Physiology Assignment 1
Physiology Assignment 1
Field Capacity is the amount of soil moisture or water content held in the soil after excess water
has drained away and the rate of downward movement has decreased. This usually takes place 2–3 days
after rain or irrigation in pervious soils of uniform structure and texture.
The physical definition of field capacity (expressed symbolically as θfc) is the bulk water content
retained in soil at −33 J/kg (or −0.33 bar) of hydraulic head or suction pressure
At “Field Capacity” (FC) the soil is wet and contains all the water it can hold against gravity. At the
“Permanent Wilting Point” (PWP) the soil is dry and the plant can no longer extract any more water. The
difference in the water content of soil between field capacity and the permanent wilting point gives the
amount of soil water available for uptake by plants. The plant available water is expected to be greater for
clayey and organic soils compared to sandy soils. If we know the plant available water and the rate at
which this water is being depleted by crops then we can determine the necessary frequency of irrigation.
Apart from irrigation scheduling, this information can also be useful in the modeling of crop growth and
prediction of yields.
For media with suctions up to 10 or even 20 kPa suction tables are in common use. The suction is usually
applied by placing the samples on a bed of very fine sand or clay in a water-tight container connected to
an adjustable overflow (Fig. 7.2).
Water retention measurements are time consuming because equilibrium between the applied suction and
the water-filled pore space is essential. Usually each measured point on a curve with 3–10 points requires
1–2 days to reach equilibrium. Frequently, the term ‘water retention’ is used for the measurement of both
the main drainage curve and the main wetting (= rewetting) curve. Common measuring points include the
water content at saturation, and at suctions of –2.5cm, –10cm, –20cm, –50 cm, –100cm and –500 cm
suction head. The air content is calculated as the difference between porosity and water content. Usually
the main drainage curve and the main wetting curve share the wettest and the driest points but differ in
between. This effect is called hysteresis. Its causes are discussed in the section on ‘Hydrophobicity’.
Hysteresis also influences the rewetting, rehydration rate and unsaturated hydraulic conductivity.
Special Cases
Various variants exist, for example using air pressure instead of water columns. Especially at higher
suction levels air will enter the sand bed of the suction table. Strict procedures for sample preparation,
saturation and measuring time have to be observed. The method is derived from soil science and its
application for media creates some interpretation problems. One interpretation problem is related to the
sample height of 5 cm, which is treated as a point sample. In fact, for small suction forces applied (from 0
to 15 cm suction force, i.e. 0–1.5 kPa), the sample height does influence the reading to a large degree.
Another interpretation problem concerns the point of 0.0 cm suction, which is defined at a free water table
up to half of the sample's height. The point of 0.0 cm suction is thus changing with the sample height.
Sample heights other than 5.0 cm are common, for example 3.5 cm for propagation plugs.
A third common interpretation problem is that many substrates used in horticulture are in practice
subjected to only a part of the measuring range mentioned. There is no point in measuring substrates such
as polyurethane foam at suctions above 20 cm of water column, since at these suctions there is no longer
water in interconnected pores.
Water retention forces in growing media are usually 10–100 times lower than the common values for soil
(10–100 kPa).
Influence on Growth
The results are indicative of the ease of the uptake of water – and nutrition – by plants as well as the
wetness in various growing systems. Growth is highest at low water retention forces, but very low
water retention forces are sometimes avoided, for example, when the amount of air-filled pores
becomes too low for proper oxygen transport. The air content recommendations for optimal
growth are found in kipp et al., 2001. Water retention forces high enough to decrease the fresh
weight growth may actually be desirable, for example to create denser, that is better quality pot
plants and in transplant production, when hardy plants are preferred by the growers.
Chemical Hydrology
The tension head at field capacity is equivalent to the height of a liquid water column in a mineral
capillary hc. The effective radius of the largest water-filled pores at field capacity r¯fc is the dependent
variable and the tension head at field capacity hfc is the independent variable.
r¯fc=2⋅γstρw⋅hfc
r¯fc=2⋅0.0728997.044⋅1.02⋅9.80665=1.49×10−5m
Soil scientists use hfc ≈−1 m for coarse texture soils and hfc ≈−3.3 m for fine-textured soils. Using these
two tension heads at field capacity, the largest water-filled pores in coarse-textured soil is about 15 μm
while the largest water-filled pores in fine-textured soil is about 4 μm. Tuller et al. (1999) provide a more
detailed discussion of water film thickness in soils, examining geometries beyond the simple round
capillary discussed here. Regardless, the results are essentially the same.
Effect of precipitation on Distribution of Plants
Water:
Water is most important environmental factor. It is essential for all life processes. The living cells have
about 80% water. It is essential for maintenance of turgidity of cells. It acts as solvent for the exchanges
of gases. All physiological processes like photosynthesis, transpiration and respiration require water.
Water covers about 73% of the surface of the earth. It is present in ponds, lakes, streams anad oceans.
Only 1% water is present in fresh water bodies like ponds, river, lakes etc. 99% water is present in
oceans. It falls from the atmosphere in form of precipitation , and water vapours.
Precipitation:
Precipitation includes rain, snow, hail, and dew. Precipitation occurs due condensation of water
vapours present in the air. Air or wind carries these vapours from oceans. Precipitation may be in the
from oceans. Precipitation may be in the form of rainfall, snow fall or hailing.
Importance of precipitation: