita FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegylán-
czolata” [Mountain ridge of Sphakia, Crete] (no description).

Remarks – Nomen nudum.

Lycaena trochylus (Lycaenidae)

Delicze toportyán (Lycaena trochylus Fr.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegylán-
czolata” [Mountain range of Sphakia, Crete], p. 176, “Cordylion, Smyrnátul a tengeröblön át
egy órányira” [Cordylion, one hour via the bay from Izmir] (no description).

Remarks – Nomen nudum.

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 An annotated list of Imre Frivaldszky’s plant and animal names 273

Lycena sephirus (Lycaenidae)

Sephir boglárka. Lycena sephirus. FRIVALD. – FRIVALDSZKY 1835a: 269, “Röpül Junius elején , a’
Balkány’ aljas mezein” [It flies in the lower meadows of Stara Planina], pl. VII, figs 1. (male
dorsum, ventrum), 2. (female dorsum, ventrum) (description).
Lycaena cephirus. FRIV. – FRIVALDSZKY 1837a: 91, “Habitat in Balcani montosis, Junio” [Inhabits
Stara Planina, in June] (redescription).
Zefir toportyán (Lycaena zephyrus FR.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a Haemus
déli s éjszaki vidékei” [The southern and northern regions of Stara Planina, Sliven] (type local-
ity restriction).

Remarks – The species was described from an unstated number of syntypical

specimens of both sexes collected in the lower regions of Stara Planina. Subse-
quently the type locality was restricted as southern and northern side of Stara
Planina, near Sliven.

Ophiusa gentilis (Noctuidae)

Rokon márva (Ophiusa gentilis), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Region
of Karlovo] (no description); 172, “Karlova’ vidékén” [In the Region of Karlovo], pl. VII, fig.
2. (description).

Remarks – Introduced as nomen nudum, but subsequently in the same work

the species was described from an unstated number of syntypical specimens of un-
stated sex from the region of Karlovo.

Ophiusa gentiliata (Noctuidae)

Nemes márva (Ophiusa gentiliata FR.) – FRIVALDSZKY 1845: 174, “Kréta… Sphakia hegylánczo-
lata” [Mountain range of Sphakia, Crete] (no description).

Remarks – Nomen nudum.

Orgyia intermedia (Lymantriidae)

Hanga Nyutag közép válfaja. Orgyia ericae GERM. Var. intermedia FRIV. – FRIVALDSZKY 1865: 88,
“Lelhelye… a puszta-peszéri erdő, és a bánsági Belo-Berdó homok sivatag.” [Its locality is the
forest of Pusztapeszér and the sandy desert of Beloberdó in the Bánság] (description); 149, “A
peszéri erdőben és a határőrvidéki Grebenácz helység melletti homoksivatagon” [In the forest

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274 Zs. Bálint & S. Abadjiev

of Peszér and in the sandy desert near the settlement Grebenácz in the frontier zone], pl. V, figs
1a (male dorsum), b (female) c (coccon), d (caterpillar) (redescription).

Remarks – Briefly diagnosed first, then the species was described in details on
the basis of an unstated number of syntypical specimens of both sexes originating
from the Hungarian Plain (“Pusztapeszér”) and the “Bánság” région (“Grebenácz:
desert Beloberdo”). FRIVALDSZKY also described the early stages in great details.

Orthosia rorida (Noctuidae)

Hintett gubancz. Orthosia rorida. FRIVALDSZKY – FRIVALDSZKY 1835a: 272, “a’ Balkány’ alján,
gyümölcsös kertekben” [In orchards of the downs of Stara Planina], pl. VII, fig. 7. (descrip-
tion).
Orthosia rorida, FRIV. – FRIVALDSZKY 1837a: 91, “Habitat in Balcani montosis, Aprilis mense” [In
the mountains of the Balkans, in April] (redescription).
Hintett gubancz (Orthosia rorida FR.) – FRIVALDSZKY 1845: 165, “Szlivno (Selimno)… a Haemus
déli s éjszaki vidékei” [The southern and northern side of Stara Planina, near Sliven] (type lo-
cality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex from the lower regions of the Balkans, collected in or-
chards in April. Subsequently the type locality was restricted to southern and
northern regions of Stara Planina, near Sliven.

Plusia eximia (Noctuidae)

Kitünő éjdísz (Plusia eximia FR.) – FRIVALDSZKY 1845: 178, “bythiniai Olympus” [Olympus in
Bythinia] (no description).

Remarks – Nomen nudum.

Polia mensueta (Noctuidae)

Nyájas derécz (Polia mensueta FRIV.) – FRIVALDSZKY 1845: 176, “Cordylion, Smyrnátul a tenger-
öblön át egy órányira” [Cordylion, one hour via the bay from Izmir] (no description).

Remarks – Nomen nudum.

Annls hist.-nat. Mus. natn. hung. 98, 2006

 An annotated list of Imre Frivaldszky’s plant and animal names 275

Scardia auranciella (Tineidae)

Sárgácska biczer. Scardia auranciella. FRIVALDSZKY – FRIVALDSZKY 1835a: 272, “Röpülése’ ideje
Május, Junius, az erdők’ szélein a’ Balkány aljasabb vidékein” [It flies in May, June at forest
edges in the downs of Stara Planina], pl. VII, fig. 6. (description).
Scardia auranciella, FRIV. – FRIVALDSZKY 1837a: 91, “Habitat in Balcani montosis, Majo, Junio”
[Inhabits Stara Planina, in May, June] (redescription).
Sárgálló biczer (Scardia auranciella FR.) – FRIVALDSZKY 1845: 167, “Philipopoly vidéke” [Region
of Plovdiv] (type locality restriction).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex flying in May and June at forest edges in the downs of
Stara Planina. Subsequently the type locality was restricted as region of Plovdiv.

Symira tendinosa (Noctuidae)

Inas czoboly (Symira tendinosa), FRIVALD. – FRIVALDSZKY 1837b: 161, “Karlova vidéke” [Region
of Karlovo] (no description).
Inas czoboly (Symira tendinosa, FR.) – FRIVALDSZKY 1845: 168, “Karlova vidéke” [Region of
Karlovo] (no description).

Remarks – Nomen nudum.

Xsylina freyeri (Noctuidae)

Freyer kérgecs. Xsylina Freyeri. FRIVALDSZKY – FRIVALDSZKY 1835a: 273, “Találtatott a’ Bal-
kány’ hegyeinek déli oldalában, Juniusban” [It was found on southern side of Stara Planina, in
June], pl. VII, fig. 9. (description).
Xylina Freyeri, FRIV. – FRIVALDSZKY 1837a: 92, “Habitat in Balcani montosis, Junio” [Inhabits
Stara Planina, in June] (redescription).

Remarks – The species was described from an unstated number of syntypical

specimens of unstated sex from the southern side of Stara Planina. The re-
description indicated June as flight period.

Xylina orientalis (Noctuidae)

Keleti kérgecs (Xylina orientalis FR.) – FRIVALDSZKY 1845: 176, “Cordylion, Smyrnátul a tenger-
öblön át egy órányira” [Cordylion, one hour via the bay from Izmir] (no description).

Remarks – Nomen nudum.

Annls hist.-nat. Mus. natn. hung. 98, 2006
276 Zs. Bálint & S. Abadjiev

Zigena sedi (Zygaenidae)

Szaka igácz. Zigena sedi. FABR. – FRIVALDSZKY 1835a: 271, “Röpülése’ ideje Junius, a’ Balkány’
déli vidékin” [Its flight period is June, in the southern regions of the Balkans], pl. VII, fig. 4.
(description).
Zygaena Sedi, FRIVALDS. – FRIVALDSZKY 1837a: 91, “Habitat in Balcani montosis, Junio” [Inhabits
Stara Planina, in June] (redescription).

Remarks – Described on the basis of an unstated number of syntypical speci-

mens of unstated sex collected in June in the southern regions of Stara Planina. The
authorship of the species-group name first was attributed to the Danish entomolo-
gist JOHANN CHRISTIAN FABRICIUS (1745–1808).

AVES

Columba decaocto (Columbidae)

Különös gerlicze faj – FRIVALDSZKY 1837b: 159, “Philippopolnak … távolabb környéki, … mint
Tatárbazarcsik’, Despoto, Rilo’s Dupincza’ magas hegyei, Szamunkov’ kies, tágas völgye”
[Wider region of Plovdiv, Pazardzhik, Rhodope, the higher parts of Rila near Dupnitsa, the pic-
turesque and wide valley of Samokov] (nomen nudum); 183, Félholdas galamb, Columba
decaocto, pl. VIII. (description).
A kaczagó galambnak válfaja, mellyet némellyek a házi kaczagó galamb eredeti fajának tartanak. –
FRIVALDSZKY 1845: 166, 167, “Philippopoly vidéke…, Despoto, Rodope, Rilo, s Staminák
hegység lánczolatai… Az említett hegyláncolaton, s a Maricza folyó mentében, valamint
délkeletre a Drinápoly felé elnyúló lapályon” [Region of Plovdiv, mountain ridges of Rhodope,
Rila and Staminaka, along the river Maritsa; and on the plain extending southeast from Edirne].

Remarks – First introduced as a nomen nudum from the region of Plovdiv,

from the settlement Pazardzhik and Rhodope Planina, from the higher mountain
ranges of Rila and from the picturesc valley near Samokov, but subsequently in the
same work the species was described on the basis of an unstated number of
syntypical specimens of unstated sex. The locality of the type material was not
given, but the localities where the species have been taken were listed previously
on p. 159, therefore those places can be applied as type locality in accordance with
the ICZN Code Art. 76.1. Later FRIVALDSZKY gave a wider distribution of the spe-
cies, but these mirrored the results of the subsequent expeditions.

Annls hist.-nat. Mus. natn. hung. 98, 2006

 An annotated list of Imre Frivaldszky’s plant and animal names 277

GAZETTEER FOR HISTORICAL NAMES

First the geographical name is given as it appears in the main body of the text
(with indication whether it belonged to historical Hungary or/and the Ottoman
Empire), then the recent official name (sometimes also its English equivalent in
brackets) is given, with indication to the present political situation.
Alföld, plain (Hungary, Ottoman Empire) = Alföld [The Great Hungarian Plain] (Hungary, Romania,
Serbia)
Balkan, mountains (Ottoman Empire) = Stara Planina (Bulgaria)
Balkans, mountains (Ottoman Empire) = Balkan Peninsula
Balkány mountains (Ottoman Empire) = Balkan Peninsula
Bánát, region (Hungary) = Banat (SW Romania and N Serbia)
Banjo, mountain (Ottoman Empire) = most probably the mountain near village Banja, in the vicinity
of Karlovo (Bulgaria)
Bánság, region (Hungary) = Banat (SW Romania and N Serbia)
Békés, county (Hungary) = Békés megye (Hungary)
Beleberdó, “desert” (Hungary) = pustina Bieloberdo (Vojvodina, Serbia)
Bihar, county (Hungary) = judeţul Bihor (Transylvania, Romania)
Bizáncz, city (Ottoman Empire) = Istanbul (Turkey)
Bohodej, mountain (Hungary) = vârful Bohodei in judeţul Bihor (Transylvania, (Romania)
Brussa, town (Ottoman Empire) = Bursa (Turkey)
Buda, mountains (Hungary) = Budai-hegység (Hungary)
Buda-Pest, city (Hungary) = Budapest (Hungary)
Bythinia, region (Ottoman Empire) = Bythinia, historical part of N Turkey
Calophir, mountains (Ottoman Empire) = Kaloferska Planina, a part of the central region of Stara Pla-
nina (Bulgaria)
Canea, town (Ottoman Empire) = Khaniá (Crete, Greece)
Carlova, town (Ottoman Empire) = Karlovo (Bulgaria)
Constantinápoly, city (Ottoman Empire) = Istanbul (Turkey)
Cordylion, mount (Ottoman Empire) = Ulu-dagh (Turkey)
Csabacsüd, village (Hungary) = Csabacsüd (Békés megye, Hungary)
Cserna, river (Hungary) = Râul Cernei (Banat, Romania)
Csorna, river (Hungary) = Râul Cernei (Romania)
Dervent, heights (Ottoman Empire) = Derventski Vazvisheniya (Bulgaria)
Despoto, village (Ottoman Empire) = unknown place, probably in Rodopi Planina (Bulgaria)
Drinápoly, town (Ottoman Empire) = Edirne (Turkey)
Dupincza, town (Ottoman Empire) = Dupnitsa (Bulgaria)
Fericse, settlement (Hungary) = Ferice (judeţul Bihor, Romania)
Filipopol, town (Ottoman Empire) = Plovdiv (Bulgaria)
Filippopol, town (Ottoman Empire) = Plovdiv (Bulgaria)
Fonácza, cave and settlement (Hungary) = Fânaţe (judeţul Alba, Romania)
Funacer Grotte, cave (Hungary) = Fânaţe (judeţul Alba, Romania)
Grebenácz, settlement (Hungary) = Grebenac (Vojvodina, Serbia)
Guklina (Ottoman Empire) = unknown place, most probably in the vicinity of Plovdiv (Bulgaria)
Haemus, mountains (Ottoman Empire) = Stara Planina (Bulgaria)
Herkulesfürdő, town (Hungary) = Bâile Herculane (Banat, Romania)

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278 Zs. Bálint & S. Abadjiev

Hortiát (Ottoman Empire) = unknown place, most probably in the vicinity of Thessaloniki (Greece)
Ida, mountain (Ottoman Empire) = Psiloritis, Crete (Greece)
Igritz, cave (Hungary) = Peştera Igriţa, judeţul Bihor, Transylvania (Romania)
Islas, village (Hungary) = Islesniţa (Romania)
Jóniai szigetek, islands (Ottoman Empire) = Ionia Nisia [Ionian Islands] (Greece)
Karlova, town (Ottoman Empire) = Karlovo (Bulgaria)
Kazán-szoros, narrows (Hungary-Ottoman Empire) = Cheile Kazane (Romania, Serbia)
Kazánlik, town (Ottoman Empire) = Kazanlak (Bulgaria)
Korniareva, mount (Ottoman Empire) = Corniareva (Banat, Romania)
Krassó, county (Hungary) = judeţul Caraş-Severin (Romania)
Kréta, island (Ottoman Empire) = Kriti [Crete] (Greece)
M. sziget, town (Hungary) = Sighetul Marmaţiei (Romania)
Macedonia, region (Ottoman Empire) = Makedonia (Greece)
Máramaros, county (Hunagray) = judeţul Maramureş (Romania)
Máramarossziget, town (Hungary) = Sighetul Marmaţiei (Romania)
Maricza, river (Ottoman Empire) = Maritsa (mainly in Bulgaria)
Mehádia, town (Hungary) = Mehadia (judeţul Caraş-Severin, Romania)
Olympus, mountain (Ottoman Empire) = Ulu-dagh (Turkey)
Philipopoly, town (Ottoman Empire) = Plovdiv (Bulgaria)
Philippololy, town (Ottoman Empire) = Plovdiv (Bulgaria)
Philippopoly, town (Ottoman Empire) = Plovdiv (Bulgaria)
Poó, puszta (Hungary) = Poó-puszta, near the town Mezőtúr (Jász-Nagykun-Szolnok megye, Hun-
gary)
Puszta-peszér, village (Hungary) = Kunpeszér (Bács-Kiskun megye, Hungary)
Pusztapeszér, village (Hungary) = Kunpeszér (Bács-Kiskun megye, Hungary)
Rákos, “desert” (Hungary) = part of Budapest (Hungary)
Rézbánya, town (Hungary) = Bâiţa (judeţul Bihor, Romania)
Rhodoppe, mountains (Ottoman Empire) = Rodopi Planina [Rhodope Mountains] (Bulgaria)
Rilo, mountains (Ottoman Empire) = Rila Planina [Rila Mountains] (Bulgaria)
Rodope, mountains (Ottoman Empire) = Rodopi Planina [Rhodope Mountains] (Bulgaria)
Rumelia, region (Ottoman Empire) = Iztochna Rumelia [historical part of Southern Bulgaria] (Bul-
garia)
Salonik, town (Ottoman Empire) = Thessaloniki (Greece)
Selimno, town (Ottoman Empire) = Sliven (Bulgaria)
Smyrna, city (Ottoman Empire) = Izmir (Turkey)
Sphakia, county (Ottoman Empire) = Sfakia (Crete, Greece)
Staminák, town (Ottoman Empire) = Asenovgrad (formerly Staminaka, Bulgaria)
Szádelő, gorge (Hungary) = Zadiel (Slovakia)
Szalonik, town (Ottoman Empire) = Thessaloniki (Greece)
Szamunkov, town (Ottoman Empire) = Samokov (Bulgaria)
Szarvas, town (Hungary) = Szarvas (Békés megye, Hungary)
Szlavónia, region (Hungary) = Slavonija (Croatia)
Szliven, town (Ottoman Empire) = Sliven (Bulgaria)
Szlivno, town (Ottoman Emire) = Sliven (Bulgaria)
Szokolovác (Hungary) = Sakalova (Vojvodina, Serbia)
Tatárbazarcsik, town (Ottoman Empire) = Pazardzhik (Bulgaria)
Tericse, settlement (Hungary) = Ferice (judeţul Bihor, Romania)
Torna, county (Hungary) = Turna (Slovakia)

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 An annotated list of Imre Frivaldszky’s plant and animal names 279

Tundzsa, river (Ottoman Empire) = Tundzha (Bulgaria)

Ulma, town (Hungary) = Ulma (Vojvodina, Serbia)

*
Acknowledgements – We express our sincere thanks for the following colleagues who helped
us in various ways in this paper: OTTÓ MERKL (Coleoptera, HNHM), GÁBOR PAPP (Bibliography
and Gazetteer, HNHM) and GYÖRGY SZOLLÁT (Plants, HNHM). We are indebted to JÁNOS
FRIVALDSZKY, a direct descendant of the “great FRIVALDSZKYs”, IMRE and JÁNOS, for his help in
autobiographic questions. The study in the Archives of the Natural History Museum (London) was
made possible for ZS. B via a fellowship grant of the Royal Society in June–August, 2003. We thank
the host PHILLIP ACKERY (Butterfly Collection) for his generosity and invaluable assistance. In the
collections of the Museum National d’Histoire Naturelle (Paris) it was also possible to work via the
curator JACQUES PIERRE (Laboratoire d’Entomologie) during the summer of 2005 for ZS. B. We also
express our gratitude to him.

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FRIVALDSZKY, E. 1834: Catalogus insectorum disponibilium Doctoris Emerici Frivaldszky, – Typis

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ból. [An essay on the wandering locuts from the point of view of natural history and economy.]
– Buda, iv + 39 pp, 2 plates.
FRIVALDSZKY, I. 1854: Wirbellose Thiere. Pp. 21–27. – In: HAEUFLER, J. V.: Buda-Pest, histo-
risch-topographische Skizzen. – Emich Gusztáv, Pest, [4] + 131 + [1] + 82 + [2] + 322 + [2] + 5
plates + 4 appendices.
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Természettudományi Közlemények 1863: 231–240.
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neten Pflanzen aus Türkei. – Flora, Allgemeine botanische Zeitung 3: 225–234.
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Zoological Nomenclature, London, xxix + 306 pp.
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naturales Musei nationalis hungarici 94: 5–29.
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Annls hist.-nat. Mus. natn. hung. 98, 2006