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==Evolutionary history==
Fossil and molecular data suggest the genus ''Carya'' may have diversified during the [[Miocene]].<ref name="Zhang2013">{{cite journal |author1=Zhang, Jing-Bo |author2=Rui-Qi Li |author3=Xiao-Guo Xiang |author4=Steven R. Manchester |author5=Li Lin |author6=Wei Wang |author7=Jun Wen |author8=Zhi-Duan Chen |title=Integrated Fossil and Molecular Data Reveal the Biogeographic Diversification of the Eastern Asian-Eastern North American Disjunct Hickory Genus (Carya Nutt.) |journal=[[PLOS ONE]] |date=2013 |url=https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0070449}}</ref> Recent
{{Unreferenced section|date=May 2019}}
The earliest ancestors of hickories are identified from [[Cretaceous]] pollen grains.{{Citation needed}} The ''Carya'' as we know it first appears in [[Oligocene]] strata 34 million years ago. Fossils of early hickory nuts show simpler, thinner shells than modern species with the exception of [[pecan]]s, suggesting that the trees gradually developed defenses to [[rodent]] seed predation.{{Citation needed}} During this time, the genus had a distribution across the Northern Hemisphere, but the [[Pleistocene Ice Age]] beginning 2 million years ago completely obliterated it from Europe.{{Citation needed}} The distribution of Carya in North America also contracted and it completely disappeared from the continent west of the [[Rocky Mountains]]. Since fossil records show North America as having the largest number of Juglandaceae species, it is likely that the genus originated there and later spread to Europe and Asia.{{Citation needed}}
==Fruit==
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