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Two days before the events of [[The Lion King]], Timon is a social outcast in his [[meerkat]] colony on the outskirts of the Pride Lands, as he frequently messes things up by accident. Though he is unconditionally supported by his [[List of The Lion King (franchise) characters#Ma|mother]], Timon dreams of a better life than his colony's bleak existence, continually hiding from predators, most notably the hyenas. One day, he is assigned as a sentry in an attempt by his mother to get him accepted, but his daydreaming leads to the near death of his uncle, [[List of The Lion King (franchise) characters#Uncle Max|Max]], at the hands of hyenas [[List of The Lion King (franchise) characters#Shenzi, Banzai, and Ed|Shenzi, Banzai and Ed]]. Now ostracized and convinced that he will never fit in with the other meerkats, Timon decides to leave to find a better life. After bidding his mother farewell, he meets the mandrill [[Rafiki (The Lion King)|Rafiki]], who teaches him about "Hakuna Matata", and advises him to "look beyond what you see". Timon takes this advice literally and observes Pride Rock in the distance. Believing Pride Rock to be his dream home, Timon ventures in that direction and encounters Pumbaa the [[warthog]] on his way. The two quickly form a bond, and Pumbaa accompanies Timon on his journey.
The pair arrive at Pride Rock during the presentation of [[Simba]] to the animals of the Pride Lands, which they mistake for a land rush and quickly scraps Pride Rock for their dream home. As they make their way through the crowd of onlookers, Pumbaa explosively passes gas due to his [[agoraphobia]], causing the nearby animals to faint, but prompting animals further away to bow to Simba. Following this, Timon and Pumbaa make multiple attempts to set up homes throughout the Pride Lands, but wind up being forced away every time upon witnessing several events from the original film, such as Simba singing "[[I Just Can't Wait to Be King]]", Mufasa's fight with the hyenas in the elephant graveyard, and [[List of The Lion King (franchise) characters#Scar|Scar]]'s conspiring with the hyenas. Eventually, the pair are caught in the wildebeest stampede that killed Mufasa, and are thrown off a waterfall in their attempt to escape. Exhausted, Timon decides to give up and realizes that Pumbaa (who sympathizes with Timon over being a social outcast) is the only friend that he has ever had. The next morning, Pumbaa discovers a luxurious green jungle that he had continuously tried to tell Timon about earlier. With their dream home found, they settle there, embracing "Hakuna Matata" as their life's philosophy.
Some time later, Timon and Pumbaa encounter Simba in a nearby desert, nearly dead. They rescue him and decide to raise him under their philosophy. Years later, Simba's childhood friend [[List of The Lion King (franchise) characters#Nala|Nala]] appears and reunites with him after chasing and mistaking Pumbaa for food. Believing Hakuna Matata to be in jeopardy, Timon and Pumbaa attempt to sabotage their dates, but fail every time. Upon witnessing Simba and Nala's argument, Simba disappears much to Timon thinking that they won. The next day, they and Nala learn from Rafiki that Simba has run off to challenge Scar and reclaim Pride Rock, meaning he would need their help. Upset that Simba left them, Timon decides to stay behind, but Pumbaa, annoyed that Timon would callously abandon Simba when he needs him, follows Simba and Nala. Timon indulges in the jungle's luxuries by himself, but loneliness starts to overwhelm him. After feeling sad and lonely, Timon is met with Rafiki, who indirectly (after Timon plays out an actual guess of how their conversation would turn out) makes him realize that his true Hakuna Matata is with the ones he loves,
Timon catches up and reconciles with Pumbaa before journeying onward to Pride Rock to join Simba and Nala. After helping Simba and Nala distract the hyenas
When the story finished, Ma, Uncle Max, Simba, Rafiki, and eventually many other Disney characters (such as [[Mickey Mouse]], [[Donald Duck]], [[Goofy]], [[Snow White (Disney character)|Snow White]] and the [[Seven Dwarfs]], [[Aladdin (Disney character)|Aladdin]] and [[Jasmine (Aladdin)|Jasmine]] with the [[Genie (Disney)|Genie]], [[Stitch (Lilo & Stitch)|Stitch]], [[Pocahontas (character)|Pocahontas]], [[Dumbo]], etc.) join Timon and Pumbaa to rewatch the film in the theater, during which Pumbaa tells Timon that he 'still doesn't do so well in crowds'.
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