The life cycle is similar to Toxoplasma. An infected dog will pass the oocysts through its feces and infect food or water. A cow or other animal will then up take the parasite. The parasite will undergo asexual reproduction in the animal's muscle until it is eaten by a dog. There, sexual reproduction will occur and oocysts will be created and passed through the feces.
Epidemiology
Dogs are often the definitive host but can act as an intermediate host as well. Cows are usually the intermediate host. No horizontal cow-to-cow transmission have been shown, although salival interactions have been suggested. Vertical transmission can occur when an infected cow gives birth to an infected calf—the calf survives the infection and grows into an adult.