PRINCIPLES OF ECOLOGY

In the previous module (module-1) you have learnt about the origin and
evolution of the
environment. You have also learnt how humans have been interacting with the
environment.
In this lesson, which is the first one in the module-2, you will learn some
important concepts
of ecology which is an established branch of science.

OBJECTIVES

After completing this lesson, you will be able to:

define the term ecology;
explain the relationship between organism and its habitat with a special
mention
of the human species;
recognize the levels of ecological organizations from organism (individual) to
population, community, ecosystem, biome and biosphere;
differentiate between habitat and niche;
describe the concept of species and explain the basic idea of adaptation,
evolution
and extinction;
explain the concept of population with reference to organisms;
highlight the characteristics of size, growth, density and dispersion of
population;
analyze the demographic factors influencing the changes in population of
organisms;
explain community structure with reference to species diversity, interspecific
interactions and ecological succession.

4.1 DEFINITION OF ECOLOGY

Ecology may be defined as the scientific study of the relationship of

living
organisms with each other and with their environment.
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The term ecology was first coined in 1869 by the German biologist Ernst
Haeckel. It has
been derived from two Greek words, oikos, meaning home or estate and
logos meaning
study. The emphasis is on relationships between organisms and the
components of the
environment namely abiotic (non-living) and biotic (living).

4.2 LEVELS OF ECOLOGICAL ORGANIZATION

Ecology not only deals with the study of the relationship of individual organisms
with their

environment, but also with the study of populations, communities, ecosystems,

biomes,
and biosphere as a whole (see Fig. 4.1)
Fig. 4.1: Levels of ecological organization and types of ecological studies
I. Study of physical and
biological features of a
particular biome.
II. Study of human activities
affecting the earth like global
climate, ozone hole etc.
A large community unit, characterized by a major
vegetation type and associated fauna, found in a specific
climatic region is a biome
Biomes refer basically to terrestrial areas. The aquatic
systems like the seas, rivers etc. are also divided into
distinct life zones on basis of salinity.
Study of
Levels of
ecological
organization
1. Organisms
(individual)
basic unit of
study
2. Population
A group of organisms
consisting of a number
of different populations
that live in defind area
and interact with each
other.
3. Community
A group of organisms
consisting of a
number of different
species that live in
an area and interact
with each other
4. Ecosystem
A communities of
organisms and their
physical
environment,
interacting as an
ecological unit.
Types of
study
Study of the form,
physiology, behavior,
distribution and
adaptation of
organism in relation
to environment.
Study of
interaction between
populations and
intraspecific
relationships.
Study of structure
and composition
of community and
interspecific
interactions
between members
of community.
Study of the
community in relation
to the structure of its
ecosystem-nutrients
cycling, climate,
energy flow etc.
studied.

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4.3 HABITAT AND ORGANISM

Habitat is the physical environment in which an organism lives. Each organism

has particular
requirements for its survival and lives where the environment provides for
those needs.
The environmental requirement of an elephant would be a forest. You would not
expect an
elephant in the ocean nor would you expect a whale in the forest? A habitat
may support
many different species having similar requirements. For example, a single
ocean habitat
may support a whale, a sea-horse, seal, phytoplankton and many other kinds of
organisms.
The various species sharing a habitat thus have the same address. Forest,
ocean, river
etc. are examples of habitat.
The features of the habitat can be represented by its structural components
namely (1)
space (2) food (3) water (4) and cover or shelter (Fig. 4.2).
Earth has four major habitats-(1) Terrestrial (2) Freshwater (3) Estuarine
(Where
rivers meet the ocean) and (4) Ocean. The human gut is the habitat of a
tapeworm and the
rotting log a habitat of a fungus.
Fig. 4.2: Structural components of a habitat

4.4 NICHE AND ORGANISM

In nature, many species occupy the same habitat but they perform different
functions. The
functional characteristics of a species in its habitat is referred to as niche in
that common
habitat. Habitat of a species is like its address (i.e. where it lives) whereas
niche can be
thought of as its profession (i.e. activities and responses specific to the
species). The
term niche means the sum of all the activities and relationships of a
species by
which it uses the resources in its habitat for its survival and
reproduction.
A niche is unique for a species while many species share the habitat. No two
species in a
habitat can have the same niche. This is because if two species occupy the
same niche they
will compete with one another until one is displaced. For example, a large
number of
Water
Space
Food
Shelter or

cover

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different species of insects may be pests of the same plant but they can coexist as they
feed on different parts of the same plant. (Fig. 4.3).
Fig. 4.3: Different species of insects feeding on different parts of the same
plant
Another example is the vegetation of the forest. The forest can support a large
number of
plant species as they occupy different niches: the tall trees, the short trees,
shrubs, bushes
and grasses are all part of the forest but because of varying heights they differ
in their
requirements for sunlight and nutrients and so can survive together (Fig. 4.4)
Fig. 4.4: The three species of warbler birds search for insects as food in the
forest
at different levels in the tree and so occupy different niches
Lacebug
Shoot
moth
larva
Serpentine
leaf miners
Root-feeding white grub
Bay-breasted warbler
Byrtle warbler
Blackburnian warbler

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Ant, insect
Fruit
Cow Lion
Oyster snail
Fish
Whale
Fox
Hen
Grass
Goat

The most important resources in the niches of animals are food and shelter
while in case of
plants, they are moisture and nutrients (phosphorous and nitrogen).Fig. 4.5

shows the
niche of human beings.
Fig. 4.5: The ecological niche of human being

INTEXT QUESTIONS 4.1

1. What does the term ecology mean?

______________________________________________________________
2. Define the term niche.
______________________________________________________________
3. Give one point of difference between habitat and niche.
______________________________________________________________

4.5 ADAPTATION

Every organism is suited to live in its particular habitat. You know that the
coconuts cannot
grow in a desert while a camel cannot survive in an ocean. Each organism is
adapted to its
particular environment. An adaptation is thus, the appearance or
behaviour or
structure or mode of life of an organism that allows it to survive in a
particular
environment. Presence of gills and fins are examples of adaptation in fishes
to aquatic
habitat. In aquatic flowering plants absence of wood formation and highly
reduced root
Crab

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system are adaptations to aquatic environment. Adaptations that can be
observed in structure
(Fig. 4.6) or behaviour or physiology of an organism. Adaptations have genetic
basis and
have been produced and perfected through evolution. This means that the
adaptions have
developed over many generations to help a species survive successfully in its
environment.
Examples of basic adaptations that help animals and plants to survive in their
respective
environments.
Shape of birds beak.
The thickness or thinness of fur.
Presence of feathers and wings in birds.
Evergreen and deciduous nature of trees.
Presence and absence of thorns on leaves and stems.
Fig. 4.6: Adaptation in the types of beaks in birds: The beaks of different birds
are
adopted for feeding on different kinds of food
What is a Species

A species is defined as; a group of similar populations of organisms whose

members are
capable of interbreeding, and to produce fertile offspring (children). A tiger, a
lion, a lotus
and a rose are examples of different species. Every species has a scientific
name, understood
by people of all over the world. Humans belong to species of Homo sapiens.
Only
members of the same species can interbreed to produce fertile offspring. Every
species
has its own set of genetic characteristics that makes the species unique and
different from
other species.
Variation
However, species are generally composed of a number of distinct populations
which freely
interbreed even though they appear to be different in appearance.
A long thin beak is useful for
pulling worms out of mud
The strong beak of parrot is
useful for curshing nuts
This flat beak helps in feeding on
fish, insects and plants in water

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Difference in colour of skin, type of hair; curly or straight, eye colour, blood
type among
different ethnic groups represent variation within human species. Similarly,
different shape
and size of cows, dogs and cats etc. are examples of variation with in each of
these species
(Fig. 4.7). In plants, tall and short pea varieties, various shape and size of
brinjals exhibit
variation among these plant species. Variations are produced as a result of
chance mutation.
Competition and natural selection determines as to which variation will
succeed and survive.
Those variations that enable a species to survive in the struggle for existence
are encouraged
and promoted.
In plants one can observe wide variation in size and shapes of mangoes,
brinjals etc.
Fig 4.7: The populations of these four types of dogs look different but all are
capable of breeding among themselves and capable of producing fertile
offspring.
All four belong to same species Canis lupus.
Evolution

A valid theory of evolution was propounded by Charles Darwin and Alfred

Wallace in
1859. This theory has been extended in the light of progress in genetics and is
known as
Neo-Darwinism. It has the following features:
1. Organisms tend to produce more off springs that can be supported by the
environment.
2. Mutation (a change in genetic material that results from an error in
replication of
DNA) causes new genes to arise in a population. Further, in a sexually
reproducing
population, meiosis and fertilization produce new combination of genes every
generation,
which is termed recombination. Thus members of the same species show
variation
and are not exactly identical. Variations are heritable.
3. An evolutionary force which Darwin termed natural selection, selects
among
variations i.e. genes that help the organism to adopt to its environment. Such
genes
are reproduced more in a population due to natural selection.
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4. Those offspring which are suited to their immediate environment have a
better chance
of surviving, reaching reproductive age and passing on the suitable
adaptations to
their progeny.
5. Evolution thus results in adaptation and diversity of the species.
Fig 4.8: Process of natural selection

4.6 SPECIES FORMATION: SPECIATION

The number of species surviving in the world today is the outcome of two
processesspeciation
and extinction.
Speciation is the process by which new species are formed and evolution is
the mechanism
by which speciation is brought about.
A species comprises of many populations. Often different populations of a
species remain
isolated due to some geographic barrier such as mountain, ocean, river, etc.
Geographic
Sexual reproduction results in
more offspring than
environment can support
Inherited variation expressed in
phenotypes of offspring
Struggle for survival
Some rabbits with thinner
coats die of exposure

Some weaker rabbits die

of disease
Some rabbits lose out in
competition for food and
die
Some slower rabbits are
caught by predators
Healthiest, strongest,
fastest, best insulated
rabbits survive and
breed
Their genes get represented more in the population. Thus National
selection brings about differential reproduction and results in
adaptation

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isolation occurs when a physical barrier develops between two populations of a
species as
you can see in fig. 4.8. The most common way a population undergoes
speciation is by
geographic isolation.
The members of a population of a species live in a particular environment and
are
capable of breeding with the member of another population of the same
species.
The population then becomes separated into two completely isolated
populations by
a barrier which prevents their interbreeding and gene exchange. The isolating
mechanism
may be a physical barrier like water, mountain, ocean represent geographical
isolation.
(Fig. 4.9)
Fig. 4.9: Geographical isolation
Large interbreeding population
of one species A
Isolation by barrier
Sub-population B Sub-population C
Mutation
Large mutant Small mutant
Natural selection
Large size enjoys selection
advantage (e.g. in cold climate)
Small size enjoys selection
advantage (e.g. in warm climate)
Further natural selectionleads to
formulate of two species
Species A Species B
Barrier removed
Species B cannot
breed with species
C or with original
parent species A
BA
RR

IE
R
BA
RR
IE
R
BA
RR
IE
R
BA
RR
IE
R

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Ecological isolation caused by differences in temperature, humidity, pH etc. in
the
environment of the two populations.
Reproductive isolation caused by interference in interbreeding between
members of
different populations of species i.e. species. When two populations of a species
are
unable to interbreed due to reproductive barrier.
Reproductive isolation may occurs due to any one or more of the following
reasons:
(i) When two different populations become sexually receptive at different times
of
the year. For example a population of frogs that breeds in May is effectively
isolated
from one that breeds in July, though both populations may occur in the same
area.
(ii) Members of different populations are not attracted by courtship behaviour
towards
one another.
(iii) Pollination mechanism fails, between flowers of two populations.
(iv) Cross fertilization is prevented as sex organs of different populations of a
species
do not match.
Mutations occur randomly in isolated populations giving rise to new variation
within
each sub-population of these mutations those that help to adapt to the
environment are
reproduced in greater numbers in the next generation due to natural selection.
In other words since no two environments are identical, natural selection
pressures
that occur on each separate sub-population are different, depending on local
conditions
such as climate, disease, predators etc. Natural selection affects each subpopulation
differently and so different variations caused by nutrition or recombination in
different

subpopulations get established. With the passage of time, the sub populations
become
more and more different from each other.
After a long period of time, the sub-populations become very different and get
isolated,
reproductively, i.e. they no longer interbreed.
Later even when the barrier is removed the sub-populations are unable to
interbreed
and thus subsequently the sub-populations become two different species.
An example of formation of new species
A current example of speciation can be seen in the two species of squirrels
Kaibab squirrels
and Abert squirrels that live on opposite sides of the Grand Canyon. Biologists
assume
that the two squirrel populations became separate species when about one
million years
ago, the Colorado river changed its course, splitting the original population of
squirrels into
two. Since the environment on opposite sides of the canyon is different,
different
characteristics were favoured on each side of the canyon, by natural selection.
After many
years of separation, the genetic differences between the populations became
so large that
the two squirrel populations became two separate species. They look different
and can no
longer interbreed (Fig 4.10).
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Fig. 4.10: The Kaibab squirrel of the north rim and the Abert squirrel of the
south
rim had common ancestors
4.6.1 Extinction
Ever since life evolved on earth, new species better suited or adapted to the
environment
have appeared and older less successful forms have died or become extinct.
Extinction is
generally a natural occurrence. It means the dying out of a variety of or a
species. The
primary reason for these extinctions is environmental change or biological
competition.
Extinction occurs when species cannot evolve fast enough to cope with the
changes taking
place in their environment. (Fig. 4.11). Many species have gone extinct during
geological
history of the earth. Fossils are, the preserved remains of animals, plants, and
other

organisms that lived in the geological past.

(a) (b)
Fig. 4.11: (a) Fossil of fern plant. (b) Fossil fish
Extinction may take place due to catastrophic natural phenomena such as
tsunami, volcanoes
etc. In recent time, human activities such as deporation, over exploitation,
environmental
Kaibab Squirrel (North Rim)
Sciurus kaibabensis
Aberr Squirrel (South Rim)
Sciurus aberti

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pollution and environmental change are other factors responsible for extinction.
Deforestation
for expansion of industries and human settlements has promoted economic
growth but at
the same time it has resulted in habitat loss for many wild plants and animals.
Pollution has
killed many an aquatic species.

INTEXT QUESTIONS 4.2

1. What is meant by the term adaptation? Answer in one sentence.

______________________________________________________________
2. Define : (i) species (ii) variation
______________________________________________________________
3. Name two sources of variation.
______________________________________________________________
4. Name the evolutionary force which brings about greater reproduction of
adaptive
variation.
______________________________________________________________
5. Explain the term (i) speciation and (ii) extinction.
______________________________________________________________

4.7 POPULATION

Population is defined as a group of freely interbreeding individuals of the

same species
present in a specific area at a given time. For example, when we say that the
population of
a city is 50,000, we mean that there are 50,000 humans in that city. However,
all populations
of humans living in any part of the world constitute the species Homo
sapiens.
A population has traits of its own which are different from those of the
individuals forming
the population. An individual is born and dies but a population continues. It
may change in
size depending on birth and death rates of the population. An individual is
either female or

male, young or old but a population has a sex ratio and age structure, which
means, the
ratio of male to female in the population and the various age groups into which
the population
may be divided.
The characteristics of any population depends on:
(i) density of the population, (ii) natality (birth rate), (iii) mortality (death rate),
(iv) dispersal,
(v) biotic potential (vi) age distribution (vii) dispersion and (viii) growth form.
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Density: The number of individuals per unit area at a given time is termed as
population
density. The density of species varies from time to time and from one place to
another.
For example, you may notice more plant and animal species in the garden
during the
monsoon season. Density of a particular organism in a region is determined by
selecting
random samples of a particular dimension size called quadrat from that region.
In case of large, mobile animals like tigers, leopards, lions, deer etc, the density
may be
determined by counting individual animals directly or by the pugmarks (foot
imprints) left
by the animals in a defined area (Fig. 4.12). Pugmarks of each individual
animals are
unique and different from one others. Study of pug marks can provide the
following
information reliably if analyzed skillfully:
Presence of different species in the area of study.
Identification of individual animals.
Population of large cats (tigers, lions etc.).
Sex ratio and age (young or adult) of large cats
Fig. 4.12: Pugmark of a lion/cat (Foot prints) of soft padded wild animals
Counting of human population is called census and is carried out by the Indian
government
every 10 years. In census however each individual is physically counted.
Natality: The rate at which new individuals are born and added to a
population
under given environmental conditions is called natality. Birth, hatching,
germination
and vegetative propagation cause an increase in the number of individuals in a
population.
Wolf
Hyaena
Lion
Pagmark of a cat

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In case of humans, natality or birth rate is usually expressed in terms of births
per
thousand per year.
Mortality: Loss of individuals from a population due to death under given
environmental conditions is called mortality. The number of individuals dead
in a year
is calculated for obtaining the mortality rate or death rate. Mortality rate in
human
population may be expressed in terms of number of persons dead per thousand
per
year.
Dispersal: The movement of individuals of a population out of a region on a
permanent
basis is termed emigration while immigration refers to the movement of
individuals into
a new area where dispersal includes both emigration and immigration of
individuals.
The population of a region is affected by dispersal. Active migration is not
possible in
plants though seeds may be dispersed over long distance by wind, water and
animals.
The density of a population thus basically depends on four factors: i) natality, ii)
mortality,
iii) immigration and iv) emigration (Fig. 4.13)
Immigration
(+)
(+) (-)
Natality Density Mortality
(-)
Emigration
Fig. 4.13: Parameters of population
Age distribution
Natural populations include individuals of all age groups. It, therefore, becomes
necessary
for us to consider age distribution of a population. Age distribution refers to the
proportions
of individuals of different age groups in a population.The population may be
broadly divided
into three age groups: pre-reproductive group: comprising of juvenile individuals or children,
reproductive group: consisting of individuals capable of reproduction ,
post-reproductive group: contains aged individuals who are incapable of
reproduction.
A rapidly growing population will usually contain a large proportion of
individuals in the

reproductive age group; a stationary population (where there is no increase or

decrease in
population) contains an even distribution of all age groups, and a declining
population
contains a large proportion of old or post-reproductive age of an individuals.
Sex ratio
Sex ratio is an important aspect of population. It refers to the ratio between
female and
male individuals in a population
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4.8 POPULATION GROWTH

The growth, stability or decline in number of individuals in a population is
influenced by its
relation with the environment.
Populations have characteristic patterns of growth with time, which is depicted
by population
growth curves. Two basic forms of population growth curves can be identified.
(i) J
shaped growth curve and the (ii) S shaped or sigmoid growth curve.
Density independent population growth
A forest fire may reduce a dense or scanty population drastically. Extreme
weather conditions
like drought, rains, floods, storms and sudden rise or fall in temperatures all act
as density
independent factors as they tend to cause sudden reduction in population
numbers. The
population growth that is depicted by a J shaped growth curve is called
density independent
growth.
Generally the J shaped growth curve is typical of the species which reproduce
rapidly
and which are greatly affected by seasonally fluctuating environmental factors
such as light,
temperature and rainfall. In this type of curve, population density increases
rapidly in
exponential (geometric) progression (total number doubles at regular intervals
of time) like
this:
2 2 2 2 2
8 16 32 64 128 till a peak is reached.
This type of exponential growth occurs in nature when a population has
abundant supply
of resources. After reaching a peak there is a sudden crash or decline due to
environmental
or other factors. Such type of growth may be exhibited by insect populations
which show

explosive growth during the monsoon season and then abruptly disappear at
the end of the
season.

INTEXT QUESTIONS 4.3

1. Define population.
______________________________________________________________
2. Name at least three characteristics of population.
______________________________________________________________
3. What are the factors on which density of a population depends?
______________________________________________________________
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4.9 COMMUNITIES AND THEIR CHARACTERISTICS

In ecology the term community, or more appropriately biotic community,

refers to the
populations of different kinds of organisms living together and sharing the
same habitat.
4.9.1 Organization of a biotic community
The characteristic pattern of the community is termed as structure of the
community and
is determined by:
the roles played by its various populations;
the range of its various populations;
the type of area that is inhabited by the populations of the community;
the diversity of species in the community;
the interactions between various populations of the community inhabiting the
area.
Members of a community also actively interact with their environment. In a
community
only those plants and animals survive which are adapted to a particular
environment. The
climate determines the type of environment, hence, the type of organisms in a
community.
For example, it is the climate of the area which determines whether a given
area becomes
a desert or a forest.
Communities created by human such as lawns or crop communities are such
man made
communication are crop communities are relatively simple and consists of only
one species
as opposed to a natural community characterized by a large number of
species. Man
made communities are very unstable and require great deal of care and
constant
manipulation and maintenance.
4.9.2 Stratification

Stratification of a community refers to the vertical layers of the vegetation.

Tropical forests
represent a good example of vertical stratification. In moist tropical rain forests
up to five
distinct strata or layers of vegetation can be formed. These include from the
forest floor to
the top (Fig. 4.14):
(i) Ground layer of mosses and liverworts
associated with dead leaves and other The bottom layer
substances rich in organic matter.
(i) Herb or grass layer, } The lower layer
(ii) Short shrub layer The middle layer
(iii) Tall shrub layer
(iv) Layer of under storey of short trees,
(v) Layer of canopy of lower tress and The upper layer
(vi) Over storey or emergent tree layer
formed by tall trees.
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Fig. 4.14: Stratification in a biotic community
As you can see in the fig. 4.14 the tropical forest the canopy dominates the
area. They
modify the light and moisture conditions for the shorter trees growing under
them, which in
turn determine the conditions for the ground vegetation. The vertical
stratification of the
plant community determines the structure of the community. The vegetation
provides a
number of habitats for the various organisms. Different layers of the
community are occupied
by different species of plants and animals. Plants and animals of each layer
differ in size,
behaviour and adaptation from those of other layers. The different layers of
organisms
minimize competition and conflict among the members of the community. The
various
species in the community compete with each other for nutrients, space, light
and other
resources. (refer again to Fig 4.4). Stratification is a practical strategy to
minimize interspecific
competition.
Community Characteristics
Species diversity
An important attribute of a community is its species diversity.
The different kinds of organisms present in a community represent its species
diversity.
The species composition or diversity differs from one community to another.

Even in the
same community, there may be seasonal variation in species composition.
Species diversity also influences the stability of the community. A stable
community is one
which is able to return to its original condition after being disturbed in some
way. Communities
with high species diversity have been found to be comparatively more stable.
Animal life mostly
flying organisms
Main habitat of
forest animals
Animals rarely
descend to forest
floor
Animal descend to
forest floor for
short period
Soil fauna
Emergent trees
Dense canaopy
of lower trees
Understory of
shorter trees
Tall shrubs
Low shrubs
Seedlings
scattered herbs
and a carpet of
rotting vegetation
Strata

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The diversity is calculated both by the number of species (richness) and the
relative
abundance of each species (evenness). Relative abundance is measure of
relative proportion
of different species occurring in a community. The greater the number of
species and more
even their distribution the greater is the species diversity.

4.10 ECOLOGICAL SUCCESSION

Biotic communities are dynamic in nature and change over a period of time.
The process
by which communities of plant and animal species in an area are replaced or
changed into
another over a period of time is known as ecological succession. Both the
biotic and
abiotic components are involved in this change. This change is brought about
both by the
activities of the communities as well as by the physical environment in that
particular area.
The physical environment often influences the nature, direction, rate and
optimal limit of
changes. During succession both the plant and animal communities undergo
change. There

are two types of successions (i) Primary succession and (ii) Secondary
succession.
4.10.1 Primary succession
Primary succession takes place an over a bare or unoccupied areas such as
rocks outcrop,
newly formed deltas and sand dunes, emerging volcano islands and lava flows
as well as
glacial moraines (muddy area exposed by a retreating glacier). where no
community has
existed previously. The plants that invade first bare land, where soil is initially
absent are
called pioneer species. The assemblage of pioneer plants is collectively called
pioneer
community. A pioneer species generally show high growth rate but short life
span (Fig
4.15)
Fig 4.15: The orderly sequence of primary succession
Bare rock Lichens Small annual
plants,
lichens
Perennial
herbs, grasses
Grasses, shrubs,
shade-intolerant trees
Shade-tolerant trees
Pioneer Stages Intermediate stages Climax community

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Primary succession is much more difficult to observe than secondary
succession because
there are relatively very few places on earth that do not already have
communities of
organisms. Furthermore, primary succession takes a very long time as
compared to
secondary succession as the soil is to be formed during primary succession
while secondary
succession starts in an area where soil is already present.
The community that initially inhabits a bare area is called pioneer
community. The pioneer
community after some time gets replaced by another community with different
species
combination. This second community gets replaced by a third community. This
process
continues sequence-wise in which a community replaced previous by another
community.
Each transitional (temporary) community that is formed and replaced during
succession is
called a stage in succession or a seral community (Fig. 4.16). The terminal
(final) stage of
succession forms the community which is called as climax community. A

climax community
is stable, mature, more complex and long lasting. The entire sequence of
communities in a
given area, succeeding each other, during the course of succession is termed
sere
(Fig 4.16).
The animals of such a community also exhibit succession which to a great
extent is determined
by plant succession. However animals of such successional stages are also
influenced by
the types of animals that are able to migrate from neighbouring communities.
A climax
community as long as it is undisturbed, remains relatively stable in dynamic
equilibrium
with the prevailing climate and habitat factors.
Succession that occurs on land where moisture content is low for e.g. on bare
rock is
known as xerarch. Succession that takes place in a water body, like ponds or
lake is
called hydrarch.
Fig. 4.16: Secondary succession on land
Annual
weeds
Grasses
and other
perennials
Shrubs Spruces Spruces Chestnut
Pines Immature oaks
Oak
Hickory
200 years (variable)
Chestnut Black
walnut
Tulip
poplar
Maple
Hickory
Oak
Intermediate Stage
Climax Community
Plowed
field
12 years
pioneer
220
years

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4.10.2 Secondary succession
Secondary succession is the development of a community which forms after
the existing
natural vegetation that constitutes a community is removed, disturbed or
destroyed by a
natural event like hurricane or forest fire or by human related events like tilling
or harvesting
land.

A secondary succession is relatively fast as, the soil has the necessary
nutrients as well as
a large pool of seeds and other dormant stages of organisms.

INTEXT QUESTIONS 4.4

1.Explain in brief (one to two sentences) the following ecological terms:

(i) Succession._______________________________________________________
(ii) Pioneer species. ___________________________________________________
(iii) Climax community _________________________________________________
(iv) Secondary succession ______________________________________________

4.11 BIOTIC INTERACTION

The biological community of an area or ecosystem is a complex network of

interactions.
The interaction that occurs among different individuals of the same species is
called
intraspecific interaction while the interaction among individuals of different
species in a
community is termed as interspecific interaction.
Interactions between organisms belonging to the same trophic level often
involve
competition. Individuals of population may compete for food, space and mates.
For
example if a mouse has been eaten by a cat, other cats competing for this
resource would
have one less mouse to prey on. The snake another predator of the mice would
also have
fewer mice to eat during the night if the cat has succeeded. Direct competition
though,
between the cat and snake is not much as they prey at different times. They
also eat a
variety of different foods. So competition may be intraspecific as well as
interspecific.
Interspecific relationship may be direct and close as between a lion and deer or
indirect
and remote as between an elephant and a beetle. This is because interactions
between two
species need not be through direct contact. Due to the connected nature of
ecosystems,
species may affect each other through intermediaries such as shared resources
or common
enemies. Specific terms are applied to interspecific interactions depending
upon whether
the interaction is beneficial, harmful or neutral to individuals of the species.
The various
possible interactions between two species are given in Table 4.1.
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Table 4.1: Possible biological interactions between two species.

S. No. Type of interaction 1 Result of Effects of interaction

species 2
I. NEGATIVE INTERACTIONS
i. Amensalism 0 one species is inhibited while the
other species is unaffected
ii. Predation + Predatorprey relationship: one
species (predator) benefits while
the second species (prey) is
harmed and inhibited.
iii. Parasitism + Beneficial to one species
(parasite) and harmful to the
other species (host).
iv. Competition 0 Adversely affects both species
II. POSITIVE ASSOCIATIONS
i. Commensalism + 0 One species (the commensal)
benefits, while the other species
(the host) is neither harmed nor
inhibited
ii. Mutualism + + Interaction is favourable to both
species
III. NEUTRAL INTERACTIONS
i. Neutralism 0 0 Neither species affects the other
+ =beneficial; = harmful, 0 =unaffected or neutral

Some types of interactions listed by the effects they have on each partner. O
is no
effect, is detrimental and + is beneficial.
4.11.1 Types of Interactions
From the table you can see that in certain types of interspecific associations at
least one of
the species is harmed by the other. Such associations are termed as negative,
in case
where both the associated species are benefited is a positive association and
when the
associated species are neither benefited nor harmed represents a neutral
interaction and
include:
1. Amensalism: This is a negative association between two species in which
one species
harms or restricts the other species without itself being adversely affected or
harmed
by the presence of the other species. Organisms that secrete antibiotics and
the species
that get inhibited by the antibiotics are examples of amensalism. For example
the
bread mould fungi Pencillium produce penicillin an antibiotic substance which
inhibits
the growth of a variety of bacteria. Pencillium benefits apparently by having
greater
availability of food when the competition because of the bacteria is removed.
2. Predation: In this type of interaction predator captures, kills and eats an
animal of
another species called the prey. The predator naturally benefits from this
relationship;
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while the prey is harmed. Predators like leopards, tigers and cheetahs use
speed, teeth
and claws to hunt and kill their prey.
3. Parasitism: In this type of interaction, one species is harmed and the other
benefits.
Parasitism involves parasite usually a small size organism living in or on
another living
species called the host from which the parasite gets its nourishment and often
shelter.
The parasite is benefited and the host is harmed. Many organisms like animal,
bacteria
and viruses are parasites of plants (Fig. 4.18a) and animals (Fig. 4.18b). Plants
like
dodder plant (Cuscuta) (Refer again to fig. 4.18a) and mistletoe (Loranthus )
are
parasites that live on flowering plants. Tap worm, round worm, malarial
parasite, many
bacteria, fungi, and viruses are common parasites of humans.
(a) (i) (ii)
Dodder, a parasitic plant is eating up a bush Dodder, a leafless parasitic plant,
growing on the leaf of a grass tree
(b) Ascaris lumbricoides infections. A mass of large round worms from a human infestation.

Fig. 4.17: Parasite-host relationship (a) Plant parasite: Dodder (Cuscuta) plant
is
a parasitic weed that obtains moisture and nourishment by attaching to a
green,
living plant. (b) Animal parasite: Ascaris or round worms are internal parasites
found in the human intestines
4. Competition: This is an interaction between two populations in which both
species
are harmed to some extent. Competition occurs when two populations or
species,
both need a vital resource that is in short supply. The vital resource could be
food,
water, shelter, nesting site, mates or space. Such competition can be: (i)
interspecific
competition-occurring between individuals of two different species occurring in
a habitat
and (ii) intraspecific competition-occurs between individuals of same species.
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Intraspecific competition occurs between members of the same species and so
it is
very intense.

5. Commensalism: In this relationship one of the species benefits while the

other is
neither harmed nor benefited. Some species obtain the benefit of shelter or
transport
from another species. For example sucker fish, remora often attaches to a
shark by
means of its sucker which is present on the top side of its head. This helps the
remora
get protection, a free ride as well as meal from the left over of the sharks meal
(Fig.
4.18). The shark does not however get any benefit nor is it adversely affected
by this
association. Another example of commensalisms is the relationship between
trees and
epiphytic plants. Epiphytes live on the surface of other plants like ferns, mosses
and
orchids and use the surface of trees for support and for obtaining sunlight and
moisture.
The tree gets no benefit from this relationship nor are they harmed.
Fig. 4.18: Commensalism: A shark with suckerfish
6. Mutualism: This is a close association between two species in which both
the species
benefit. For example of protocorporation the sea anemone, a cnidarian gets
attached
to the shell of hermit crabs for benefit of transport and obtaining new food
while the
anemone provides camouflage and protection by means of its stinging cells to
the
hermit crab (Fig. 4.19).
Fig. 4.19: Sea anemone, attached to a shell inhabited by a hermit crab
However, some mutualisms are so intimate that the interacting species can no
longer live
without each other as they depend totally on each other to survive. Such close
associations
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and Issues
are called symbiosis. An example of such close mutualistic association is that
of termite
and their intestinal flagellates. Termites can eat wood but have no enzymes to
digest it.
However, their intestine contains certain flagellate protists (protozoans) that
have the
necessary enzymes to digest the cellulose of the wood eaten by termites and
convert it into
sugar. The flagellates use some of this sugar for their own metabolism while
enough is left
for the termite. Both termite and flagellates cannot survive without each other.

Another
familiar example of symbiosis is seen in pollination of flowers where flowering
plants are
cross pollinated by the bees which benefit by getting nectar from the plants
and both
cannot survive without the other.
7. Neutralism: Neutralism describes the relationship between two species
which do
interact but do not affect each other. It is to describe interactions where the
fitness of
one species has absolutely no effect what so ever on that of other. True
neutralism is
extremely unlikely and impossible to prove. When dealing with the complex
networks
of interactions presented by ecosystems, one can not assert positively that
there is
absolutely no competition between or benefit to either species. Since true
neutralism is
rare or non-existent, its usage is often extended to situations where interaction
are
merely insignificant or neglible.

INTEXT QUESTIONS 4.5

1. Define (a) ecological succession, (b) symbiosis

______________________________________________________________
2. What type of competition exists between members of a deer herd in an
area?
______________________________________________________________
3. What type of relationship is represented by a garden spider feeding on a
grasshopper?
______________________________________________________________
4. What type of relationship is represented by a flower being pollinated by a
butterfly?
______________________________________________________________
5. Which term best fits the relationship of a person who has a lice feeding on
his scalp?
______________________________________________________________
6. Which term means two species live together with each providing a benefit to
the other
through the relationship?
______________________________________________________________
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and Issues

WHAT YOU HAVE LEARNT

Ecology may be defined as the scientific study of the relationships between

each other
and with their environment. The term ecology was coined by Ernst Harckel in
1869 .

 Ecology encompasses study of individual, organisms, population, community,

ecosystem, biome and biosphere which form the various levels of ecological
organization.
Habitat is the physical environment in which an organism lives (it corresponds
to address
of an organism).
Niche refers to the functional position of a species in its habitat.
Species is a group of populations whose individual members are capable of
interbreeding
with each other to produce a fertile offspring.
Evolution is the change which gives rise to new species. Mutation and
recombination
are sources of variation or differences in the genetic make up or gene pool of
a
species. Natural Selection is the mechanism proposed by Darwin and Wallace
which
interacts with variation to cause greater reproduction of these genes which
help in
adaptive to the environment.
Thus Evolution results in adaptation.
Evolution leads to speciation or formation of new species. Isolation is the
factor which
supports specimen. Isolation is of two major types (i) geographical isolation (ii)
reproductive isolation.
Many species, however, have been lost forever and not a single individual
belonging
to these species which once existed are now present. Extinction may occur due
to
catastrophic events in nature or due to human activities.
Population is a group of interbreeding individuals found in a specific time in a
particular
geographical area. The characteristics of a population become evident through
the (i)
population density (ii) birth rate or natality (iii) death rate or mortality (iv)
dispersion
(immigration and migration) (v) age distribution (vi) sex ratio.
Ecological succession is the successive growth of primary succession occurs
in an
area where there is no previous community. Secondary succession forms on
existing
natural vegetation.
Biotic interaction refers to the interaction taking place between individuals
belonging
to the same species (intra specific) or different species (interspecific).
Examples are (i)
competition (ii) predation (iii) parasitism (iv) mutualism (v) symbiosis (vi)
commensalism
(vii) neutralism
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TERMINAL EXERCISE

1. Define the terms : Ecology, niche, species, extinction.

2. What do you understand by Variation and Natural Selection? In what way
do they
interact to cause evolution?
3. What is the role of isolation in the formation of new species and keeping
them distinct.
4. In what ways have humans caused the extinction of species?
5. What do you understand by (i) natality (ii) speciation (iii) mutation (iv)
extinction
6. Explain ecological succession.
7. State and explain community characteristics.
8. What are (i) climax community and (ii) pioneer species?
9. Write an essay on biotic interaction.
10. Define biotic Interaction. Describe any one type of positive, negative and
neutral
quotation.

ANSWER TO INTEXT QUESTIONS

4.1
1. Ecology means the scientific study of the relationship of living organisms
with each
other and with their environment.
2. The term niche means the sum of all activities and relationship a species has
while
obtaining and using the resources it needs to survives and reproduce.
3. Habitat is the physical environment where an organism lives while niche is
the sum of
all activities and relationship of a species
4.2
1. The appearance or behaviour or structure or mode of life of an organism that
allows it
to survive in particular environment.
2. Species a group of similar populations of organisms whose members are
capable of
inter breeding to produce fertile offspring.
Variation- differences in structure due to differences in gene combinations.
3. (i) Gene combination (ii) Mutation
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4. Natural selection
5. Speciation- is the process by which new species are formed and extinction is
dying
out of a variety of or a species.

4.3
1. A Group of freely interbreeding individuals of the same species present in a
specific
area at a given time.
2. i) Density of the population
ii) Natality
iii) Mortality (any other)
3. Mortality, natility, immigration, emigration
4.4
1. (i) Succession is the orderly change of organisms in an environment over
time.
(ii) Pioneer species is the name use for the first time assemblage of plants that
inhabit
as area undergoing changes during succession. They are the first species in
successional
process.
(iii) Climax community is the final stage of succession. It is a relatively stable,
long
lasting community.
(iv) Secondary succession is the term used for the orderly series of change s
that
begins with the disturbance of an existing community and leads to a climax
community
4.5
1. The interacting species can no longer time without each other as they
depend totally
on each other to survive.
2. Intraspecific competition
3. Predation as it is preying upon or eating the grasshopper.
4. Mutualism as both are helped by the relationship.
5. Host
6. Mutualism