The Iberian wolf differs from the more common Eurasian wolf with its slighter frame, white marks on the upper lips, the dark marks on the tail and a pair of dark marks in its front legs that give it its subspecies name, signatus ("marked"). The subspecies differentiation may have developed at the end of the PleistoceneIce Ages due to the isolation of the Iberian Peninsula when glacier barriers grew in the Pyrenees and eventually reached the Gulf of Biscay in the West and the Mediterranean in the East.