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{{Short description|Asexual reproductive structure}}
[[Image:DroseraRoseanaGemmae.jpg|thumb|Pygmy sundew ''[[Drosera roseana]]'' with gemmae.]]
[[Image:DroseraRoseanaGemmae.jpg|thumb|Pygmy sundew ''[[Drosera roseana]]'' with gemmae.]]
A '''gemma''' (plural ''gemmae'') is a single [[cell (biology)|cell]], or a mass of cells, or a modified bud of tissue, that detaches from the parent and develops into a new individual. This type of asexual reproduction is referred to as [[fragmentation (reproduction)|fragmentation]]. It is a means of [[asexual reproduction|asexual]] propagation in [[plant]]s. These structures are commonly found in [[fungi]], [[alga]]e, [[Marchantiophyta|liverwort]]s and [[moss]]es, but also in some [[flowering plant]]s such as pygmy [[sundew]]s and some species of [[butterwort]]s.<ref name="Sundew">{{cite web | url=http://www.cascadecarnivores.com/index.php?cPath=34 | title=Pygmy Sundew Gemmae | publisher=Cascade Carnivores | work=Pygmy Sundew Gemmae | date=2011 | accessdate=December 14, 2011}}</ref><ref name="Hughes">{{cite book | title=On conidia of fungi, and gemmae of algae, bryophytes and pteridophytes | publisher=NRC Research Press | author=Hughes, Stanley John}}</ref> Vascular plants have many other methods of asexual reproduction including [[Bulbils]] and [[Turion (botany)|Turions]].
A '''gemma''' (English plural ''gemmas'', Latin plural ''gemmae'') is a single [[cell (biology)|cell]], or a mass of cells, or a modified bud of tissue, that detaches from the parent and develops into a new individual. This type of asexual reproduction is referred to as [[fragmentation (reproduction)|fragmentation]]. It is a means of [[asexual reproduction|asexual]] propagation in [[plant]]s. These structures are commonly found in [[fungi]], [[alga]]e, [[Marchantiophyta|liverwort]]s and [[moss]]es, but also in some [[flowering plant]]s such as pygmy [[sundew]]s and some species of [[butterwort]]s.<ref name="Sundew">{{cite web | url=http://www.cascadecarnivores.com/index.php?cPath=34 | title=Pygmy Sundew Gemmae | publisher=Cascade Carnivores | date=2011 | accessdate=December 14, 2011}}</ref><ref name="Hughes">{{cite book | title=On conidia of fungi, and gemmae of algae, bryophytes and pteridophytes | publisher=NRC Research Press | author=Hughes, Stanley John}}</ref>{{pn|date=March 2023}} Vascular plants have many other methods of asexual reproduction including [[bulbils]] and [[Turion (botany)|turions]].


== In mosses and liverworts ==
== In mosses and liverworts ==
[[File:Thallose liverwort (Marchantia and Lunularia spp.) showing clonal plantlets in gemma cups.jpg|thumb|Liverworts ''[[Marchantia polymorpha|Marchantia]]'' with round cups, and ''[[Lunularia]]'' with crescent cups, both containing gemmae. Gemmae dislodged by rain are visible at the bottom of the image]]
[[File:Thallose liverwort (Marchantia and Lunularia spp.) showing clonal plantlets in gemma cups.jpg|thumb|Liverworts ''[[Marchantia polymorpha|Marchantia]]'' with round cups, and ''[[Lunularia]]'' with crescent cups, both containing gemmae. Gemmae dislodged by rain are visible at the bottom of the image.]]


The production of gemmae is a widespread means of asexual reproduction in both liverworts and mosses. In liverworts such as ''[[Marchantia polymorpha|Marchantia]]'', the flattened plant body or thallus is a [[haploid]] [[gametophyte]] with gemma cups scattered about its upper surface. The gemma cups are cup-like structures containing gemmae. The gemmae are small discs of haploid tissue, and they directly give rise to new gametophytes. They are dispersed from gemma cups by rainfall.<ref>Smith, AJE (1989) ''The Liverworts of Britain and Ireland'', Cambridge University Press, Cambridge.</ref>
The production of gemmae is a widespread means of asexual reproduction in both liverworts and mosses. In liverworts such as ''[[Marchantia polymorpha|Marchantia]]'', the flattened plant body or thallus is a [[haploid]] [[gametophyte]] with gemma cups scattered about its upper surface. The gemma cups are cup-like structures containing gemmae. The gemmae are small discs of haploid tissue, and they directly give rise to new gametophytes. They are dispersed from gemma cups by rainfall.<ref>Smith, AJE (1989) ''The Liverworts of Britain and Ireland'', Cambridge University Press, Cambridge.</ref>{{pn|date=March 2023}}

The gemmae are bilaterally symmetrical and are not differentiated into dorsal and ventral surfaces. The mature gemmae fall on the ground and if conditions are suitable their germination starts immediately. The surface of the gemma which comes in contact of the soil gives out many rhizoids. This surface eventually becomes the lower(ventral) surface of the thallus. Meanwhile, the apical cells present in the two lateral notches become active and form two thalli in opposites directions.

[[Endogenous]] gemmae are also produced in liverworts, these are ovoid or ellipsoidal shaped, 2 celled at leaf tips or margins. Examples such as ''[[Bazzania kokawana]]'' ([[Fossombroniaceae]]), ''[[Endogemma|Endogemma caespiticia]]'' and also ''[[Riccardia]]'' species.<ref>Kishan Gopal Ramawat, Jean-Michel Merillon and K. R. Shivanna (Editors) {{google books|qXvSBQAAQBAJ|Reproductive Biology of Plants (2016)|page=70}}</ref>


The gemmae are bilaterally symmetrical and are not differentiated into dorsal and ventral surfaces. The mature gemmae fall on the ground and if conditions are suitable their germination starts immediately. The surface of the gemma which comes in contact of the soil gives out many rhizoids. This surface eventually becomes the lower(ventral) surface of the thallus. Meanwhile, the apical cells present in the two lateral notches become active and form two thalli in opposite directions.
==References==
==References==
{{reflist}}
{{reflist}}
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==External links==
==External links==
{{Commons category|Gemmae}}
{{Commons category|Gemmae}}
{{Wiktionary|gemma}}
* [https://web.archive.org/web/20110703113022/http://kentsimmons.uwinnipeg.ca/2152/lb7pg3.htm Marchantiales] - diagrams and micrographs of liverwort gemmae
* [https://web.archive.org/web/20110703113022/http://kentsimmons.uwinnipeg.ca/2152/lb7pg3.htm Marchantiales] diagrams and micrographs of liverwort gemmae


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[[Category:Plant reproduction]]
[[Category:Plant reproduction]]



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Latest revision as of 10:47, 21 December 2024

Pygmy sundew Drosera roseana with gemmae.

A gemma (English plural gemmas, Latin plural gemmae) is a single cell, or a mass of cells, or a modified bud of tissue, that detaches from the parent and develops into a new individual. This type of asexual reproduction is referred to as fragmentation. It is a means of asexual propagation in plants. These structures are commonly found in fungi, algae, liverworts and mosses, but also in some flowering plants such as pygmy sundews and some species of butterworts.[1][2][page needed] Vascular plants have many other methods of asexual reproduction including bulbils and turions.

In mosses and liverworts

[edit]
Liverworts Marchantia with round cups, and Lunularia with crescent cups, both containing gemmae. Gemmae dislodged by rain are visible at the bottom of the image.

The production of gemmae is a widespread means of asexual reproduction in both liverworts and mosses. In liverworts such as Marchantia, the flattened plant body or thallus is a haploid gametophyte with gemma cups scattered about its upper surface. The gemma cups are cup-like structures containing gemmae. The gemmae are small discs of haploid tissue, and they directly give rise to new gametophytes. They are dispersed from gemma cups by rainfall.[3][page needed]

The gemmae are bilaterally symmetrical and are not differentiated into dorsal and ventral surfaces. The mature gemmae fall on the ground and if conditions are suitable their germination starts immediately. The surface of the gemma which comes in contact of the soil gives out many rhizoids. This surface eventually becomes the lower(ventral) surface of the thallus. Meanwhile, the apical cells present in the two lateral notches become active and form two thalli in opposites directions.

Endogenous gemmae are also produced in liverworts, these are ovoid or ellipsoidal shaped, 2 celled at leaf tips or margins. Examples such as Bazzania kokawana (Fossombroniaceae), Endogemma caespiticia and also Riccardia species.[4]

References

[edit]
  1. ^ "Pygmy Sundew Gemmae". Cascade Carnivores. 2011. Retrieved December 14, 2011.
  2. ^ Hughes, Stanley John. On conidia of fungi, and gemmae of algae, bryophytes and pteridophytes. NRC Research Press.
  3. ^ Smith, AJE (1989) The Liverworts of Britain and Ireland, Cambridge University Press, Cambridge.
  4. ^ Kishan Gopal Ramawat, Jean-Michel Merillon and K. R. Shivanna (Editors) Reproductive Biology of Plants (2016), p. 70, at Google Books
[edit]