Spandrel

A spandrel, less often spandril or splaundrel, is the space between two arches or between an arch and a rectangular enclosure.

There are four or five accepted and cognate meanings of spandrel in architectural and art history, mostly relating to the space between a curved figure and a rectangular boundary - such as the space between the curve of an arch and a rectilinear bounding moulding, or the wallspace bounded by adjacent arches in an arcade and the stringcourse or moulding above them, or the space between the central medallion of a carpet and its rectangular corners, or the space between the circular face of a clock and the corners of the square revealed by its hood. Also included is the space under a flight of stairs, if it is not occupied by another flight of stairs. This is a common location to find storage space in residential structures.

In a building with more than one floor, the term spandrel is also used to indicate the space between the top of the window in one story and the sill of the window in the story above. The term is typically employed when there is a sculpted panel or other decorative element in this space, or when the space between the windows is filled with opaque or translucent glass, in this case called spandrel glass. In concrete or steel construction, an exterior beam extending from column to column usually carrying an exterior wall load is known as a spandrel beam.

Spandrel (biology)

In evolutionary biology, a spandrel is a phenotypic characteristic that is a byproduct of the evolution of some other characteristic, rather than a direct product of adaptive selection.

The term originated during the Roman era as an architectural word for the roughly triangular space between the tops of two adjacent arches and the ceiling. These spaces were not actually utilized until later on, when artists realized they could make designs and paint in these small areas, enhancing the overall design of the building. Stephen Jay Gould, a paleontologist at Harvard, and Richard Lewontin, a population geneticist, borrowed the word to apply to secondary byproducts of adaptations that were not necessarily adaptive in themselves.

Origin of the term

The term was coined by the Harvard paleontologist Stephen Jay Gould and population geneticist Richard Lewontin in their influential paper "The Spandrels of San Marco and the Panglossian Paradigm: A Critique of the Adaptationist Programme" (1979).