{{sfn | Beutel | Leschen | 2016 | p=}}

* {{cite book | last=Beutel | first=Rolf G. | last2=Leschen | first2=Richard A.B. | title=Coleoptera, Beetles. Morphology and Systematics | publisher=Walter de Gruyter GmbH & Co KG | publication-place=Berlin : Boston | date=2016-03-21 | isbn=978-3-11-037392-9}}

<ref name="r310">{{cite book | last=Beutel | first=Rolf G. | last2=Leschen | first2=Richard A.B. | title=Coleoptera, Beetles. Morphology and Systematics | publisher=Walter de Gruyter GmbH & Co KG | publication-place=Berlin : Boston | date=2016-03-21 | isbn=978-3-11-037392-9 | page=}}</ref>