Rhynie chert

Coordinates: 57°20′12″N 002°50′29″W / 57.33667°N 2.84139°W / 57.33667; -2.84139

The Rhynie chert is an Early Devonian sedimentary deposit exhibiting extraordinary fossil detail or completeness (a Lagerstätte). It is exposed near the village of Rhynie, Aberdeenshire, Scotland; a second unit, the Windyfield chert, is located some 700 m away. The Rhynie chert contains exceptionally preserved plant, fungus, lichen and animal material preserved in place by an overlying volcanic deposit. The bulk of the fossil bed consists of primitive plants (which had water-conducting cells and sporangia, but no true leaves), along with arthropods, lichens, algae and fungi.

This fossil bed is remarkable for two reasons. First, the age of the site (Pragian, Early Devonian, formed about 410 million years ago) places it at an early stage in the colonisation of land. Second, these cherts are famous for their exceptional state of ultrastructural preservation, with individual cell walls easily visible in polished specimens. Stomata have been counted and lignin remnants detected in the plant material, and the breathing apparatus of trigonotarbids—of the class Arachnida—(known as book lungs) can be seen in cross-sections. Fungal hyphae can be seen entering plant material, acting as decomposers and mycorrhizal symbionts.