Plant litter
Litterfall, plant litter, leaf litter, tree litter, soil litter, or duff, is dead plant material, such as leaves, bark, needles, and twigs, that have fallen to the ground. This detritus or dead organic material and its constituent nutrients are added to the top layer of soil, commonly known as the litter layer or O horizon ("O" for "organic"). Litter has occupied the attention of ecologists at length for the reasons that it is an instrumental factor in ecosystem dynamics, it is indicative of ecological productivity, and may be useful in predicting regional nutrient cycling and soil fertility.
Characteristics and variability
Litterfall is characterized as fresh, undecomposed, and easily recognizable (by species and type) plant debris. This can be anything from leaves, cones, needles, twigs, bark, seeds/nuts, logs, or reproductive organs (e.g. the stamen of flowering plants). Items larger than 2 cm diameter are referred to as coarse litter, while anything smaller is referred to as fine litter or litter. The type of litterfall is most directly affected by ecosystem type.
For example, leaf tissues account for about 70 percent of litterfall in forests, but woody litter tends to increase with forest age. In grasslands, there is very little aboveground perennial tissue so the annual litterfall is very low and quite nearly equal to the net primary production.