Cactus Explorer 17 - Complete

The
Cactus Explorer
The first free on-line Journal for Cactus and Succulent Enthusiasts
1 Siccobaccatus
2 Morangaya pensilis
Number 17 3 Espostoa in Tenerife
ISSN 2048-0482 4 Barranco Rambla de Ruiz

December 2016 5 Juab and Utah County
The Cactus Explorer ISSN 2048-0482
IN THIS EDITION
Number 17 December2016
Regular Features Articles
Introduction 3 A naturalised population of

News and Events 4 Espostoa melanostele on Tenerife 21
In the Glasshouse 9 Travel with the cactus expert (16) 25
Journal Roundup 14 Where lizards dare: an excursion to Barranco
On-line Journals 15 Rambla de Ruiz (Tenerife) 29
The Love of Books 18 Juab and Utah County, Utah, throughout the
Society Pages 51 year 2015 36
Plants and Seeds for Sale 55 A Happy Medium? Morangaya pensilis. 41
Books for Sale 62
Cover Picture: Siccobaccatus dolichospermaticus See page 9

The No.1 source for on-line information about cacti and succulents is http://www.cactus-mall.com
The best on-line library of succulent literature can be found at:
https://www.cactuspro.com/biblio/en:accueil
Invitation to Contributors
Please consider the Cactus Explorer as the place to publish your articles. We welcome
contributions for any of the regular features or a longer article with pictures on any aspect of
cacti and succulents. The editorial team is happy to help you with preparing your work. Please
send your submissions as plain text in a ‘Word’ document together with jpeg or tiff images with
the maximum resolution available.
A major advantage of this on-line format is the possibility of publishing contributions quickly
and any issue is never full! We aim to publish your article quickly and the copy deadline is just a
few days before the publication date. There will usually be three issues per year, published when
sufficient material is available. Please note that advertising and links are free and provided for
the benefit of readers. Adverts are placed at the discretion of the editorial team, based on their
relevance to the readership.
Publisher: The Cactus Explorers Club, Briars Bank, Fosters Bridge, Ketton, Stamford, PE9 3BF U.K.
The Cactus Explorer is available as a PDF file downloadable from www.cactusexplorers.org.uk
The Editorial Team:
Organiser:Graham Charles [email protected]
Paul Hoxey [email protected] This issue published on
Zlatko Janeba [email protected] 5th December 2016
Martin Lowry [email protected]
Opinions expressed in the articles are those of the authors, and not necessarily those of the editorial team.
©The Cactus Explorer may be freely distributed whilst the copyright of the text and pictures remains with the
authors and photographers. Permission is required for any use other than reading, printing or storage.
2
ISSN 2048-0482 The Cactus Explorer
INTRODUCTION
Number 17 December 2016
Time to think about buying seeds matucanas in preparation for my next project,
a book about the genera Matucana and Oroya.
Now that another growing season has come
The only specialist book on the subject, written
to an end here in England, I can turn my
by Rob Bregman, was published in 1996. It is
attention to the garden and indoor activities. I
now quite difficult to find for sale, and there
have just finished producing Bradleya 34
have been significant new discoveries since it
which, like last year, was very short of material
was written so it’s time for a new one.
at the start of August but after a hectic couple
of months, ended up at a record size of 240 I would really appreciate hearing from
pages. It is a rich legacy of which the BCSS anyone who has found matucanas in habitat or
should be proud. I say a big ‘Thank You’ to the has good pictures of documented plants in
many members who subscribe to it, for cultivation that I could use in the book.
supporting a valuable resource of knowledge Raising plants from seeds must surely be
for the future. one of the best parts of our hobby. It gives you
The contents of Bradleya are the usual broad the chance to grow plants which are not easily
mix of subjects but my particular interest in available any other way. Earlier, I mentioned
cacti is especially catered for by a very useful the genus Cleistocactus. Anyone reading
review of Cleistocactus by Martin Lowry and a Martin’s article in Bradleya will realise that only
learned explanation of what constitutes a a few of the many species are readily available
cephalium by Root Gorelick. You can see more for sale in the trade. At the end of this edition,
information about the contents and how to buy you will find lots of advertisements for seeds
Bradleya 34 on page 7. and many of these dealers offer species that are
rarely offered for sale as plants.
Thinking of the succulent experts who
generously contribute articles to Bradleya It is really exciting to grow seedlings of
reminds me of the sad death of Heidi plants you have never seen before. I remember
Hartmann earlier this year. She was a regular when the new Lobivia species discovered by
contributor of detailed scientific studies to Walter Rausch were first offered as seeds and
Bradleya and I very much enjoyed working watching the young plants develop into plants
with her to get everything just right. Her I didn’t recognize!
friendly and very polite emails sometimes Please remember to send me pictures of
referred to the health problems which had unusual plants that have flowered in your
affected her in recent years. collection to share with other readers.
Then, only last month, I heard of the sad
death of Darrel Plowes at the age of 91. Just a
GrahamCharles
few days before I had been in communication P.S. Those of you who attend the Cactus
with him about a paper he was working on for Explorers Club Weekend will be pleased to
Bradleya about the stapeliad genus Angolluma. know that I have booked it for next year, the
His health was already failing and regrettably weekend of September 15–17th 2017. The
he never finished the work. You can find an invitations to regular participants will be sent
appreciation of Darrel on page 4. out in January.
At last, I have got around to rationalising If you have not already told me and would
my gymnocalycium collection. I accumulated like to be advised when the next issue of the
hundreds of speciments which I used to Cactus Explorer is available for down-
research and illustrate my book. Now I need load, please send me your E-mail address to
the space to grow other plants, notably be added to the distribution list.
3
NEWS AND EVENTS

†Plowes, Darrel Charles Herbert A. Castellanos & H. Lelong

(1925-2016) Publications devoted to cacti
b. Estcourt, Natal, 4 Apr 1925; This, the second e-book (416 pages)
published by Au Cactus Francophone is
Mutare, Zimbabwe, 19 Oct 2016.
devoted to articles by Castellanos and
Lelong. The introduction presents the life of
The celebrated 91-year old veteran of both authors who were husband and wife.
stapeliads, Darrel Plowes, finally After the series of articles, some original
succumbed to a 2-day bout of pneumonia pictures by Castellanos are reproduced.
on the night of 19 Oct., leaving two This very useful reference for Argentinian
children from his first marriage, Carolyn cacti can be downloaded (76Mbyte) from
& Robert, and Mrs. Nina Bauer, with https://www.cactuspro.com/biblio_fichiers/p
whom he had been sharing a home df/KieslingRoberto/CastellanosLelong.pdf
during his final few years.
Following wartime service, Darrel Bradleya Index
graduated from Witwatersrand
University in 1948 with a degree in
agriculture, soil conservation and ecology,
An index to Bradleya
and then emigrated in 1949 to Zimbabwe, 1–34 has been
where he occupied several positions, first compiled by Roy
as a Research Officer, then Superintend- Mottram and can be
ent, and finally as a Regional Director of downloaded as a
African Agriculture in Zimbabwe’s searchable PDF file
Manicaland Province.
When not involved in his career work,
he devoted himself to studying the
natural history of Zimbabwe, sometimes
www.cactusexplorers.org.uk/Explorer17/Bra
being named for several plants and
dleya Index 1–34.pdf
animals of his own discovery, and became
a leading light in the National Trust of Thank you Roy!
Zimbabwe, from where tributes are
already pouring in. His particular Cactáceas y Suculentas
speciality in stapeliads came rather later Mexicanas
in life, but he soon made up for that by The Mexican journal has been published
becoming an internationally recognised since 1955 with the aim of sharing knowledge
authority in their classification, describing about succulent plant families, especially the
numerous new taxa. His level of Cactaceae, and to promote interest and
classification was considered by many to research on different aspects of these amazing
be too liberal, but he was a careful, hands- plants.
on observer and his work could never be The journal is available on line and free at:
dismissed lightly. web.ecologia.unam.mx/cactsucmex/
Any comments please contact: Dra. Mariana
Roy Mottram
Rojas-Aréchiga ([email protected])
4
Number 17 December 2016 ISSN 2048-0482 The Cactus Explorer
What a Wonderful Day!
The BCSS National Show, August 20th 2016

This is the day when the BCSS feels at its best. It is not At a time when we hear about the worry of reducing
just the plants on show, nor the plants for sale, but the membership and branches struggling to keep going, it is
chance to meet friends and enjoy the atmosphere of a encouraging to see how many people will turn out to
successful event. This one must have hosted the biggest this outstanding event. What a shame it is only every 4
sale of succulent plants ever held in the UK and the years. It is said that it cannot be held more often because
whole event was improved by the longer opening hours of all the work, but I wonder if someone would cham-
and the opportunity for exhibitors to buy plants on the pion the cause for a more frequent event?
Friday afternoon. GC
All of Duke’s books are printed on high-quality 150 GSM, Lumi Silk Art paper, and all are high-quality-hardcover bound.
Duke Benadom’s Superb Succulents is Duke’s Echinocereus is 8! " 11” (22 " Duke’s Southwest Deserts is 8! " 11” (22
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full-color photographs, 236 pages, based photographs, 500 pages, based on photographs, 384 pages, based on
on the popular, long-running column, decades of personal research on the decades of peregrination throughout our
Superb Succulents as seen in the Cactus genus Echinocereus. southwest deserts.
& Succulent Journal. $59.95 $79.95 $74.95 $74.95
Volume 1 of The Cycads is on cycads of Asia & Australia. It Volume 2 is on cycads of Africa & The Americas. It includes
includes 460 pages and 473 photographs, covering the genera 500 pages and 820 photographs, covering the genera of Dioon,
of Bowenia, Cycas, Lepidozamia, & Macrozamia. Ceratozamia, Encephalartos, Stangeria, & Zamia.
$89.95 $89.95
Cycad volumes are $89.95 each, but are discounted to $154.95 per set (both volumes).
Collector’s editions of Superb Succulents, Echinocereus, & Southwest Deserts are available at www.SuperbSucculents.net for
an additional $30 each. The Collector’s edition is essentially the same as the standard edition, but with foil stamping on the spine and
on the front & back and with 135gsm Egyptian Dynic Saifu cloth over 3mm Graphic Board on the book. There is also a slipcase with
4-color gloss laminate on 130gsm art paper. In summary, it’s the hardbound edition with a special cloth cover, gold or silver foil
imprinting (depending on the cloth color), and with a quality slip case. These sets also come individually shrink wrapped.
5
Order these great books from www.superbsucculents.com
The Cactus Explorer ISSN 2048-0482 Number 17 December2016
The Haworthia Society

Haworthiad on DVD
A single searchable file of 3,572 pages contains all
the issues of Haworthiad from 1986 to 2015
(Volumes 1 to 29) including the Special Editions.
The DVD may be ordered through our PayPal Shop

http://haworthia.org/shop.php or alternatively,
contact the Treasurer: [email protected]
Members £30.00 (UK), £31.00 Europe,

£32.00 (Rest of World),
Non-members £40.00 (UK0, £41.00 (Europe),
£42.00 (Rest of World).
All prices include postage and packing.
6
Bradleya 34
Yearbook of the British Cactus & Succulent Society

The 2016 issue of Bradleya is now on sale.
23 well-illustrated articles to enjoy including
7 about cacti. 240 pages.
You can buy Bradleya 34, by sending £21

(£23 overseas) including post & packing to
the BCSS Publications Manager, Brenfield, Bolney
Road, Ansty, West Sussex, RH17 5AW, UK.
Payment accepted by:

£ sterling cheques drawn on a UK bank
(payable to BCSS)
PayPal ([email protected]) or
Credit/debit card (Visa, Mastercard or Maestro)
The articles in this edition are:

An extension to the known distribution of What is a cephalium?
Austrocylindropuntia pachypus in Peru
Tinospora fragosa subsp. fragosa adaptation to its
The correct author citation and date of publication hostile African habitat
of the name Agave yuccifolia
Notes on the geographical distribution range of
A further species of Furcraea: Furcraea selloana, Sedum mucizonia, a miniature, annual succulent,
naturalised in South Africa in continental Portugal
Cylindropuntia fulgida var. fulgida is naturalised and Correction of a nomenclatural problem in
spreading in Zimbabwe Cotyledon, and notes on the taxonomy of
miscellaneous succulent Hyacinthaceae
Widely cultivated, large-growing yuccas: notes on
Yucca elephantipes, Yucca gigantea and Yucca Permanent open flowers in Nelia – an approach to
guatemalensis the secret
Cactus survey at the Floresta Nacional of A synopsis of the genus Cleistocactus
Contendas do Sincorá, Bahia, Brazil
Notes on the cacti of the Sandia valley, Puno, Peru
Austrocylindropuntia vestita recorded from the Little and a description of a new species of Echinopsis
Karoo in SA’s Western Cape province
A review of Agave mitis
Reinstatement of Aloe candelabrum, a tree-like aloe
Kalanchoe winteri, a new species from the Wolkberg
of KwaZulu-Natal province, South Africa
Centre of Endemism, South Africa
Occurrence of the little-known Kalanchoe leblanciae
Sansevieria ballyi and newly found field notes
confirmed in South Africa
Dracaena transvaalensis, the dragon tree of the
A review of Euphorbia mlanjeana: its habitats on
Limpopo Province, South Africa
Mount Mulanje (Malawi) and new localities in
Mozambique Tylecodon florentii, a new cliff dwelling species from
the Kookrivier cliffs, Richtersveld, SA
The taxonomy and type of Kalanchoe sexangularis
7
Guided Trip to Baja California

If you would like to see cacti in amazing scenery then Baja is a good choice. One of our regular
Cactus Explorers, Daina Delladdio, lives in Mexico and will be leading a trip to see cacti and
wildlife from 12th to 26th February 2017.
It will be a 15 day adventure trip dedicated to the discovery of Baja California and its
surroundings. This is where you'll experience the extraordinary biodiversity and enchanting
landscapes of this area: remote beaches, multicoloured deserts, breathtaking vistas, centennial
cacti, endemic plants, and the wonderful grey whales that often approach you.
For information on the itinerary please see this download or contact [email protected]
https://www.facebook.com/events/1211997358867677/
Copiapoa decorticans Copiapoa ahremephiana Copiapoa atacamensis Copiapoa laui
Your chance to buy cards illustrated with reproductions of pictures from

CACTACEAS En la flora silvestre de Chile
by Adriana Hoffmann and Helmut Walter.
Each card is 124 x 174mm and is blank inside for your own message.
Set of any four designs of your choice: £10 plus postage (UK: £0.64; EU: £1.52; World: £2.25)
Orders and PayPal payments to Roger Ferryman email: [email protected]
Eriosyce aspillagae maechleri Eriosyce napina challensis Copiapoa krainziana Eriosyce spinibarbis
8
IN THE GLASSHOUSE
Siccobaccatus, a magnificent cactus genus from Brazil

Kamiel Neirinck tells us about one of the most remarkable and beautiful colum-
nar cacti. Translation by R. Fonteyne. Photographs by the author except where stated.
Fig.1 Siccobaccatus dolichospermaticus flowering in the collection of K.J. Neirinck.

The genus Siccobaccatus was set up by Braun Siccobaccatus dolichospermaticus, the type
and Esteves and described by them in species, with reddish spination, was described
Succulenta in 1990. There are only three very in 1974 by Buining and Brederoo as an
nice species which were discovered from the Austrocephalocereus. Buining and Horst first
early 1970s onwards. In habitat these columnar found this plant in west Bahia in Brazil. Later
cacti reach several meters high and have a in the 1980s it was also found in adjacent
striking appearance because of their size, the Minas Gerais. In 1999, also in southwest Bahia,
nice colour of the epidermis, the spination and thousands of plants were discovered by Braun
the lateral cephalium. They do not branch and and Esteves on a huge 120km long limestone
always grow on very remote, isolated places massif. Thus, they are far from extinction.
on bambui-limestone rock outcrops. Siccobaccatus estevesii, which grows twice as tall
9
Photo: Graham Charles
Fig.2 Siccobaccatus dolichospermaticus in habitat near to Bom Jesus de Lapa. GC1014.03.

as S. dolichospermaticus, was discovered
somewhat later in Goias by Eddie Esteves and
Photo: Graham Charles

described in his honour by Buining and
Brederoo.
Two more subspecies of S. estevesii were
described in the 1980s: ssp. grandiflorus from
the state of Tocantins and ssp. insigniflorus
from southeast Minas Gerais, the latter raised
to species level in 2008. Taylor and Zappi
consider Siccobaccatus as a subgenus of
Micranthocereus, but this is not supported by
Braun and Esteves, nor by molecular studies
which suggest a closer relationship with
Coleocephalocereus. This is why seeds and plants
of Siccobaccatus species are sometimes offered
under the genus name Micranthocereus.
Siccobaccatus dolichospermaticus is very difficult
in culture and is preferably grafted, while S.
estevesii can more easily be grown on its own
roots. "Siccobaccatus" means "with dry fruits",
and "dolichospermaticus" means "with
elongated seeds".
As with most cerei, the flowers are not Fig.3 Siccobaccatus dolichospermaticus
spectacular. Compared with the general GC1014.03 near to Bom Jesus de Lapa.
10
appearance of the plants they are rather
modest. They flower at night and have no
scent. Insects and birds are attracted for
pollination by the white colour. In Europe
Siccobaccatus dolichospermaticus starts to flower
from a lateral cephalium at a height of one
metre.
The species are:
Siccobaccatus dolichospermaticus.
Siccobaccatus estevesii with subspecies estevesii
and subspecies grandiflorus.
Siccobaccatus insigniflorus.
Siccobaccatus dolichospermaticus
Siccobaccatus dolichospermaticus was first
described by Buining and Brederoo as
Austrocephalocereus dolichospermaticus in Kakteen
und andere Sukkulenten in 1974, then later
included by Braun and Esteves in their new
genus Siccobaccatus, published in Succulenta in
1990.
Description
Body: columnar, only branching or offsetting
when the plant is damaged, up to 2m high,
sometimes 3m and higher, up to 8cm thick,
glaucous. The root system is branched off in
rock crevices. Cephalium: up to 6cm wide,
lateral, uninterrupted, with cream-coloured
4cm long wool and many 5cm long bristles.
The cephalium can already be formed at a
height of 40cm. Ribs: about 30, 9–10mm wide
and 10mm high. Areoles: oval, 7mm long and
5mm wide, 2–3mm apart, initially covered
with light felt, later naked. Radial spines:
flexible, 4–7mm long, yellow or reddish.
Central spines: 6–8, yellowish to light brown, up
to 25mm long. Flowers: tubular, more
campanulate towards the top, 40mm long and
25mm wide, naked, night-flowering. Fruits:
7mm long, 9mm wide, elongate-turbinate,
cream-coloured, dry berry with flower-
remains. Ripe fruits lose their lid, the open
fruit with the free seeds remains in the
cephalium, hence the name Siccobaccatus. Seeds:
oblong, 2mm long and 0.5mm wide, the colour
Fig.4 Siccobaccatus dolichospermaticus in the is chestnut brown.
collection of Gerhard Heimen.
11
Distribution Literature
Brazil, federal state Bahia, type locality west BRAUN, P.J. & EDDIE ESTEVES PEREIRA (1990).
of Bom Jesus de Lapa at a height of 460m, Siccobaccatus Braun et Esteves: gen. nov. Een
amongst others together with Melocactus nieuw cactusgeslacht uit Brazilie. Succulenta
levitestatus, several Pilosocereus species and 69(1): 3–8.
bromeliads. The plants grow on limestone BRAUN, P.J. & EDDIE ESTEVES PEREIRA (2001).
rocks. Also found in Minas Gerais by Braun Kakteen und andere Sukkulenten in
and Esteves. Brasilien. Schumannia 3.
Cultivation BRAUN, P.J. & EDDIE ESTEVES PEREIRA (2008).
Even in South and Central Brazil top Siccobaccatus insigniflorus. A new status for a
cuttings do not root. 30 years ago I grafted marvelous columnar cactus from Brazil.
such an imported cutting (at that time it was Cactus & Succulent Journal (US) 80(1):36–41.
allowed to import them) on a thick Trichocereus BUINING, A.F.H. & BREDEROO, A.J. (1974).
macrogonus. The stock, and finally the plant Austrocephalocereus dolichospermaticus spec.
itself, rotted when it was planted into full nov. Kakt. & Sukk. 25(4):76–78.
ground. Hence, propagation by seed raising is
recommended. Minimum temperature in CHARLES, G. (2009). Micranthocereus, a view
winter is 10°C. If not grafted, the plant must from the field of a popular genus of
grow in a mainly mineral substrate. In a sunny columnar cacti. Cactus & Succulent Journal
place in the greenhouse one gets an intense (US) 81(3):150–155.
blue cereus with reddish spination. Very TAYLOR, N. & ZAPPI, D. (2004). Cacti of Eastern
attractive are high grafts which stimulate a fast Brazil.
development of the cephalium. When
cultivated plants loose their roots it is quite a
task to root these again. They grow slowly on
their own roots. It is recommended not to
disturb the root ball when repotting.
Remarks
On the type locality this cereus is in danger
and almost exterminated. The discovery of a
new 120km long habitat keeps Siccobaccatus
dolichospermaticus out of the danger zone. Local
people regularly cut down the columns to
burn them after drying.
Conclusion
A beautiful attractive columnar cactus that
can only rarely be admired in its full glory in
our collections.
Kamiel Neirinck
12
Copiapoa solaris does flower in cultivation!

At least it might if you live in California. Photographs by Duke Benadom
The species was first found in 1956 by

Friedrich Ritter and then described by him in
1961 as a new species and a new genus Pilo-
copiapoa. When Ritter wrote his 4 volume Kak-
teen in Südamerika, he relegated Pilocopiapoa to
subgenus status under Copiapoa (1980).
Ritter’s type locality is El Cobre, a mine at
There are a few cacti that achieve cult status the place where the coast road going north
and certainly Copiapoa solaris is one. It is the from Paposo turns inland. The distribution of
most northerly occurring large clumping the species extends north and south of this
species that makes clusters comprising dozens place, from Quebrada Botija in the south to
of heads. This part of Chile is extremely arid north of Antofagasta.
and has probably got even drier over recent A recent molecular study (Larridon et al.,
decades. Many of the plants in habitat are 2015) showed that C. solaris is basal to all the
dead, either through drought or perhaps from species of Copiapoa. It is certainly a distinctive
infections caused by their weakened state. species and one of the easier copiapoas to
Only in favourable times do the plants pro- recognise.
duce a few flowers, hence seeds are hard to
find. References
There are old plants in cultivation but they CHARLES, G. (1998). Copiapoa. The Cactus File
are difficult to flower and I am unaware of a Handbook 4. Cirio Publishing.
reliable report of a single plant flowering in the LARRIDON, I. ET AL.(2015).An integrative ap-
UK. I said the same thing in my Copiapoa book proach to understanding the evolutionand
(1998) and, in response, Ralph Martin pub- diversity of Copiapoa (Cactaceae), a threat-
lished a picture of his flowering plant (Martin, enedendemic Chilean genus from the Ata-
1998), except that I don’t believe the plant pic- cama Desert. American Journal of Botany
tured is really C. solaris, but rather a form of C. 102(9):1506–1520.
coquimbana .... sorry Martin! MARTIN, R. (1998). Copiapoa solaris in flower.
Duke Benadom recently sent me pictures of British Cactus & Succulent Journal 16(4): 214.
his C. solaris plants flowering in California. RITTER, F. (1961). Ein neues Kakteen-Genus aus
They clearly show the characteristic spination Chile. Kakt. and. Sukk. 12(2):20.
of this slow-growing beauty. RITTER, F. (1980). Kakteen in Südamerika, Band 3.
pp.1046–1053. GC
13
JOURNAL ROUNDUP
Kaktusblüte
I am always impressed by the enthusiasm of our
German friends for their cactus hobby. They have
produced many local regional publications over the
years but the one I really enjoy receiving is Kaktusblüte.
It is published by the Ortsgruppe Rhein-Main-Taunus
and the Kakteenfreunden Mainz/Wiesbaden.
It started back in 1985 with a simply produced
newsletter but soon progressed to a well printed black
and white publication. High quality colour pictures
were introduced in 1988 and the articles were of a
quality usually associated with a national journal.
The journal has been produced annually ever since.
The issue for 2016, shown right, includes well-
illustrated articles about travelling in Mexico;
Echinocereus from the Trans-Pecos, Texas; Epiphyllum
crenatum; Echinocereus from the north Sierra Madre
Occidental; the Mossen Costa i Llobrera gardens in
Barcelona; the genus Nolina; Melocactus warasii and
Lithops optica rubra.
The cover pictures is Ariocarpus ‘confusus’.
The publication (and back issues) can be bought for
about 6€ each from the book dealer Gottfried Gutte
http://www.cactusbooks.com
14
ON-LINE JOURNALS
On-line Journals for you to download free

Publishing journals on the web is now very popular. Creating them is a lot of work so perhaps
that is why some have ceased publication. Here are some links for you to download and enjoy.
Xerophilia
The eighteenth issue of Xerophilia appeared in October
2016. It is published in English as well as the language of
the original article. The quality of the contents is varied
and impressive.
Contents include: Sacred sites of the Wixaritari
Community; A new old plant: Turbinicarpus nikolae; Cacti
hybrids of Orogrande; Rebutia sensu Buining & Donald;
Notes on Mammillaria boelderiana; Spring in Mexico - part
1; Growing Lithops in Mexico; Ant Palnts of Papua New
Guinea; Vertical Garden Aeoniums;
The magazine may be downloaded free as a pdf from
http://xerophilia.ro
Contact: [email protected]
ECHINOCEREUS Online-Journal
The German language on-line journal for Echinocereus
lovers. The goals of this journal are to study the genus
Echinocereus, to publish articles about the continuous
research on these plants (classification, morphology,
evolution) as well as to protect the genus Echinocereus by
reproduction from seeds and distribution of the seedlings.
In this issue, 2016 04, there are well-illustrated articles:
A long overdue correction: Echinocereus salm-dyckianus
Scheer; Critically: variability and classification of the
Echococereus x lloydii hybrids
The downloaded pdf file allows printing, but does not
permit copying of the content. For those of us who do not
understand German very well, the publishers also
provide a downloadable MS Word document of the text
making it possible to copy and paste it into a translation
program. This is a major benefit of online journals and I
thank them for this useful feature.
See website: www.echinocereus.eu
15
Sansevieria Online
The online journal for the growing number of
enthusiasts for this genus. A small group of
Sansevieria enthusiasts publish the first
Sansevieria online journal in German. They
welcome contributions on systematics,
morphology, physiology, evolution etc.
This issue includes: The inflorescence of
Sansevierias; a new cultivar: Sansevieria
cylindrica ‘Boncel’; Shield bugs (Pinnaspis
strachani) as companions of Sansevieria; Not
only sansevierias in the Botanical Garden of
the University of Potsdam: a plant paradise in
Welterbe "Park Sanssouci"; Flower pictures of
Sansevieria burmanica.
The publisher of this online journal have set
themselves the goal of contributing more to
clarify this wonderful genus.
Download the PDF from www.sansevieria-
online.de where you can also find a special
issue containing field number lists and an
index to the journal.
Schütziana
The latest issue of Schütziana, the specialist on-
line journal for Gymnocalycium enthusiasts,
features:
Gymnocalycium friedrichii (Werderm.) Pažout ex
Schütz – Evaluation from a different perspective
Gymnocalycium bruchii (Spegazzini) Hosseus
subsp. deminii Gapon et Neuhuber
The text of this valuable publication is available
in English, German, Russian and Japanese.
The pictures and distribution maps give a clear
insight into the plants found in habitat and culture.
You can download free all the issues from:
www.schuetziana.org
16
Sukkulenten (formerly Avonia News)

Free German language on-line newsletter of
"Avonia", the quarterly journal of the German
Society for other Succulents.
From 2015, the on-line journal has been called
“Sukkulenten”
This issue has a wonderfully illustrated article
about Pseudolithos by the late Darrel Plowes.
See website: www.fgas-sukkulenten.de
Annual seed list for members and much more.
Special interest groups for Aloe (incl. Haworthia
etc.), Ascleps, Euphorbia, Mesembs and Yucca/winter-
hardy Succulents.
For membership and further information contact:
Dr. Jörg Ettelt: Morgenstr. 72, D-59423 Unna,
[email protected] or
Wilfried Burwitz: Postfach 100206, D-03002
Cottbus, [email protected]
Succulentopi@
The 15th issue of this free online journal

appeared a year ago. I wonder why no further
issues have appeared. It would be a great pity if
this valuable publication ceased to be
published.
It was the first online journal published in
French. The quality is excellent in every respect.
Back issues are available as a free PDF
downloads from:
http://www.cactuspro.com/succulentopia
17
THE LOVE OF BOOKS

Making Sense of German Journal Chronology

Germany was the most active country for cactus study at the end of the 19th century and
during much of the 20th. Graham Charles lists the major journals which were published there
under various names during this golden age. Pictures by Graham Charles
The many titles of cactus journals can be

confusing so I hope the following list will be a
useful reference. It is based on information
provided to me by Jörg Köpper who used to be
a book dealer from Wuppertal, Germany. I am
also grateful to Gordon Rowley for additional
information.
1923: The title of the DKG publication changed

to Zeitschrift für Sukkulentenkunde (ZfS). Each
volume of this new publication ran for two
years and comprised a total of 16 issues. Three
volumes (Vol.1: 1923/24, Vol.2: 1925/26 and
Vol.3: 1927/28) were published.
Vols. 1–3, 1923/24–1926/27
1891: the first German cactus magazine 1929: The title changed again, this time to
Monatsschrift für Kakteenkunde (MfK) started Monatsshrift der DKG, again with monthly is-
with No.0 and ran until No.12. It was founded sues running from January to December. Only
on the initiative of a private cactus lover and four volumes were published under this name
then carried on as a publication of the until 1932.
Deutsche Kakteen-Gesellschaft after 1892. This Vols. 1–4, 1929–1932
rare volume was reprinted by the DKG
(German Cactus Society) in 1992. 1932: A free saleable
Vol.1 1891/2 cactus journal called Der
Kakteenfreund was pub-
1892: The Deutsche Kakteen-Gesellschaft, DKG
lished for 4 years. (12 is-
was founded and the Monatsschrift für
sues monthly until 1934,
Kakteekunde became its official publication until
then just 6 issues bi-
1922. It was founded by Karl Schumann who
monthly in 1935)
also edited it from 1892 until his death in 1904.
Vols. 1–4, 1932–1935
Each volume of the publication comprised 12
issues published monthly and ran from
January to December. GC has some of these
old German journals
Vols. 2–32, 1892–1922
for sale, please see
Index to vols. 1–XX. 1912 the advert on p. 61.
18
1933: The title of the 1942 with 2 issues each year (except 1940 with
DKG publication only one issue) and was terminated with a
changed to Kakteenkunde double issue for 1943/44.
(monthly, 12 issues per 1937, Einleitender
year) and ran until 1936 Sonderteil, parts 1–2
as the official organ of
the DKG then, until 1938, parts 1–2
1938, as a free sale 1939, parts 1–2
journal for cactus lovers.
1940, part 1
In 1939 it again became
the official organ of the 1941, parts 1–2
DKG with at first 4 1942, parts 1–2
issues and then from 1940 to 1943 just 3 issues
each year. 1943/44, part 1
Vols. 1–4, 1933–1936 (DKG)

Vols. 5–6, 1937–1938 (free sale journal) 1938: A second DKG
publication was started
Vols. 7–11, 1939–1943 (DKG) named Beiträge zur
Sukkulentenkunde und
Pflege. This ran from
1938 to 1942 with 3
issues each year and it
finished in 1943 with
one last issue.
1938–1942, each with
parts 1–3
1943, 1 issue
1949: During World
War II all DKG
publications stopped
1936: The DKG published its first yearbook and only when the
called Jahrbuch der DKG 1935/36 (144 pages). Deutsche Kakteen-
Gesellschaft was
revived in West
1937: The official DKG publication was named Germany in 1949 was a
Kakteen und andere Sukkulenten for the first new journal, again
time. It ran for just two years, with twelve named Kakteen und
issues in the first and only 3 in the second. andere Sukkulenten
Vol.1, 1937 (1–12) (KuaS), started. It was
published irregularly
Vol.2, 1938 (1–3)
but since 1957 it has had 12 issues every year
until today.
1937: Also in 1937 the DKG started a new Vol.1, 1949/50 (5 issues)
publication called CACTACEAE – Jahrbücher
Vols. 2-3, 1951–52 (each with 4 issues)
der Deutschen Kakteen-Gesellschaft. The first part
of this publication was an issue named Vols. 4-6, 1953–55 (each with 3 issues)
Einleitender Sonderteil 1937 followed by part 1 Vol. 7, 1956 (6 issues)
and 2 of 1937. Then the publication ran until
Vol. 8, 1957 to today (each with 12 issues)
19
1949: Another DKG
publication called
Small Opuntias
Nachrichtenblatt der A new book from John Pilbeam
Deutschen Kakteen- and Mike Partridge
Gesellschaft was
started and appeared
irregularly until 1956
Vol.1, 1949/50 (10
issues)
Vol.2, 1951 (7 issues)
Vol.3, 1952 (9 issues
with Nos. 1–12)
Vol.4, 1953 (7 issues with Nos. 1–8)
Vols.5–7, 1954–56 (each with 6 issues)
1965: A good quality

magazine titled
Stachelpost with b/w
and colour pictures
was founded with
No.1 (1965) and ran There is no doubt that small opuntias are
until No.51 in 1974. currently very popular. They are easy to
Vols. 1–10, 1965–1974 grow, most are cold tolerant, and many will
(Nos.1–51) produce their beautiful flowers while in small
pots. And, if you see one you like, the chances
are that you will be able to get a cutting!
Reference This well-illustrated book provides a
EGGLI, U. (1995). Biography of Succulent Plant useful guide to the genera and species which
periodicals. Fričiana 60. may be described as small and also a few
which are not so small. Also included is
Die Echinocereus scheeri Gruppe Maihuenia which is probably here because it
Another well-illustrated will never find a place anywhere else!
book from the German The pictures, which were supplied by
Arbeitsgruppe Echinocereus many enthusiasts, are the joy of this book and
providing an in-depth they comprehensively illustrate the available
study of ten taxa in section species. You can also find advice on
Scheeria of subgenus cultivation, all in all making it easy to
Triglochidiata. recommend this book to the keen cactophile.
This well-produced hard- 150 pages 275 x 210mm, 250 photos of
bound book has 176 pages species flowering in habitat and cultivation,
233 x 170mm, and 219 colour pictures, maps, with maps and description references.
herbarium specimens and seed SEMs. £38 UK, £43 EU, or £48 everywhere else
Available to non-members for 24.50€ + car- including postage.
riage, 2€ less for members. German text. Text Payment by UK cheque to John Pilbeam,
file available for use in translation programs. 51, Chelsfield Lane, Orpington, Kent, BR5
Available from Ulrich Dosedal 4HG, England or pay by PayPal to
[email protected] [email protected]. GC
20
A NATURALISED POPULATION OF
ESPOSTOA MELANOSTELE
(CACTACEAE: CACTOIDEAE)
ON TENERIFE (CANARY ISLANDS)
Filip Verloove, Botanic Garden of Meise, describes his discovery of a population of Espostoa
melanostele on Tenerife. The island is well known for the suitability of its climate for the
cultivation of cacti, and these plants look remarkably similar to those growing in their natural
habitat. Photographs by the author.
The genus Espostoa Britton & Rose is entirely Pseudoespostoa Backeb., but all these are now
South American and includes from about nine usually accepted as a single genus (Mauseth,
up to 16 species, depending on the species 1999). Molecular studies, however, will
delimitation (e.g. Mabberley, 2008; Sánchez de certainly shed new light on its boundaries.
Lorenzo Cáceres, 2000). Its center of diversity Schlumpberger & Renner (2012) already
obviously lies in Peru with two species found demonstrated that, in its current (i.e., broad)
in Ecuador. At one time, species currently circumscription, Espostoa is paraphyletic and
accepted in Espostoa were accommodated in a that Vatricania Backeb., another segregate, is
few segregated genera, among which was not closely related.
21
22
Species of Espostoa are highly appreciated about this record are provided and the plants
for their decorative qualities due to their are copiously illustrated.
splendid white fleece. No wonder that several Espostoa melanostele is a species endemic to
species are grown as ornamentals. In Europe, central Peru where it is found both in the
at least four species are more or less widely coastal and the warmer inland valleys from
available: E. blossfeldiorum (Werderm.) Buxb., E. about 800m up to over 2,000m (Charles, 1999).
lanata (Kunth) Britton & Rose, E. melanostele It is columnar and branched, usually from the
(Vaupel) Borg and E. senilis (F. Ritter) N.P. base, with several erect stems to 10cm across.
Taylor (Hunt, 1998; Sánchez de Lorenzo Most are densely covered in whitish, fuzzy
Cáceres, 2000). However, in continental hairs that cloth the entire stem (this is
Europe none of the species is suited for particularly attractive in young stems). In E.
cultivation out of doors: they are classified as melanostele, the main central yellowish spines,
“G2 – needs a heated glasshouse even in south the longest up to 10cm long, extend well
Europe” (Hunt, l.c.). beyond the hairs. The whitish to brownish
In June 2016, a magnificent colony of a (pseudo-) cephalium is lateral, usually on one
species of Espostoa was discovered on a bare, side of the stem and bears numerous
sun-exposed slope near Playa de San Juan in (nocturnal) flowers with a whitish perianth
Tenerife (Canary Islands, Spain). This species that is often slightly suffused with pink. The
was identified as E. melanostele (syn.: Pseudo- plants found in this Canarian locality are,
espostoa melanostele (Vaupel) Backeb.), a species despite being mature, relatively small (all well
that – to our knowledge – had not been below 100cm) and have a pale-coloured
recorded before growing wild outside of its cephalium rather than the dark brown
area of origin. In this short note some details cephalium of E. melanostele s.str. These are
23
traits typical of subsp. nana (F. Ritter) G.J. Acknowledgements
Charles, a taxon that was originally given I sincerely wish to thank Graham Charles
species rank (E. nana F. Ritter). Young plants in (Stamford, England) for confirming the
cultivation, however, are indistinguishable identity of the Canarian plant.
from E. melanostele (Charles, 1999). Therefore, it
is now accepted at most at subspecies rank or Filip Verloove, Botanic Garden of Meise,
merely sunken in the synonymy of the latter Nieuwelaan 38, B-1860 Meise, Belgium.
(e.g. Hunt, 1998). fi[email protected]
On a bare and sun-exposed stony slope on References
the verge of a shallow barranco, close to the CHARLES, G. (1999) The genus Espostoa Br. & R.
sea cliffs near Playa de San Juan, Espostoa Brit. Cact. Succ. J. 17(2): 68–79.
melanostele occurs with several tens of
HUNT, D.R. (1998) Espostoa. In: WALTERS, S.M. ET
individuals (probably close to one hundred
AL. (EDS.), The European Garden Flora vol. 3.
altogether). The plants in this population are
Cambridge University Press, Cambridge:
of variable age and include young individuals
232–233.
as well as sexually mature ones with flower-
bearing cephalia. Espostoa melanostele has a MABBERLEY, D.J. (2008) Mabberley’s plant-book
reputation for being slow-growing. Even well (3rd ed.). Cambridge University Press,
grown individuals are not more than 25cm tall Cambridge: XVIII + 1021p.
after ten years in cultivation (Charles, 1999); MAUSETH, J.D. (1999) Comparative anatomy of
hence, the presence of numerous mature plants Espostoa, Pseudoespostoa, Thrixanthocereus
in this locality suggests a relatively old and Vatricania (Cactaceae). Bradleya 17: 27–
introduction and the species looks perfectly 37.
established there.
SáNCHEZ DE LORENZO CáCERES, J.M. (2000) Flora
As for all cacti currently found in the wild in Ornamental Española, vol. II Cactaceae-
the Canary Islands, Espostoa melanostele Cucurbitaceae. Mundi-Prensa.
evidently must have been introduced on
purpose initially, most likely as a garden SCHLUMPBERGER, B.O. & RENNER, S.S. (2012)
ornamental (although it is sometimes grown Molecular phylogenetics of Echinopsis
for its edible, sweet and juicy fruits as well). In (Cactaceae): Polyphyly at all levels and
Playa de San Juan it probably naturalised from convergent evolution of pollination modes
garden debris or may even originally have and growth forms. Amer. J. Bot. 99(8): 1335–
been planted intentionally. Nearby, on the 1349.
other side of the barranco, an extraordinary VERLOOVE, F., SMITH, G.F. & BARONE, R. (in
collection of introduced cacti and other preparation) New alien succulents from the
succulents is present and this is obviously an Canary Islands (Gran Canaria and Tenerife;
illegal, abandoned former plantation. It Spain).
includes several exceptional alien species that
ultimately managed to naturalise locally, e.g.
Cylindropuntia fulgida (Engelm.) F.M. Knuth
var. mamillata (Schott ex Engelm.) Backeb.,
Euphorbia grandialata R.A. Dyer, Opuntia
macrocentra Engelm., Tephrocactus articulatus
(Pfeiff.) Backeb., etc. (see Verloove et al., in
preparation), but not Espostoa melanostele.
24
TRAVEL WITH THE CACTUS EXPERT (16)

Zlatko Janeba continues his popular series of articles about exploring the American South
West. Photographs by the author.
As already mentioned, Josef Busek was involved The garden was quite busy as there was a fair
in some business negotiations for the whole with rock plants going on during those days
morning on 12th May 2006. Thus, I was left on my (Fig.2). But imagine my disappointment when I
own. I was recommended to visit the Denver figured out the tropical greenhouses were closed
Botanic Gardens and I decided to use the for reconstruction. I was only allowed to visit the
opportunity while being in Denver. First I went to outdoor exhibitions, but still, there was plenty to
wash my dirty and dusty car and to do some discover. I could see numerous frost and winter
shopping to replenish our food stock for the next hardy plants growing in the rock gardens,
couple of days. Then I headed to the gardens. especially various agaves, yuccas, North American
cacti of genera Opuntia and Echinocereus, hardy
Denver Botanic Gardens (Fig.1) is a famous
Patagonian Maihuenia poeppigii (Fig.3) as well as
public botanic garden with an area of some 23
flowering South African succulents, like Delosperma
acres. It was founded in 1951 and currently
nubigenum (Fig.4).
features a variety of theme gardens. The garden is
well-known especially for its large collection of Moreover, I could enjoy an outdoor exhibition
plants from cold temperate climates around the of numerous sculptures called “Living Bronze”,
world, which also includes my favourite frost- open from 20th April to 31st October 2006. Twenty-
hardy cacti and other succulents. six bronze sculptures by artist Robert Wick were
Fig.1 The entrance to the Denver Botanic Gardens. Fig.2 Many people were interested in the sale of a
great selection of rock plants in the Denver Botanic
Gardens.
Fig.4 Flowering Delosperma nubigenum or “Hardy

Yellow Ice Plant” in the outdoor exposition in the Fig.3 Even Maihuenia poeppigii from Patagonia is
Denver Botanic Gardens. doing well in rock garden in Denver, Colorado.
25
Fig.6 Escobaria missouriensis was just about to

flower. Southwest of Denver, Colorado (May 12,
2006).
mostly prairies and woodlands. They were recently
also discovered in the Mexican state of Coahuila.
And although the distribution range of E. missouri-
ensis is wide, it is not always so easy to find as this
is a relatively small-bodied and cryptic species.
They can be easily spotted only when in flower or
when bearing shining orange-red spherical and
juicy fruits. Often they can be seen decorated both
with flowers and with fruits at the same time. In
cultivation they belong among the most tolerant
cacti and are fully winter hardy. Although
inconspicuous for their relatively unattractive
Fig.5 Shiva (1990, Bronze 8’ 6’’). Shiva is the Hindu spination, they attract a lot of attention among
god of creation and destruction. Multiple arms are laymen when in flower and fruit. Interestingly,
American botanists do not recognize the genus
broken. A huge earth and sea formation behind her
Escobaria and keep it as Coryphantha missouriensis. I,
head carries much life as evidenced by the living on the other hand, prefer to call these interesting
plants. cacti Neobesseya missouriensis.
placed in the landscape of the American West and We drove further in a southwesterly direction
various living plants were actually inserted into along US Route 285 and after some time we
numerous cavities of the sculptures (Fig.5). So after stopped in a pine forest at an elevation over 2500m
all, I enjoyed my visit to the Denver Botanic (Fig.7), where we saw numerous small yuccas with
Gardens anyway and I took copious photos. dry open seed capsules (could be Y. angustifolia). I
Later I was told by Josef that we had been have also found a single specimen of Pediocactus
invited to a chalet of another colleague of his near simpsonii with delicate pink flowers (Fig.8).
Fairplay, that is situated in the Rocky Mountains Only about 4 miles further the pine forest
not too far from Denver. We took off after lunch thinned out and we stopped to search through
and headed along US Route 285 towards the montane meadows at an elevation of some 2500m.
southwest. We stopped just southwest of Denver, We encountered the same yucca species we had
near the exit to the Red Rocks, and there we seen at the previous spot, and also several clusters
discovered several Escobaria missouriensis speci- of Escobaria vivipara without any signs of flowers
mens with flower buds (Fig. 6). They grew in red (Fig.9). But the most common cactus on the open
sandy soil among grasses together with a small grasslands was Pediocactus simpsonii. We saw
yucca species. dozens of pediocacti but the flowering season there
I am always more than happy to see Escobaria was already over. The largest pediocacti there were
missouriensis or related plants in the field. This only about 6 cm in diameter.
group of cacti is quite widespread in the We made our next stopover just about 1 mile
southwestern and cental states of the U.S.A. before the Kenosha Pass (north of it) at an elevation
(Arizona, Colorado, Kansas, Montana, Nebraska, of some 2900m. This is about 5 miles northeast of
New Mexico, North Dakota, Oklahoma, South Jefferson in the Rocky Mountains (Colorado). This
Dakota, Texas, Utah, and Wyoming). They inhabit area is often recomended to search for pediocacti.
26
Fig.7 A habitat along the US 285 at elevation of some 2500 m. We found numerous yuccas and flowering
Pediocactus simpsonii in the open pine forest.
Fig.8 Pediocactus simpsonii with delicate pinkish flowers along the US 285 (2500 m), Colorado.
27
Fig.9 Large clump of Escobaria vivipara along the Fig.10 Population of copious small Pediocactus
US 285 (2500m elevation), Soutwest of Denver, simpsonii along the US 285 (2890m elevation), just
Colorado. North of Kenosha Pass, near Jefferson in Col-
orado.
population of P. simpsonii at such a high altitude.
Unfortunately, it was already getting late and cold,
so the flowers were mostly closed at that time.
Pediocactus simpsonii, together with the above
mentioned Neobesseya missouriensis and also
Escobaria (or Coryphantha in the USA) vivipara,
belongs to globular cacti with the widest
distribution range in the southwest of the USA. P.
simpsonii (in a broad sense) has been reported from
Arizona, Colorado, Idaho, Montana, Nevada, New
Mexico, Oregon, South Dakota, Utah, Washington,
and Wyoming. It is mostly an inhabitant of
Fig.11 Pediocactus simpsonii with pinkish flowers woodlands and montane grasslands at altitudes
closing for the night, along the US 285 (2890m ele- usually between 1500m to 3500m. The large
vation), just North of Kenosha Pass, near Jefferson distribution range is reflected in high variability of
in Colorado. these plants which actually led to descriptions of
several subspecific taxa (variety and subspecies).
We climbed up a grass meadow next to the road
These are not, however, accepted as good taxa
but we could not find any cacti there. The grass
anymore.
seemed to be too dense to allow any cacti to grow
there. After a while we reached a rocky place Then we headed further and just before sunset
(composed of granite rocks and gravel) within the we arrived at the chalet of Josef’s colleague that is
grassland and there, finally, we discovered a rich located near Fairplay at elevation over 3000m.
population of Pediocactus simpsonii growing in Nobody was there yet, but Josef had been given
slightly sandy to gravelly substrate among grasses. keys, so we could comfortably settle inside the
The plants were really almost everywhere, most of house. The owner’s family arrived later that
them bearing nice pink flowers (Figs.10 & 11). We evening and we spent a very pleasant evening
also saw miniature species of Sedum with reddish- together sitting around a warm fireplace.
brown tiny leaves. It was indeed a wondeful Zlatko Janeba
experience to see such a plentiful and healthy
28
WHERE LIZARDS DARE:
AN EXCURSION TO BARRANCO RAMBLA

DE RUIZ (TENERIFE)
Marco Cristini describes an interesting day looking at succulents on the popular holiday
island of Tenerife. Photographs by the author.
The Canary Islands are one of the most richer areas are surely those of Anaga (in the
spectacular places on Earth for succulento- East) and Teno (in the West), but also in the
philes. This remote corner of Europe is home to central part of the northern coast there are quite
a rich endemic flora, whose species are a few interesting places. In this article I will
cultivated in gardens and greenhouses shortly describe one of them, Barranco Rambla
throughout the world. But one thing is to grow de Ruiz.
aeoniums or monanthes in Northern Italy, in This narrow valley is situated between
England or in the USA, another is to see them Puerto de la Cruz and San Juán de la Rambla.
in their habitat. Reaching it is very easy: the barranco is
In August 2016 I went for the first time to perpendicular to highway TF-5, one of the most
Tenerife, in order to see face to face as many important roads of Northern Tenerife. It is
Canarian succulents as possible. I was not impossible to miss it, because shortly before
disappointed. Although I arrived at the peak of there is an unambiguous brown road sign.
the dry season, during my holiday I spotted at After a sheer embankment one has to turn left
least 17 different species of Crassulaceae. The (coming from Puerto de la Cruz) and to park
Fig. 1. Barranco Rambla de Ruiz
29
Fig.2 A stressed Aeonium canariense var. Fig.3 Aeonium arboreum var. holochrysum.
canariense.
Fig.4.Aeonium arboreum var. holochrysum. Fig.5. Aeonium arboreum var. holochrysum.
Fig.6. Aeonium arboreum var. holochrysum in a Fig.7. Aeonium arboreum var. holochrysum on the
shaded place. path.
30
Fig. 8. Barranco Rambla de Ruiz.

the car in a nice picnic area, where whole due, in my opinion, to lack of water.
families of El Rosario, the tiny town which is in When I arrived in Tenerife the flowering
front of the barranco, spend their Sunday season had already ended for the majority of
afternoons. An elegant stairway situated near Crassulaceae. I spotted lots of inflorescences of
the entrance leads to the beginning of the path. Aeonium urbicum and a few of Aeonium
When I was still climbing on the steps I saw haworthii, but I rarely saw a dried inflorescence
the first succulent, Aeonium arboreum var. of A. canariense var. canariense. I don't know the
holochrysum, which had only tiny and compact reasons of this circumstance, maybe the spent
rosettes because of the drought. Many leaves inflorescences of this species break off more
were bordered or veined with red, so some easily or perhaps few plants flowered in 2016.
plants had a somewhat marbled appearance. But let us return to Barranco Rambla de Ruiz.
Then, after few minutes, I saw the real “king” After the first A. canariense var. canariense the
of the barranco, Aeonium canariense var. path becomes steeper and if it is hot (as, for
canariense. This succulent is quite common in example, during my visit) one or two bottles of
Tenerife, I saw it for example near Puerto de la water are necessary.
Cruz (on Sendero del Agua), in Barranco de The landscape, however, is unforgettable and
Igueste and in Chinamada, where I spotted a aeoniums are ubiquitous. Every five steps you
solitary plant near the village. In Barranco can see a plant or, more often, a nice group of
Rambla de Ruiz, however, it is very abundant plants, hanging on rocks, growing under trees
and it is found along the path, allowing the or thriving along the path. Also Aeonium
tourist to take lots of photos. I observed that arboreum var. holochrysum is common in the
this aeonium mostly grows under bushes, trees barranco, but it was less conspicuous than A.
or cliffs, in sheltered positions, where the canariense var. canariense and also less
tropical sun cannot burn it. In spite of this quite photogenic. The valley's cliffs are covered also
a few specimens had red leaves, a sign of stress by Opuntia ficus-indica, an invasive plant native
31
Fig.10. Aeonium canariense var. canariense.
Fig.9. Aeonium arboreum var. holochrysum.
Fig.11. Aeonium canariense var. canariense. Fig.12. Aeonium canariense var. canariense.
Fig.13. Aeonium canariense var. canariense. Fig.14. Aeonium canariense var. canariense.
32
Fig.15 Aeonium canariense var. canariense. Fig.16 Aeonium canariense var. canariense.
Fig.17 Aeonium canariense var. canariense. Fig.18 Aichryson laxum.

of Mexico which, since the discovery of cultivated fields. Also Aichryson laxum is quite
America, has been spreading in Europe, Africa common. I observed it, often together with M.
and Asia. I saw it in almost every corner of laxiflora, in Anaga, in the Teno area and in
Tenerife, often competing with Euphorbia Barranco Badajoz (near Güimar).
canariensis. The former succulent, however, is a After having dutifully photographed
good omen, at least for me. In fact I noticed that Monanthes laxiflora I went on walking, but not
wherever I spotted Opuntia ficus-indica also a for long. The path in fact reaches quickly a little
few Crassulaceae grow. square and this is the end of the journey or, at
After an hour of climbing, the city-dweller least, it was the end of my journey, because one
and not-very-athletic tourist is authorized to be can continue the excursion on an asphalted
a little tired, but fatigue is quickly forgotten road leading towards La Vera and Los Barros,
once you look at the valley and, on the horizon, but that day I had seen enough plants. So I sat
at the Atlantic Ocean. Moreover, new botanical on a bench and admired a Dracena draco planted
wonders are at hand! Near the end of the path I in the middle of the square, while groups of
found Monanthes laxiflora and a very dried white clouds crossed the barranco eager to
specimen of Aichryson laxum. The former reach the Ocean. When I rested thinking of all
species forms little mounds of tightly packed succulents I had just seen, I spotted one of the
stems and it grows well on rocky outcrops and most curious inhabitants of Tenerife, Gallot's
stone-walls. I saw it also in Barranco de Igueste, lizard (Gallotia galloti).
in the Anaga region, in Barranco de Bucarón This reptile is quite a bit bigger than the
(near Los Silos) and in Teno Alto, where the average Italian lizard, so when it crossed my
succulent thrives on old walls bordering
33
Fig.19. Monanthes laxiflora.
Fig.20. Dracena draco in the middle of the square

at the path's end.
Fig.21. Gallotia galloti in the Barranco Rambla de

Ruiz.
Fig.22. Gallotia galloti on the Sendero del Agua.
Fig.23. Gallotia galloti on Sendero del Agua near

the Ocean.
34
Fig.24. Map.
path for the first time, at the beginning of the
holiday, I was a bit startled. By then, however, I
had grown accustomed to them, so I cautiously
came closer and I photographed the little
dinosaur. Canarian lizards do not fear people Fig.25.Opuntia ficus-indica in the Barranco Rambla
too much. Naturally if you go towards them de Ruiz.
quickly they run away, but if you do not make
canariense var. canariense and the astute Gallotia
a sudden gesture they ignore you. Moreover, I
galloti, respectively king and queen of this arid
observed that they like crackers and, above all,
but unforgettable realm.
bananas. If you eat a frugal lunch in a dry
barranco, you can find yourself literally Bibliography
surrounded by hungry lizards, ready to fight E. CARBONELL (2007). Cuadernos de succulencia.
for a crumb. Once an inquisitive reptile even Ciappe, Barcelona.
entered my rucksack! During the way back I U. EGGLI (ED.) (2003). Illustrated Handbook of
spotted and heard many lizards along the path. Succulent Plants: Crassulaceae. Springer
When they move on a cliff, in fact, they cause Verlag, Berlin.
miniature landslides. I wondered how much A. LEROY (2013). Let us visit the Macaronesian
they are responsible for the island's erosion... Islands: Tenerife. The Cactus Explorer 7 :47–
The descent of the barranco is easier, but not 53.
too much quicker than the ascent. The path can H.Y. LIU (1989). Systematics of Aeonium
be tricky and one has to pay attention if you do (Crassulaceae), Special Publications Number 3.
not want to admire an Opuntia ficus-indica up National Museum of Natural Science,
too close. Moreover, the descent offers the Taichung (Taiwan).
possibility of seeing (and photographing) again J. LODé (2010). Plantas Suculentas de las Islas
the wonderful succulents which grow along the Canarias. Turquesa, Santa Cruz de Tenerife.
barranco. After more or less an hour I returned R. SCHULZ (2007). Aeonium in habitat and
to the picnic areas, tired but happy. cultivation. Schulz Publishing, San Bruno
(California).
Surely Barranco Rambla de Ruiz is not the
most succulent-rich valley of Tenerife, but it is
Marco Cristini
worth a visit. The tourist can enjoy a typical
Canarian habitat and know better some of the
island's inhabitants, like the superb Aeonium
35
JUAB AND UTAH COUNTY, UTAH,

THROUGHOUT THE YEAR 2015
It must be a joy living close to where cacti grow naturally. One such place is Salt
Lake City, Utah. Article by Petr Šimon. Photographs by the author.
My wife and I thoroughly enjoyed the the cemetery and stopped after a couple
year we lived in Salt Lake City, Utah. of miles to explore a small hill. The field
During this time we travelled frequently was covered with sparse vegetation
and spent almost every weekend which predominantly consisted of
enjoying the natural beauty of Utah and Sagebrush (Artemisia sp.). However, we
the neighboring states. During our very did not find any cactus plants.
first trips searching for cacti we We then drove to a second locality
explored localities near Eureka in both one mile east of Eureka on highway 6.
Juab and Utah Counties. We were able We parked our car on a shoulder close
to visit this region three times in to some rocky hills. After a few steps,
February, April, and July. we found our first Echinocereus
Our first visit happened on the last triglochidiatus v. mojavensis on a rock,
weekend of February (Feb. 28th, 2015). under a juniper tree (Juniperus sp.),
Based on my map studies, we started at (Fig.2). The plants were dry and limp
the area west of Eureka. We drove past but in good shape, undamaged. We
Figure 1. View from slope looking over the snowy landscape west of Eureka.
36
Figure 2: Rocky place under a juniper tree with

snow and Echinocereus triglochidiatus v. mojaven-
sis.
Figure 3: A lot of buds on Echinocereus triglochidia-

tus v. mojavensis at a locality north of Highway no.
6 near Eureka, Utah (April 11th, 2015).
decided to search the larger area, but as Figure 4: Mid-April and flowering time at the Pedio-
we travelled further from our car, we cactus simpsonii locality near Eureka, Utah.
saw fewer cactus plants. Soon the of Echinocereus triglochidiatus v.
weather became unfriendly including a mojavensis. These plants were well
snowstorm and freezing temperatures, hydrated and prepared for flowering.
so we decided to end our trip. The same day, we visited a second
Our next visit to this area was in April locality south of Highway no. 6.
(Apr. 11th, 2015). We decided to explore Following a short walk, we found a very
other localities east of Eureka. The first nice place at the top of a hill. The most
one was north of Highway no. 6. common plants there included sparse
Unfortunately, the whole area was growth of juniper (Juniperus sp.) and
destroyed by human activity. Off-road pine (Pinus sp.) trees with sagebrush
driving and geological research had (Artemisia sp.) and lupin (Lupinus sp.) in
damaged the land rapidly and flower, but only one cactus species grew
profoundly. Despite this deterioration, there. It was Pediocactus simpsonii with a
we found a couple of plants of interest diameter larger than 10cm and with
on a hill. We studied our first Pediocactus yellow flowers. The amount of plants
simpsonii. The second slope of the same was breathtaking, and the adult plants
hill was covered by a very dense growth were all in flower. We studied this
37
Figure 5: Detail of a yellow flowering plant of Pediocactus simpsonii with tapered petals at a locality near
Eureka, Utah.
locality in more detail including the
number of juvenile plants that grew
there.
Since all the adult plants were in
bloom, it was a perfect time to compare
the flowers. It is common that collectors
select plants according to their literature
description which contains only limited
information about the natural
variability that is known among them.
We explored the variation of the shape
and colour of the petals.
The first plant had tapered petals
with a yellowish interior and a pinkish
tint on outer edge (Fig.5). Another plant
had lighter petals than the previous one.
Also, their colour was more uniform
than the first plant. However, the most
significant difference was the shape of
the petals, which were rounded (Fig.6).
Figure 6: Detail of whitish flowering plant of Pedio- The last plant with flowers, I want to
cactus simpsonii with rounded petals, locality near
Eureka, Utah. show you, had a petal shape between
38
Figure 7: Detail of pinkish-flowering Pediocactus Figure 8: Opuntia sp. with fruits, west of Nephi,
simpsonii with slightly tapered petals, locality near Juab County, Utah.
Eureka, Utah.
Figure 9: Sparse junipers and Pediocactus simp- Figure 10: Summer view at a locality near Eureka,
sonii at a locality west of Nephi, Juab County, Utah. Juab County, Utah.
the two previous plants (Fig.7). It could and Utah Counties was on Sunday July
be described as almost rounded but 12th 2015. The first stop was at a locality
slightly tapered. Additionally, its colour west of Nephi, Juab County, Utah. We
was vibrant. It didn’t have any yellow, found information about this locality
instead transitioning from a light pink from a herbarium. After a short walk
in the center to a dark pink at the into nature, we successfully found an
flower’s exterior. opuntia with dry fruit (Fig.8).
Our last visit to the localities in Juab The most frequent plants at this
39
Figure 11: Echinocereus triglochidiatus v. mojaven-

sis in summer, east of Eureka, Utah County, Utah,
and removed plants in distance. Figure 12: How bulldozers take care of cacti.
locality were grasses, sagebrush the private property of a mining
(Artemisia sp.), and juniper trees company, who had removed all the
(Juniperus sp.). As we continued further trees and plants using bulldozers.
from the road, we found desirable cacti It is difficult to understand how this
near the top of a small hill. It was could occur in a world where CITES
Pediocactus simpsonii as described in rules exist to protect these species
herbarium files (Fig.9). (Fig.12).
We also visited another locality in As you read, the landscape and
Utah county near Eureka, Utah. In vegetation change dramatically through
comparison with the previous locality the year, so it is very nice to visit
west of Nephi, this area had little various localities in freezing winter,
vegetation and was full of stones (Fig flowering time in mid spring, and the
10). This arid microclimate increased the dry period in late summer. Each visit
dryness of the cacti we found here. was distinct and will be deep in our
On our way back to the Salt Lake hearts.
City, we stopped on road No.6 east of Petr Šimon
Eureka, Juab County, Utah. However,
we only found a limited number of
viable cacti, which grew only in rocky
areas (Fig.11). All the nearby fields were
40
A HAPPY MEDIUM?
A never ending discussion about Morangaya pensilis (K. Brandegee) Rowley!
Michael Lange discusses its relationships and provides information about its
distribution in Baja California Sur Photographs by the author unless otherwhis stated.
‘Pensilis’ is a cactus, with a hard to explain compared it with Heliocereus. And not to forget
phylogenetic background. It makes botanists my own misinterpretation (Lange in Blum et
breathless and leads us around by our noses al., 1998: 45) and the creation of Morangaya as a
like a dancing bear at a middle age market. subgenus within Echinocereus is obsolete now;
Some say one thing, others like a different the former genetically based statement by
solution. I personally think this species likes Wallace & Forquer (1995) is now revised by the
itself best in the position defined as a (un-) better knowledge published by Barcenas et al.
happy medium (Rowley, 1974). (2011). And thus is confirmed by Sanchez et al.
For a long time M. pensilis seemed to have
been the only Echinocereus growing in the
manner of an Aporocactus from the top hanging
down, we shall not wonder why this character
was over stressed in the genus definition for
Morangaya by Rowley (1974). He also
Figure 2 Morangana pensilis in cultivation.

Photograph by Peter Hallmann
Figure 1 Morangana pensilis in cultivation.

Photograph by Peter Hallmann
Figure 3 Morangana pensilis in cultivation. Figure 4 Morangana pensilis in cultivation.

Photograph by Peter Hallmann Photograph by Peter Hallmann
41
Figure 7a Stenocereus kerberi

Photograph by J. Lodé
Figure 5 Morangaya pensilis fruit in cultivation.
Figure 6 Morangaya pensilis in habitat at 600m, north of San Jose del Cabo. Photograph by P. Breslin
42
Figure 7c Stenocereus gummosus fading flower.
Figure 7b Stenocereus gummosus

Photograph by P. Hallmann
Figure 7d Stenocereus gummosus

Photograph by P. Hallmann Figure 8a Morangaya pensilis flower
(2014) again: Morangaya does not belong to a
monophyletic genus Echinocereus Engelmann!
Looking back, you will eventually admit
that the proposed origin as explained by Blum
et al. (1998: 39) is rather close to the updated
point of view. Nevertheless I am still expecting
the next change for the phylogenetic position
of the taxon ‘pensilis’. Because today’s position
proposed by analysis (Barcenas et al., 2011: 5)
as a sister taxon to Stenocereus eruca (indeed
very close to the former genus Machaerocereus)
is eventually telling the beginning of the next
part of the whole (never ending?) story. Well,
comparing the flower shape of these
Stenocereus species and not to forget the less
specialized flowers of the Rathbunia-group one
will discover some parallels. Also the flower
Figure 8b Morangaya pensilis flower section
43
Figure 10 Morangaya pensilis small buds
Figure 9 Stenocereus alamosensis small bud
Figure 11b Morangaya pensilis in culture

ecology (daytime and short-lasting anthesis)
tells us something, does it not? Even the
growing habit is quite similar but one should
not over-estimate such features when
identifying relatives; as we learned from the
Aporocactus example!
Comparing morphological and ecological
details
Growth
The people from the cactus community in
general are well infomed about the shape and
style of growth of the proposed sister-taxon
Figure 11a Morangaya pensilis in culture Stenocereus eruca (syn. Machaerocereus)
44
Figure 12 Morangaya pensilis fruit, ripe and juicy.

although we will never ever get the very
special spination in our greenhouses.
We will hardly see Stenocereus gummosus in
flower within European collections even if we
will find it at all. So we do not know much
about its natural variability.
Figure 13 Stenocereus alamosensis fruit in
Unfortunately we have a scanty knowledge habitat. Photograph by T. VanDevender
of the daytime-flowering octopus-cacti
(Stenocereus « cina »-group synonym Rathbunia,
see Lange (2016) in the Cactus Explorer 16
and
http://de.wikipedia.org/w/index.php?title=Dat
ei:Teguise_Guatiza_-_Jardin_-
_Stenocereus_alamosensis_03_ies.jpg&filetime
stamp=20111028180537)
But there is some light thanks to a German
picture-postcard from the 1980s and a well
illustrated publication by Niestradt (2000).
Another report by Earle (1972) briefly
mentions the space-demanding growth with a
single bending top of a specimen; a potential
start for a new plant of the entire clone.
All these species share the opportunity of
producing adventives roots in order to
establish entire individuals. This is an
interesting way of becoming immortal!
Flower
The so-called night flowering species
normally share the character of keeping their
flowers (depending on temperature) open until
the following late morning. This is probably
the reason for contradicting data about the
time of flowering and for the flower colour. Figure 14 Fruit of Stenocereus gummosus Baja
Hence those normally white flowers change to California in 1997
45
Figure 15a Morangaya pensilis seed. SEM from Figure 15b Morangaya pensilis seed. SEM from
population „La Soledad“ SEM by G. Mettenleiter population „La Soledad“ SEM by G. Mettenleiter
Figure 16a Morangaya pensilis seed. recieved as Figure 16b Morangaya pensilis seed. recieved as
Lau 050 SEM by G. Mettenleiter Lau 050 SEM by G. Mettenleiter
Figure 17a Morangaya pensilis seeds under light- Figure 17b Morangaya pensilis seeds under light-
microscope: ex W. La Haye s.n., ca. 2.000m microscope: ex W. La Haye s.n., ca. 2.000m
(see Corbett, 1998: 69 Abb. 194) (see Corbett, 1998: 69 Abb. 194)
46
Figure 18a Stenocereus alamosensis SB 1834 Figure 18b Stenocereus alamosensis SB 1834
seed SEM by G. Mettenleiter seed SEM by G. Mettenleiter
Figure 19a Stenocereus gummosus collected July Figure 19b Stenocereus gummosus collected July
1997 SEM by G. Mettenleiter 1997 SEM by G. Mettenleiter
light pink when ageing. Interestingly, we find from undocumented and from plants with
specimens with really red daytime flowers in field data. Right from the beginning of my
the Rathbunia-group and also one species with investigation, I was able to compare the
a white night-time flower (S. standleyi). original SEM for Lau050, kindly provided by
Morangaya itself occupies a middle way Dr. G.R.W. Frank (see Frank, 1986), with
between these: The flowers open in late undocumented (historical) material: I directly
afternoon, stay open all night and fade the next recognized that there are huge differences
day during early afternoon ; this is altogether among them. Over the years I ordered more
about 20 hours of anthesis. and more SEM-pictures and even today it is
not easy to describe a typical seed character for
Fruit the species. So it is best illustrated!
Its not that easy to get Morangaya to flower While seeds of most of the related
and it is even more problematic to get nice Stenocereus taxa are hard to come by, here in
fruits and seeds. I have to admit, I have never Europe they are best compared by the
ever investigated a Rathbunia fruit ! But here illustrations provided by Gibson, Barthlott &
are some pictures. Hunt (2000: 70 Fig. 15.5) bzw. Arroyo-
Seed Cosultchi et al. (2006: 986 Fig. 3d).
For about 25 years, I collected seeds, both As a result I have to confirm that it is nearly
impossible to compare SEMs with pictures
47
Figure 20 Morangaya pensilis from Sierra Cacachi-

las, cultivated specimen. Foto Leon de la Luz.
For habitat pictures E of La Paz & N of road to San
Juan de Los Planes; vicinity of Cerro El Llano,
Sierra Cacachilas, all originated by Dr. J. Rebman
see: www.bajaflora.org
(http://www.bajaflora.org/BajaPhotoSearch.aspx)
Map 1 Generalized distribution of Morangaya pen-

silis in Baja California Sur [black spots]: Recently
documented populations from north to south: Sierra
Cacachila, ca.1000m (Leon de la Luz, 2005), [white
doted] presumed type location (Picacho) in Sierra
de la Laguna ca.1800m (Römer, 2002, Wolf &
Wolf, 1999), Sierra de la Laguna 600m (Breslin,
pers. com.), San Pedro de la Soledad ca.900m
(Römer, 2002) and San Jose del Cabo, ca.600m
(Breslin, pers. com.).
Figure 21 A specimen of Morangaya pensilis at the
produced by a conventional microscope and so called Todos Santos (600m) site in flower
seeds from different sources do not look Photograph by P. Breslin.
similar ! & 8) respectively to S. stellatus (see Barthlott &
As a basis for interpretation of the seed Hunt, 2000: 70 Abb. 15.1-2). But there is clearly
characters we should remind ourselves of the a difference compared to the one species of
diagnosis given by George Engelmann (1859: “Rathbunia” available as SEM showing flat
28) describing Echinocereus seeds. polygonal cells, being convex in the ventral
part and more concave in the lateral part of the
Following the character of the seed’s surface
seed. Stenocereus gummosus instead is
(testa) it is possible to consider Morangaya as a
surprisingly smooth (see Barthlott & Hunt,
member of the genus Echinocereus (or as a
2000).
member of Stenocereus whose seed characters
are not cited herein). There is a clear affinity to Habitat
the seeds of S. standleyi (see Gibson:287 Abb 6 Although “Machaerocereus” is well known
48
Figure 22 The situation at a well know habitat of Figure 23 In habitat, ca.2011; the thicker bodies
Morangaya pensilis at ca.1800m, Sierra de la La- belong to the hybrid Photograph by P. Hallmann
guna Photograph by P. Hallmann
Figure 25 P. Hallmann is touching the hybrid, so it

Figure 24 P. Berresford visited the same location in is real! Photograph by P. Hallmann
2001 and also photographed the hybrid
from Baja California, S. gummosus is also well
distributed along the coastal area in the State
of Sonora. So we have this species with a very
wide distribution and the well known S. eruca
with a very limited distribution. The
rathbunias inhabit the western mainland of
Mexico in a stripe between Sonora and Colima.
What I consider a kind of sensation and the
background for this report is not the
recognition of Morangaya as a separate genus
but the discovery of a very few populations of
Figure 26 P. Hallmann, in habitat, ca.2011; the
the latter. ‘Pensilis’ was only known from its
thicker bodies belong to the hybrid
type area in Sierra de la Laguna but as of today Photograph by P. Hallmann
we have some reports of a more northern area, some surprises!
Sierra Cacachilas (Leon de la Luz, 2005) and
more southern area near to the Cape Region A Natural Hybrid
(Breslin pers. com.). That leads me to the One of those surprises is very obviously the
question, if the species will once become recognision of a natural hybrid in a well
located in the Sierra Gigantea to! known population of Morangaya in the Sierra
Considering that Baja is well known to Laguna (if it is not a genetic mutant). So we
many botanists we should still be prepared for can only know it by the photographic
49
documentation from the Sierra de la Laguna. FRANK, G.R.W. (1986). Die Echinocereen von
But various cactophiles have been able to bring Baja California (3). Kak. and. Sukk. 37(7): 130–
pictures of that entire plant, but none of them 133.
was aware of that entity! Its coming out was GIBSON, A.C. (1988-1989). The systematics and
just on my desk when putting many pictures evolution of Subtribe Stenocereinae. CSJ 60–
together, circumscribing the morphology by 61
in-situ specimens. LANGE, M. (1998). Stenocereus gummosus. –
In the local cereoid plant community only Kak.and.Sukk. 49(5): 1998/10 Karteiblatt
Stenocereus thurberi and Pachycereus schottii LANGE, M. (2016). (Steno) Cereus
appear, so one of those is the potential sexual pseudosonorensis Gürke. The Cactus Explorer
partner. (online journal) 16: 28–34.
LEON DE LA LUZ, J.L. (2005). Evaluation of the
Summing it all up, it is one thing to conservation status of Morangaya pensilis.
philosophize about Morangaya in the Onyx 39(2): 219–222.
greenhouse or at a desk or under a microscope. NIESTRADT, W. (1982). Rathbunia alamosensis
It is another thing and physically much more (Bildpostkarte) - Planet-Verlag Berlin A4-
stressing to find it in nature. So my great 302542/82-III/26/13 12821
thanks are sent out to our Mexican, US- NIESTRADT, W. (2000). Rathbunia sonorensis aus
American, French, English, Austrian and der Sonora-Wüste. - Kak.and.Sukk. 51(10):
German colleagues for providing top secret Titel & 260.
data to me and very nice pictures to all NIESTRADT, W. (2000a). in litt. 31.10.2000 (2
readers. Especially I want to thank the late Dr. pages)
Richard Chr. Römer and the holder of his POREMBSKI, S. (1992). Stenocereus eruca. –
picture heritage. Richard made the “Cuesta del Kak.and.Sukk. 43(6): 1992/18 Karteiblatt
Infierno” a few times more than a decade ago. RöMER, R.CHR. (2002). Zur Blütezeit am
Acknowledgements Fundort des E. pensilis. Kak.and.Sukk. 53(4):
99–104.
P. Berresford, P. Breslin, Dr. L. Diers, Dr.
ROWLEY, G.D. (1974). The unhappy medium:
G.R.W. Frank, M. Haberkorn, P. Hallmann, W.
Morangaya, a new genus of Cactaceae.
La Haye, Dr. J. Leon de la Luz, J. Lodé, H.-J.
Ashingtonia 1(4): 44–45.
Ness, U. Raudonat (†), B. Roczek, Dr. R. Chr.
SANCHEZ, D., ARIAS S. & TERRAZAS, T. (2014).
Römer (†), T. VanDevender and and G. Charles
Phylogenetic relationships in Echinocereus
for improving the translation.
(Cactaceae, Cactoideae). Systematic Botany
Literature 39(4): published online 3. September 2014
ARROYO-COSULTCHI, G., TERRAZAS, T., ARIAS, S. WALLACE, R.S. & FORQUER, E.D. (1995).
& ARREOLA-NAVA, H.J. (2006). The Molecular evidence for systematic
systematic significance of seed morphology placement of E. pensilis. Haseltonia 3: 71–76.
in Stenocereus (Cactaceae). Taxon 55: 983–992. WOLF, F. & WOLF, R. (1999). Baja California und
BARCENAS, R.T., YESSON, CH. & HAWKINS, J.A. seine Inseln. F. & R. Wolf und Arbeitskreis
(2011). Molecular systematic of the für Mammillarienfreunde e.V.,
Cactaceae. Cladistics 27: 1–20. Sulz/Wienerwald & Homburg/Saar. ISBN 3-
BARTHLOTT, W. & HUNT, D. (2000). Seed- 926573-10-4
diversity in the Cactaceae. Succulent Plant
Research 5. Sherborne, 173 S. M. Lange
CORBETT, T.L. (1998). The Echinocereus Index. [email protected]
Cactus & Co 2(3) Special Issue.
EARLE, W.H. (1972). Octopus Cactus.
Saguaroland Bulletin 26: 80–81.
50
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standard and
+ English summaries. published 4 times per
Also available are a se- year since 1977.
ries of separate books about particular groups
of Echinocereus. Articles in English as
well as German.
http://www.arbeitsgruppe-echinocereus.de
http://www.mammillaria.eu/en_index.html
51
The Sedum Society The Tephrocactus Study Group

Publishes articles principally about the
smaller South American Opuntias, including
such genera as Cumulopuntia, Punotia,
Maihueniopsis, Tephrocactus and Pterocactus. The
smaller North American Opuntias are also
sometimes included.
Since 2013, the journal has no longer been
printed but articles may be viewed free on the
new website. An annual meeting is also held.
Website: http://www.cactus-mall.com/sedum/ Secretary: John Betteley, 25, Old Hall Gardens,
Download information leaflet here Coddington, Newark, Notts. NG24 2QJ U.K.
Tel: +44(0)1636 707649
[email protected]
Internoto
The specialist society for http://www.tephro.com
the study of Notocactus.
(German with English Succulenta
summaries) First published in 1919,
A well-produced journal this is the journal of the
published 4 times per Dutch Cactus Society,
year since 1980. Succulenta.
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a year, this journal has
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history.
IN T E R N AT I O N A L S A N S E VIE R I A S O C IET Y
Dutch with English

Learn about the enormous variety of Sansevierias an
and how to
summaries.
grow them by joining the INTERNATI
INTERNATIONAL SANSEVIERIA
SOCIETY.. We have members world
SOCIETY
colour journal three times per subscripti
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journal also covers the related Dr
Subscription £25 or €28 per year (UK an

countries), £29 or US$46 for
for airmail d
International
outside of Europe. For further details write
to: Alan Butler, via della Campana 7,
00048 Nettuno (RM), Italy or e-mail
Cactus
[email protected]. You You can also
subscribe on line at www.sansevieri
Adventures
international.org where you can also fin
local representatives.
The Mammillaria Society The first European

cactus magazine.
Published in three
The UK-based specialist
editions: English,
society for the study of
French and Spanish.
Mammillaria and allied
Well-illustrated
genera.
articles about plants
in habitat and culti-
Back issues on DVD
vation.
Extensive annual seed list.
4 issues per year, Published twice per year by the famous
seed list and meetings. cactus and succulent enthusiast Joël Lodé.
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52
INTER NATIONAL ASCLEPI AD S O CIET Y

The INTERNATIONAL ASCLEPIAD SOCIETY is for all those
interested in the Asclepiads and all members of the Apocynaceae
family, particularly the succulent species. Asklepios journal 3
times a year, seed lists, meetings, CDs
SUBSCRIPTION: £17 (€20) per year for the UK and
other European countries, £21 (US$33) for airmail
outside Europe.
Write to: Tim Marshall, 17 High Street, Wighton,
Wells-next-the-Sea, Norfolk NR23 1AL, UK
[email protected]
Or e-mail: [email protected]
Or visit our web site: www.asclepiad-international.org
for on-line subscription or details of local
representatives.
Piante Grasse
First published in 1981,
this is the journal of the
Italian Society A.I.A.S.
Published 4 times a year,
it has a long history as
the principal journal of
Italy.
Italian language.
Subscription, including
Special Issues: 40€ or
outside Italy: 50€.
http://www.cactus.it
53
"Avonia", the quarterly journal of the German Society for other Succulents.
Written in German with English summaries, non-German manuscripts in original
language. Includes colour photographs, excellent drawings and articles on all aspects of
other Succulents. See our website: www.fgas-sukkulenten.de
Annual subscription: Germany: 30 € incl. P&P Other countries: 35 € incl. P&P
Also available is the free colour online journal "Sukkulenten", Annual seed list for
members and much more. Special interest groups for Aloe (incl. Haworthia a.s.),
Ascleps, Euphorbia, Mesembs and Yucca/winter-hardy Succulents.For membership
and further information please contact:
Dr. Jörg Ettelt: Morgenstr. 72, D-59423 Unna, [email protected] or
Wilfried Burwitz: Postfach 100206, D-03002 Cottbus,
[email protected]
Kaktusy is an international (Czecho-Slovak) journal

about cacti and succulents with a lot of interesting articles
(travelling, descriptions, growing, exhibitions, books, tax-
onomy) published since 1965. It is in the Czech language
with summaries in English and German.
Volume 2013 has 292 pages (4 regular issues and 2 spe-
cial issues), 485 colour photos and 50 B&W photos.
Price: 180 CZK + postage (about EUR 8 + postage or
about USD 11 + postage)
.http://www.cactus.cz/english/kaktusy/kaktusy_2013/
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men indes mehr und mehr n der Pflanzen neh- 1994 + 2005). However
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See our website for information:

Ing.
eber sc
R e i s d b ra s i l i
ü
Im ssü
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www.gymnocalycium.info
.gymnocalycium.info
54
> [email protected] <
PLANTS AND SEEDS FOR SALE

Your place to advertise spare plants, seeds, journals, books etc. FREE!
Entries are free so please send me the text of your advert which can include
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Seeds from Aymeric de Barmon (The picture above is his glasshouse)

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Seeds of Chilean Cacti František Nechvil Seeds

Collected in habitat or in our collection. I offer cactus seeds of first-rate quality,
Visit our website to download our lists: all from my own collection.
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If you are in Chile, you are welcome to visit
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New seed list for 2015/2016
Ingrid Schaub & Ricardo Keim, Olmué, Chile
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Plants offer: http://www.cact.cz/nabidka-ros-
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Seeds from Ralf Hillmann
New seed list available in December.

All seeds are from habitat source or
pollinated in my own collection.
Specialising in Aylostera, Mediolobivia,
Rebutia, Sulcorebutia, Lobivia, Echinopsis,
Gymnocalycium, Notocactus, Cereoids, Bercht Seed List
Opuntiae, Mesembs, Puya, Crassulaceae etc. Already familiar to many of you, for others a voyage
For more information please contact of discovery. Many seeds you are looking for can be
found in this seed list. A large number of the seeds is
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A comprehensive list of seeds from the Czech The list specialises in South American cacti such as
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http://www.cactus-hobby.eu Wigginsia, but above all an almost complete selection
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An extensive seed list from Prochazka, strong many seeds of North American cacti in this list.
on Mexican cacti: www.bercht-cactus.nl
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The new offer of the seeds (mostly with Mesa Garden

locality data) is available from Jan Jecminek P.O. Box 72, Belen, New Mexico, USA
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Large list of Cactus and Succulent seeds,
www.cactus-succulent.com
particularly strong on Mexican and U.S.
Lifestyle Seeds
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Note the new emil address
Supplier of South African seeds
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email: [email protected] email: [email protected]
56
Mondocactus Echinocereus Online Services

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Seeds Martina & Andreas Ohr
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SEEDLIST of 325 forms of Echinocereus
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species. Many of the plants are provided with transfer – Special discounts on Book Sets
detailed information regarding their natural
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Mondocactus is the virtual store of De
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Echinocereus Online Web
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The Genus – Culture – Habitat – Field Numbers –
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Supplementary information
Echinocereus pectinatus www.pectinatus.de

Online shop for Cacti, Succulents and other Flower galleries of Echinocereus pectinatus
Exotic plants. We offer a good range of plants,
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(also English language)
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57
Opuntia Web.
Richard and Wendy Edginton

The Nurseries, Station Road, Flordon,
Norwich, Norfolk, NR15 1QR UK
We stock a wide range of cacti from North and This website about the opuntias of the USA
South America, raised from seed at our nursery. has been redone and is better than ever. There
Send a stamped addressed envelope for our list or are 1,500 photographs of the various species
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William’s CaCtus The oldest cactus nursery

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Pontefract, W. Yorks WF9 2RS England
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58
Southfield Nurseries - Cactusland

Bourne Road, Morton, Bourne,
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Tel: +44 (0)1778 570168
×Chamaelobivia ‘Lincoln Gem’ ×Chamaelobivia ‘Lincoln Firecracker’
×Chamaelobivia ‘Lincoln Pink Diamond’ Echinopsis ‘Bourne Razzle Dazzle’ Echinopsis ‘Shades of Bourne’
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All of our plants are propagated here in our own nursery and we have many thousands of cacti ready potted for
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Mail order service.
Visitors welcome (please check opening hours)
Enquiries: [email protected]
Website: http://www.cactusland.co.uk
Conophytum Plants
Chris Rodgerson, 35, Lydgate Hall Crescent,
Sheffield S10 5NE England
Nursery grown plants and cuttings.
See my website for information about me, the
plants and a list of plants for sale in season.
Toobees Exotics Web: http://www.cactus-succulent.com

e-mail: [email protected]
Bob and Beryl Potter, 20 Inglewood,
Woking, Surrey GU21 3HX UK Abbey Brook Cactus Nursery
South African and Madagascan succulents.
You are welcome to visit:
Visitors are welcome by appointment at our Wednesday to Friday 13.00 –16.00
nursery in Blackhorse Road, Woking.
Saturday & Sunday 13.00 – 17.00
Please send stamped, addressed envelope for list Old Hackney Lane, Darley Dale, Matlock,
or visit our new website Derbyshire DE4 2QJ UK
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Tel: +44(0)1629 580306
email: [email protected] Email: [email protected]
Telephone: +44 (0)1483 722600 www.abbeybrookcacti.com
59
J & J Cactus and Succulents
Joyce Hochtritt
600 N. Pine St., Midwest
City, OK 73130, USA
Brookside Nursery
(+1) 405-737-1831
Contact address: via della Campana 7, 00048 Nettuno (RM), Italy
[email protected]
Website: www.brookside-nursery.com
E-mail: [email protected]
www.jjcactus-succulents.net
Tel: 00-39-335-6159058
We are open to visitors
Visits By Appointment Only

but strictly by prior appointment.
Visit our web site

at www.brookside-nursery.com for more
Corona Cactus Nursery

information and our plant list.
We offer an efficient mail order service and
probably the most comprehensive list
of Sansevierias, and Aloaceae in Europe.
We also stock a wide range of other
• Specializing in collector cacti and succulents

succulents, especially asclepiads,
• Mail Order - We ship to most countries,

euphorbias and caudex plants.
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The Cactus Man • The majority of our plants are seed grown
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• Quarterly newsletter, cultivation articles,
Good Selection of Cacti & Succulent Plants photo galleries and more ....
Opuntia Specialist
http://www.coronacactus.com
See Website for Days of Opening & Events
18 Bodgara Way, Liskeard, Cornwall PL14 3BJ UK
Connoisseurs’
Email : [email protected]
Web : www.thecactusman.co.uk
Cacti
ebay: thecactusmanuk
Opuntias (plants & cuttings); large cacti always wanted
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Interesting Website and on-line shop cacti/index.html
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PlaNtlife
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Kakteen - Piltz Cactus & Succulent Nursery

Cacti, Succulents and Seeds Beechwood, 50, Friday Street, Eastbourne,
http://www.kakteen-piltz.de East Sussex, BN23 8AY U.K.
Nursery open for visits. Stuart & Jane Riley
Tel +44(0)1323 761943 FAX +44(0)1323 724973
Monschauer Landstrasse 162,
email: mailto:[email protected]
D-5160 Düren - Birgel, Germany
milenaudisio
● Cactus & succulent Nursery in italy ● seed grown plants
● specializing in Gymnocalycium and lithops ● mail order service
Via Casalgrasso 1/a, 10022 Carmagnola (To) Italy

Web: http://www.milenaudisio.it Email: [email protected]
60
Marie’s Pots
Tony Irons Cacti

Have a look at our range
of high quality new and used pots,
BEF, Optipot, Plantpak etc.
Reliable Mail Order Service
17, White Lodge Park,
Portishead, Somerset BS20 7HH web: www.mariespotsdirect.co.uk
Good Quality Home Grown Plants and Imports email: [email protected]
Lithops Plants available online
Arid Plants
Plus check out Ebay offers
Visitors welcome by appointment.
Website: http://www.tonyironscacti.co.uk
Many cacti and succulents from arid
regions around the world.
Small seedlings to specimen plants.
Mail order to UK and Europe

Tel: +44 (0)7973 630359
Plants, Books and Accessories
Email: [email protected]
Excellent web site and catalogue Web: http://www.arid-plants.co.uk
www.kuentz.com
Ets. Kuentz - 327 rue du Général Brosset - Visitors welcome by appointment.
83600 Fréjus - FRANCE Colchester, Essex, UK
Books and Journals for Sale

I have a number of books and journals
from the library of Harry Middleditch which
have been donated by his family to help
fund the creation of the Rowley Library
Trust.
Please contact me for details of condition
and very reasonable prices:
Backeberg, C.: Die Cactaceae, original edition,
5 (of 6) parts (1958–62), missing Part III.
Britton & Rose: The Cactaceae, Dover reprint
(1963) 4 volumes in two books.
Borg, J.: Cacti, First edition (1937).
Zeitschrift für Sukkulentenkunde Band 1–3
(1923–28) 3 bound volumes.
Monatsshrift der DKG Band 1–3(1929–31) 3
bound volumes.
Kakteenkunde (1938–38) 1 bound volume.
Cactaceae, Jahrbücher der DKG (1937–42) 12
softbound volumes.
Also many years of Monatsschrift für Kak-
teenkund.
Graham Charles
61
BOOKS FOR SALE

Keith’s Cactus Books
For the widest range of books on Cacti and Other Succulents

And a wide range of other plant and gardening books Delivery worldwide
Please visit www.keithscactusbooks.co.uk - As easy to browse as a paper catalogue
Please click on the book image to be directed to my website for details of the book
GYMNOCALYCIUM
IN HABITAT AND CULTURE
Copies of my book are still available from
dealers around the world or from me.
If you would like me to sign it, please ask!
Graham Charles Gymno Book Website
CHUCK EVERSON, BOOKSELLER

“Your Cactus Bookstore Specialist”
1444 E. Taylor Street Vista, CA 92084-3308 USA
Specializing in literature concerning cacti, succulents,
epiphytes, deserts, bromeliads, hoyas, palms, cycads,
and ornamental plants from around the world.
We have the latest books at the most reasonable
prices.
We also buy and sell previously owned books on
cacti and succulents.
WE SHIP WORLDWIDE. VISA/MASTERCARD IS ACCEPTED.
Check with us for the best possible times to visit.
Our 33 years of experience tells you that you are in
good hands. www.cactusbookstore.com
Tel: 760-758-4290 [email protected]
If you have not already told me and would like to be advised when the next issue of the Cactus
Explorer is available for download, please send me your E-mail address and I will add you to
the distribution list.
Contributions to any of the regular features, articles, adverts for events, plants etc. are all very
welcome. Thank you for your support!
62

Cactus Explorer 17 - Complete

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What is the purpose of the journal?

What is the purpose of the journal?

What are some of the regular features included in the journal?

What are some of the regular features included in the journal?

The

Number 17 3 Espostoa in Tenerife

ISSN 2048-0482 4 Barranco Rambla de Ruiz

Regular Features Articles

Introduction 3 A naturalised population of

Cover Picture: Siccobaccatus dolichospermaticus See page 9

NEWS AND EVENTS

†Plowes, Darrel Charles Herbert A. Castellanos & H. Lelong

What a Wonderful Day!

The BCSS National Show, August 20th 2016

The Haworthia Society

The DVD may be ordered through our PayPal Shop

Members £30.00 (UK), £31.00 Europe,

Yearbook of the British Cactus & Succulent Society

You can buy Bradleya 34, by sending £21

Payment accepted by:

The articles in this edition are:

Guided Trip to Baja California

Copiapoa decorticans Copiapoa ahremephiana Copiapoa atacamensis Copiapoa laui

Your chance to buy cards illustrated with reproductions of pictures from

Siccobaccatus, a magniﬁcent cactus genus from Brazil

Fig.1 Siccobaccatus dolichospermaticus flowering in the collection of K.J. Neirinck.

Fig.2 Siccobaccatus dolichospermaticus in habitat near to Bom Jesus de Lapa. GC1014.03.

Photo: Graham Charles

Copiapoa solaris does ﬂower in cultivation!

At least it might if you live in California. Photographs by Duke Benadom

The species was ﬁrst found in 1956 by

On-line Journals for you to download free

Sukkulenten (formerly Avonia News)

The 15th issue of this free online journal

THE LOVE OF BOOKS

Making Sense of German Journal Chronology

The many titles of cactus journals can be

1923: The title of the DKG publication changed

Vols. 1–4, 1933–1936 (DKG)

1965: A good quality

TRAVEL WITH THE CACTUS EXPERT (16)

Fig.4 Flowering Delosperma nubigenum or “Hardy

Fig.6 Escobaria missouriensis was just about to

AN EXCURSION TO BARRANCO RAMBLA

Fig. 1. Barranco Rambla de Ruiz

Fig.4.Aeonium arboreum var. holochrysum. Fig.5. Aeonium arboreum var. holochrysum.

Fig. 8. Barranco Rambla de Ruiz.

Fig.10. Aeonium canariense var. canariense.

Fig.9. Aeonium arboreum var. holochrysum.

Fig.17 Aeonium canariense var. canariense. Fig.18 Aichryson laxum.

Fig.19. Monanthes laxiflora.

Fig.20. Dracena draco in the middle of the square

Fig.21. Gallotia galloti in the Barranco Rambla de

Fig.22. Gallotia galloti on the Sendero del Agua.

Fig.23. Gallotia galloti on Sendero del Agua near

JUAB AND UTAH COUNTY, UTAH,

Figure 2: Rocky place under a juniper tree with

Figure 3: A lot of buds on Echinocereus triglochidia-

Figure 11: Echinocereus triglochidiatus v. mojaven-