AVE no voladora

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"Flightless" vuelve a dirigir aquí. Para el sello discográfico, consulte Flightless
(sello discográfico) .

Los pingüinos son un ejemplo bien conocido de aves no voladoras.

Un kiwi Okarito ( Apteryx rowi ) también conocido como Rowi

Los avestruces son las aves no voladoras más grandes que existen, así como las aves
más grandes que existen en general.

An extinct Moa. Until the arrival of humans, New Zealand's only mammals were bats
and seals, resulting in many bird species evolving to fill the open niches. While
many of New Zealand's flightless birds are now extinct, some, such as the Kiwi,
Kakapo, Weka and Takahe are still around today.
Las aves no voladoras son aves que a través de la evolución perdieron la capacidad
de volar . [1] Hay más de 60 especies existentes, [2] incluidas las conocidas
ratites ( avestruces , emúes , casuarios , ñandúes y kiwis ) y pingüinos . El ave
no voladora más pequeña es el carril de la isla inaccesible (longitud 12,5 cm, peso
34,7 g). El ave no voladora más grande (tanto más pesada como más alta), que
también es el ave viva más grande, es el avestruz (2,7 m, 156 kg).

Muchas aves domesticadas, como el pollo y el pato domésticos , han perdido la

capacidad de volar durante períodos prolongados, aunque sus especies ancestrales,
el ave de la selva roja y el ánade real , respectivamente, son capaces de volar por
períodos prolongados. Algunas aves particularmente criadas, como el pavo blanco de
pecho ancho , se han vuelto totalmente incapaces de volar como resultado de la
reproducción selectiva ; las aves fueron criadas para producir una enorme carne de
pechuga que pesa demasiado para que las alas de las aves puedan soportar el vuelo.

La falta de vuelo ha evolucionado en muchas aves diferentes de forma independiente.

[3] Había familias de aves no voladoras, como las ahora extintas Phorusrhacidae ,
que evolucionaron para convertirse en poderosos depredadores terrestres. Llevando
esto a un extremo mayor, las aves del terror (y sus parientes los batornítidos ),
eogruidas , geranoides , gastornitiformes y dromornítidos(todos extintos) todos
desarrollaron formas corporales similares (patas largas, cuellos largos y cabezas
grandes), pero ninguno de ellos estaba estrechamente relacionado. Además, también
comparten rasgos de ser aves gigantes, no voladoras, con alas vestigiales, patas
largas y cuellos largos con algunas de las ratites, aunque no están emparentadas.
[4] [5]

Contenido
1 Orígenes de la falta de vuelo
1.1 Evolución independiente de la falta de vuelo en Palaeognathes
2 Cambios morfológicos y conservación de energía.
2.1 Presencia continuada de alas en aves no voladoras
3 Lista de aves no voladoras
3.1 Ratitas
3.2 Galliformes (aves de caza)
3.3 Anseriformes (aves acuáticas)
3.4 Aegotheliformes (chotacabras mochuelo)
3,5 Mesitornithiformes (mesites)
3.6 Columbiformes (palomas, palomas)
3,7 Gruiformes (grúas, rieles y fochas)
3.8 Podicipediformes (somormujos)
3.9 Charadriiformes (aves playeras y aliados)
3.10 Sphenisciformes (pingüinos)
3.11 Suliformes (piqueros, cormoranes y aliados)
3.12 Pelecaniformes (pelícanos, garzas, ibis y aliados)
3,13 Strigiformes (búhos)
3,14 Coraciiformes (martines pescadores y aliados)
3,15 Falconiformes (halcones y caracaras)
3,16 Psittaciformes (loros)
3,17 Passeriformes (aves que se posan)
4 Referencias
5 enlaces externos
Orígenes de la falta de vuelo
Divergences and losses of flight within ratite lineage occurred right after the K-
Pg extinction event wiped out all non-avian dinosaurs and large vertebrates 66
million years ago.[6] The immediate evacuation of niches following the mass
extinction provided opportunities for Palaeognathes to distribute and occupy novel
environments. New ecological influences selectively pressured different taxa to
converge on flightless modes of existence by altering them morphologically and
behaviorally. The successful acquisition and protection of a claimed territory
selected for large size and cursoriality in Tertiary ancestors of ratites.[7]
Temperate rainforests dried out throughout the Miocene and transformed into
semiarid deserts, causing habitats to be widely spread across the growingly
disparate landmasses. Cursoriality was an economic means of traveling long
distances to acquire food that was usually low-lying vegetation, more easily
accessed by walking.[7] Traces of these events are reflected in ratite distribution
throughout semiarid grasslands and deserts today.[8]

Gigantism and flightlessness are almost exclusively correlated.[clarification

needed] This is mostly observed in islands lacking predators and competition.
However, ratites occupy environments that are mostly occupied by a diverse number
of mammals.[9] It is thought that they first originated through allopatric
speciation caused by breakup of the supercontinent Gondwana.[10] However, later
evidence suggests this hypothesis first proposed by Joel Cracraft in 1974 is
incorrect.[11] Rather ratites arrived in their respective locations via a flighted
ancestor and lost the ability to fly multiple times within the lineage.

Gigantism is not a requirement for flightlessness. The kiwi do not exhibit

gigantism, along with tinamous, even though they coexisted with the moa and rheas
that both exhibit gigantism. This could be the result of different ancestral
flighted birds arrival or because of competitive exclusion.[10] The first
flightless bird to arrive in each environment utilized the large flightless
herbivore or omnivore niche, forcing the later arrivals to remain smaller. In
environments where flightless birds are not present, it is possible that after the
K/T Boundary there were no niches for them to fill. They were pushed out by other
herbivorous mammals.[9]

New Zealand had more species of flightless birds (including the kiwi, several
species of penguins, the takahe, the weka, the moa, and several other extinct
species) than any other such location. One reason is that until the arrival of
humans roughly a thousand years ago, there were no large land predators in New
Zealand; the main predators of flightless birds were larger birds.[12]

Independent evolution of flightlessness in Palaeognathes

Ratites belong to the superorder Palaeognathae, which include the volant tinamou,
and are believed to have evolved flightlessness independently multiple times within
their own group.[4][6][7][9] Some birds evolved flightlessness in response to the
absence of predators, for example on oceanic islands. Incongruences between ratite
phylogeny and Gondwana geological history indicate the presence of ratites in their
current locations is the result of a secondary invasion by flying birds.[13] It
remains possible that the most recent common ancestor of ratites was flightless and
the tinamou regained the ability to fly.[14] However, it is believed that the loss
of flight is an easier transition for birds than the loss and regain of flight,
which has never been documented in avian history.[7] Moreover, tinamou nesting
within flightless ratites indicates ancestral ratites were volant and multiple
losses of flight occurred independently throughout the lineage. This indicates that
the distinctive flightless nature of ratites is the result of convergent evolution.
[15]

Morphological changes and energy conservation

Two key differences between flying and flightless birds are the smaller wing bones
of flightless birds[16] and the absent (or greatly reduced) keel on their
breastbone. (The keel anchors muscles needed for wing movement.)[17]

Adapting to a cursorial lifestyle causes two inverse morphological changes to occur

in the skeleto-muscular system: the pectoral apparatus used to power flight is
paedorphically reduced while peramorphosis leads to enlargement of the pelvic
girdle for running.[10] Repeated selection for cursorial traits across ratites
suggests these adaptions comprise a more efficient use of energy in adulthood.[7]
The name "ratite" comes from the Latin ratis, raft, a vessel with no keel. Their
flat sternum is distinct from the typical sternum of flighted birds because it
lacks a keel, like a raft. This structure is the place where flight muscles attach
and thus allow for powered flight.[15] However, ratite anatomy presents other
primitive characters meant for flight, such as the fusion of wing elements, a
cerebellar structure, the presence of a pygostyle for tail feathers, and an alula
on the wing.[11] These morphological traits suggest some affinities to volant
groups. Palaeognathes were one of the first colonizers of novel niches and were
free to increase in abundance until the population was limited by food and
territory. A study looking at energy conservation and the evolution of
flightlessness hypothesized intraspecific competition selected for a reduced
individual energy expenditure, which is achieved by the loss of flight.[18]

Some flightless varieties of island birds are closely related to flying varieties,
implying flight is a significant biological cost.[18] Flight is the most costly
type of locomotion exemplified in the natural world. The energy expenditure
required for flight increases proportionally with body size, which is often why
flightlessness coincides with body mass.[8] By reducing large pectoral muscles that
require a significant amount of overall metabolic energy, ratites decrease their
basal metabolic rate and conserve energy.[18][19] A study looking at the basal
rates of birds found a significant correlation between low basal rate and pectoral
muscle mass in kiwis. On the contrary, flightless penguins exhibit an intermediate
basal rate. This is likely because penguins have well-developed pectoral muscles
for hunting and diving in the water.[18] For ground feeding birds, a cursorial
lifestyle is more economical and allows for easier access to dietary requirements.
[7] Flying birds have different wing and feather structures that make flying
easier, while flightless birds' wing structures are well adapted to their
environment and activities, such as diving in the ocean.[20]

A number of bird species appear to be in the process of losing their powers of

flight to various extents. These include the Zapata rail of Cuba, the Okinawa rail
of Japan, and the Laysan duck of Hawaii. All of these birds show adaptations common
to flightlessness, and evolved recently from fully flighted ancestors, but have not
yet completely given up the ability to fly. They are, however, weak fliers and are
incapable of traveling long distances by air.[21]

Continued presence of wings in flightless birds

Although selection pressure for flight was largely absent, the wing structure has
not been lost except in the New Zealand moas.[10] Ostriches are the fastest running
birds in the world and emus have been documented running 50 km/hr.[8] At these high
speeds, wings are necessary for balance and serving as a parachute apparatus to
help the bird slow down. Wings are hypothesized to have played a role in sexual
selection in early ancestral ratites and were thus maintained. This can be seen
today in both the rheas and ostriches. These ratites utilize their wings
extensively for courtship and displays to other males.[11] Sexual selection also
influences the maintenance of large body size, which discourages flight. The large
size of ratites leads to greater access to mates and higher reproductive success.
Ratites and tinamous are monogamous and mate only a limited number of times per
year.[22] High parental involvement denotes the necessity for choosing a reliable
mate. In a climatically stable habitat providing year round food supply, a male's
claimed territory signals to females the abundance of resources readily available
to her and her offspring.[19] Male size also indicates his protective abilities.
Similar to the emperor penguin, male ratites incubate and protect their offspring
anywhere between 85 and 92 days while females feed. They can go up to a week
without eating and survive only off fat stores. The emu has been documented fasting
as long as 56 days.[8] If no continued pressures warrant the energy expenditure to
maintain the structures of flight, selection will tend towards these other traits.

The only known species of flightless bird in which wings completely disappeared was
the gigantic, herbivorous moa of New Zealand, hunted to extinction by humans by the
15th century. In moa, the entire pectoral girdle is reduced to a paired
scapulocoracoid, which is the size of a finger.[23]

List of flightless birds

Many flightless birds are extinct; this list shows species that are either still
extant, or became extinct in the Holocene (no more than 11,000 years ago). Extinct
species are indicated with a dagger (†). A number of species suspected, but not
confirmed to be flightless, are also included here.

Longer-extinct groups of flightless birds include the Cretaceous

patagopterygiformes, hesperornithids, the Cenozoic phorusrhacids ("terror birds")
and related bathornithids, the unrelated eogruids, geranoidids, gastornithiforms,
and dromornithids (mihirungs or "demon ducks"), and the plotopterids.

Ratites

Common ostrich

North Island brown kiwi

Ostriches
Common ostrich, Struthio camelus
Somali ostrich, Struthio molybdophanes
Asian ostrich, Struthio asiaticus †
Emus
Emu, Dromaius novaehollandiae
King Island emu, Dromaius (novaehollandiae) minor †
Kangaroo Island emu, Dromaius (novaehollandiae) baudinianus †
Tasmanian emu, Dromaius novaehollandiae diemenensis †
Cassowaries
Dwarf cassowary, Casuarius bennetti
Southern cassowary, Casuarius casuarius
Northern cassowary, Casuarius unappendiculatus
Moa (Dinornithiformes) †, several species
Elephant birds (Aepyornithiformes) †, several species
Kiwis
Southern brown kiwi, Apteryx australis
Great spotted kiwi, Apteryx haastii
North Island brown kiwi, Apteryx mantelli
Little spotted kiwi, Apteryx owenii
Okarito kiwi, Apteryx rowi
Rheas
Greater rhea, Rhea americana
Lesser rhea, Rhea pennata
Galliformes (game birds)
New Caledonian giant scrubfowl, Sylviornis neocaledoniae †
Noble megapode, Megavitornis altirostris †
Viti Levu scrubfowl, Megapodius amissus †
Anseriformes (waterfowl)

Campbell teal
Auckland Island teal, Anas aucklandica
Campbell teal, Anas nesiotis
Steamer ducks
Fuegian steamer duck, Tachyeres pteneres
Falkland steamer duck, Tachyeres brachypterus
Chubut steamer duck, Tachyeres leucocephalus
Amsterdam wigeon, Anas marecula †
Bermuda flightless duck, Anas pachyscelus †
Finsch's duck, Chenonetta finschi †
New Zealand merganser, Mergus australis †
Moa-nalo †
Turtle-jawed moa-nalo, Chelychelynechen quassus †
Small-billed moa-nalo, Ptaiochen pau †
O'ahu moa-nalo, Thambetochen xanion †
Maui Nui large-billed moa-nalo, Thambetochen chauliodous †
Nēnē-nui, Branta hylobadistes † (possibly flightless or very weak flier)
Giant Hawaiʻi goose, Branta rhuax †
Mihirung, Genyornis newtoni †
California flightless sea-duck or Law's diving goose, Chendytes lawi †
Kaua'i mole duck, Talpanas lippa †
New Zealand geese, Cnemiornis gracilis and C. calcitrans †
Aegotheliformes (owlet-nightjars)
New Zealand owlet-nightjar, Aegotheles novaezealandiae †
Mesitornithiformes (mesites)
Brown mesite Mesitornis unicolor (possibly flightless, has not been seen flying)
[24]
Columbiformes (pigeons, doves)

Dodo
Dodo, Raphus cucullatus †
Rodrigues solitaire, Pezophaps solitaria †
Viti Levu giant pigeon, Natunaornis gigoura †
Saint Helena dove, Dysmoropelia dekarchiskos †
Henderson ground dove, Gallicolumba leonpascoi †
Gruiformes (cranes, rails, and coots)

Weka
Cuban flightless crane, Grus cubensis †
Red rail, Aphanapteryx bonasia †
Rodrigues rail, Erythromachus leguati †
Woodford's rail, Nesoclopeus woodfordi (most likely flightless)
Bar-winged rail, Nesoclopeus poecilopterus † (probably flightless)
Weka, Gallirallus australis
New Caledonian rail, Gallirallus lafresnayanus (likely †)
Lord Howe woodhen, Gallirallus sylvestris
Calayan rail, Gallirallus calayanensis
Pink-legged rail, Gallirallus insignis
Guam rail, Gallirallus owstoni
Roviana rail, Gallirallus rovianae (flightless, or almost so)[25]
Tahiti rail, Gallirallus pacificus †
Dieffenbach's rail, Gallirallus dieffenbachii †
Wake Island rail, Gallirallus wakensis †
numerous other unnamed Gallirallus rails from various Pacific islands
Chatham rail, Cabalus modestus †
Snoring rail, Aramidopsis plateni
Invisible rail, Habroptila wallacii
New Guinea flightless rail, Megacrex inepta
Aldabra (white-throated) rail, Dryolimnas (cuvieri) aldabranus
Réunion rail, Dryolimnas augusti †
Sauzier's wood rail or Cheke's wood rail, Dryolimnas chekei †
Inaccessible Island rail, Atlantisia rogersi
Saint Helena rail, Aphanocrex podarces †
Ascension crake, Mundia elpenor †
Saint Helena crake, Porzana astrictocarpus †
Laysan rail, Porzana palmeri †
Hawaiian rail, Porzana sandwichensis †
Small Maui crake, Porzana keplerorum †
Liliput crake, Porzana menehune †
Great Oʻahu crake, Porzana ralphorum †
Great Maui crake, Porzana severnsi †
Small Oʻahu crake, Porzana ziegleri †
Kosrae crake, Porzana monasa †
Henderson crake, Porzana atra
Mangaia crake, Porzana rua †
Tahiti crake, Porzana nigra †
numerous other unnamed Porzana crakes from various Pacific islands
Lord Howe swamphen, Porphyrio albus †
North Island takahē, Porphyrio mantelli †
Takahē, Porphyrio hochstetteri
Samoan woodhen, Gallinula pacifica
Makira woodhen, Gallinula silvestris
Tristan moorhen, Gallinula nesiotis †
Gough Island moorhen, Gallinula comeri
Tasmanian native hen, Tribonyx mortierii
Giant coot, Fulica gigantea (adults only; immature birds can fly)
Hawkins' rail, Diaphorapteryx hawkinsi †
Snipe-rail, Capellirallus karamu †
Antillean cave rail, Nesotrochis debooyi †
Hispaniolan cave rail, Nesotrochis steganinos †
Cuban cave rail, Nesotrochis picapicensis †
Adzebills, Aptornis otidiformis and A. defossor †
Podicipediformes (grebes)
Junín grebe, Podiceps taczanowskii
Titicaca grebe, Rollandia microptera
Atitlán grebe, Podilymbus gigas † (reportedly flightless)[26]
Charadriiformes (shorebirds and allies)

Great auk
Great auk, Pinguinus impennis †
Sphenisciformes (penguins)
King penguin, Aptenodytes patagonicus
Emperor penguin, Aptenodytes forsteri
Adélie penguin, Pygoscelis adeliae
Chinstrap penguin, Pygoscelis antarctica
Gentoo penguin, Pygoscelis papua
Little blue penguin, Eudyptula minor
Magellanic penguin, Spheniscus magellanicus
Humboldt penguin, Spheniscus humboldti
Galapagos penguin, Spheniscus mendiculus
African penguin, Spheniscus demersus
Yellow-eyed penguin, Megadyptes antipodes
Waitaha penguin, Megadyptes waitaha †
Fiordland penguin, Eudyptes pachyrhynchus
Snares penguin, Eudyptes robustus
Erect-crested penguin, Eudyptes sclateri
Northern rockhopper penguin, Eudyptes moseleyi
Southern rockhopper penguin, Eudyptes chrysocome
Royal penguin, Eudyptes schlegeli
Macaroni penguin, Eudyptes chrysolophus
Chatham penguin, Eudyptes warhami †
Suliformes (boobies, cormorants and allies)

Flightless cormorant
Flightless cormorant, Nannopterum harrisi
Pelecaniformes (pelicans, herons, ibises and allies)
Ascension night heron, Nycticorax olsoni †
Jamaican ibis, Xenicibis xymphithecus †
Hawaiian flightless ibises, Apteribis glenos and A. brevis †
Strigiformes (owls)
Cuban giant owl, Ornimegalonyx spp. † (possibly flightless)
Cretan owl, Athene cretensis † (probably flightless)
Andros Island barn owl, Tyto pollens † (possibly flightless)
Coraciiformes (kingfishers and allies)
Saint Helena hoopoe, Upupa antaios †
Falconiformes (falcons and caracaras)
Jamaican caracara, Caracara tellustris †
Psittaciformes (parrots)
Kakapo, Strigops habroptilus
Passeriformes (perching birds)
Lyall's wren, Xenicus lyalli †
Long-billed wren, Dendroscansor decurvirostris †
North Island stout-legged wren, Pachyplichas jagmi †
South Island stout-legged wren, Pachyplichas yaldwyni †
some Scytalopus tapaculos (possibly flightless, never seen flying)
Long-legged bunting, Emberiza alcoveri †
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External links
TerraNature pages on New Zealand flightless birds
Kiwi en Te Ara - la Enciclopedia de Nueva Zelanda
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