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{{
{{Speciesbox
|
| image_caption = Male at Centennial Park, Sydney
| status = LC
| status_system = IUCN3.1
| status_ref = <ref name="iucn status 12 November 2021">{{cite iucn |
| genus = Anas
| species = superciliosa
| authority = [[Johann Friedrich Gmelin|Gmelin]], 1789
| range_map = PBD range.svg
|subdivision_ranks=Subspecies▼
| range_map_caption = Natural range of the Pacific black duck
▲| subdivision_ranks = Subspecies
| subdivision = See text
}}
The '''Pacific black duck''' ('''''Anas superciliosa''
==Taxonomy==
[[File:Forster Anas superciliosa.jpg|thumb|Watercolour made by [[Georg Forster]] on [[James Cook]]'s [[Second voyage of James Cook|second voyage to the Pacific Ocean]]. This painting is the [[holotype]] for the species.]]
The Pacific black duck was [[Species description|formally described]] in 1789 by the German naturalist [[Johann Friedrich Gmelin]] in his revised and expanded edition of [[Carl Linnaeus]]'s ''[[Systema Naturae]]''. He placed it with all the other ducks, geese and swans in the [[genus]] ''[[Anas]]'' and coined the [[binomial nomenclature|binomial name]] ''Anas superciliosa''.<ref>{{ cite book | last=Gmelin | first=Johann Friedrich | author-link=Johann Friedrich Gmelin| year=1789 | title=Systema naturae per regna tria naturae : secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis | edition=13th | volume=1, Part 2 | language=Latin | location=Lipsiae [Leipzig] | publisher=Georg. Emanuel. Beer | page=537 | url=https://www.biodiversitylibrary.org/page/2656030 }}</ref> Gmelin based his description on the "Supercilious duck" that had been described in 1785 by the English ornithologist [[John Latham (ornithologist)|John Latham]] in his ''A General Synopsis of Birds''.<ref>{{ cite book | last=Latham | first=John | author-link=John Latham (ornithologist) | year=1785 | title=A General Synopsis of Birds | volume=3, Part 2 | publisher=Printed for Leigh and Sotheby | location=London | page=497, No. 45 | url=https://www.biodiversitylibrary.org/page/40079318 }}</ref><ref>{{ cite book | editor1-last=Mayr | editor1-first=Ernst | editor1-link=Ernst Mayr | editor2-last=Cottrell | editor2-first=G. William | year=1979 | title=Check-List of Birds of the World | volume=1 | edition=2nd | publisher=Museum of Comparative Zoology | place=Cambridge, Massachusetts | page=472 | url=https://www.biodiversitylibrary.org/page/16109112 }}</ref> The naturalist [[Joseph Banks]] had provided Latham with a water-colour drawing of the duck by [[Georg Forster]] who had accompanied [[James Cook]] on his [[Second voyage of James Cook|second voyage to the Pacific Ocean]]. His picture was drawn at [[Dusky Sound]], a fiord on the southwest corner of New Zealand. This picture is the [[holotype]] for the species and is now held by the [[Natural History Museum, London|Natural History Museum]] in London.<ref>{{ cite journal | last=Medway | first=David G. | date=1976 | title=Extant types of New Zealand birds from Cook's voyages. Part 1: Historical and type paintings | journal=Notornis | volume=23 | issue=1 | pages=45-60 [54, "Grey duck"] | url=https://www.birdsnz.org.nz/wp-content/uploads/2021/12/Notornis_23_1.pdf }}</ref><ref>{{ cite journal | last=Lysaght | first=Averil | date=1959 | title=Some eighteenth century bird paintings in the library of Sir Joseph Banks (1743-1820) | journal=Bulletin of the British Museum (Natural History), Historical Series | volume=1 | issue=6 | pages=251-371 [290, No. 77] | doi=10.5962/p.92313 | url=https://www.biodiversitylibrary.org/page/2238991 | doi-access=free }}</ref> The genus name ''Anas'' is the Latin word for a duck. The specific epithet ''superciliosa'' is from Latin meaning "supercilious" or "eye-browed", a reference to the prominent [[supercilium]] or eye-stripe.<ref>{{cite book | last=Jobling | first=James A. | year=2010| title=The Helm Dictionary of Scientific Bird Names | publisher=Christopher Helm | location=London | isbn=978-1-4081-2501-4 | pages=[https://archive.org/stream/Helm_Dictionary_of_Scientific_Bird_Names_by_James_A._Jobling#page/n46/mode/1up 46], [https://archive.org/stream/Helm_Dictionary_of_Scientific_Bird_Names_by_James_A._Jobling#page/n374/mode/1up 374]}}</ref>
Two [[subspecies]] are now recognised:<ref name=ioc>{{cite web| editor1-last=Gill | editor1-first=Frank | editor1-link=Frank Gill (ornithologist) | editor2-last=Donsker | editor2-first=David | editor3-last=Rasmussen | editor3-first=Pamela | editor3-link=Pamela Rasmussen | date=January 2022 | title=Screamers, ducks, geese & swans | work=IOC World Bird List Version 12.1 | url=http://www.worldbirdnames.org/bow/waterfowl/ | publisher=International Ornithologists' Union | access-date=4 July 2022 }}</ref>
* ''A.
* ''A. s. superciliosa'' Gmelin, JF, 1789 − Australasian duck, breeds in Indonesia, southern New Guinea, Australia and New Zealand, where it is known as the grey duck or pārera.<ref name=nzbirds>{{Cite web |title=Grey duck {{!}} Pārera {{!}} New Zealand Birds Online |url=https://nzbirdsonline.org.nz/species/grey-duck |access-date=2022-06-24 |website=nzbirdsonline.org.nz}}</ref>
A third subspecies, ''rogersi'' from Australia, has sometimes been recognised but it not distinguishable either genetically or phenotypically from the [[nominate subspecies|nominate race]].<ref name=nzbirds/>
==Description==
This sociable [[duck]] is found in a variety of wetland habitats, and its nesting habits are much like those of the [[mallard]], which is encroaching on its range in New Zealand.<ref name=Rhymer1996/> It feeds by upending, like other ''Anas'' ducks.
It has a dark body, and a paler head with a dark crown and facial stripes. In flight, it shows a green [[speculum feathers|speculum]] and pale underwing. All plumages are similar. The size range is 54–61 cm; males tend to be larger than females, and some island forms are smaller and darker than the main populations.<ref name=Madge/> It is not resident on the [[Marianas]] islands, but sometimes occurs there during migration. The now
Like its relatives the [[mallard]] and [[American black duck]], the Pacific black duck is one of a number of duck species that can quack, with the female producing a sequence of raucous, rapid quacking which decreases in volume.<ref>{{Cite web|url=https://collections.
==Behaviour==
===Breeding===
The nest is usually placed in a hole in a tree, but sometimes an old nest of a corvid is used and occasionally the nest will be placed on the ground. The clutch of 8–10 pale cream eggs is incubated only by the female. The eggs hatch after 26–32 days. The [[precocial]] downy ducklings leave the nest site when dry and are cared for by the female. They can fly when around 58 days of age.<ref name=hanzab>{{cite book | editor1-last=Marchant | editor1-first=S. | editor2-last=Higgins | editor2-first=P.G. | year=1990 | chapter=''Anas superciliosa'' Pacific Black Duck | title=Handbook of Australian, New Zealand & Antarctic Birds. Volume 1: Ratites to ducks; Part B, Australian pelican to ducks | location=Melbourne, Victoria | publisher=Oxford University Press | isbn=978-0-19-553068-1 | pages=1320–1332 | chapter-url=https://nzbirdsonline.org.nz/sites/all/files/030_Grey%20Duck.pdf }}</ref>
▲* ''A. superciliosa pelewensis'' {{small|[[Gustav Hartlaub|Hartlaub]] & [[Otto Finsch|Finsch]], 1872}} – Island black duck, breeds on the southwest Pacific islands and northern [[New Guinea]]
===Feeding===▼
The New Zealand subspecies has declined sharply in numbers, at least in its pure form, due to competition from and hybridisation with the introduced mallard.<ref name=Gillespie/> Rhymer ''et al.'' (1994) say their data "points to the eventual loss of identity of the grey duck as a separate species in New Zealand, and the subsequent dominance of a hybrid swarm akin to the Mariana Mallard." Studies of their three species of parasitic feather lice support this prediction<ref>{{Cite journal|title=The ectoparasites of hybrid ducks in New Zealand (Mallard x Grey Duck)|last=Bulgarella|first=M|date=2018|journal=International Journal for Parasitology: Parasites and Wildlife|volume=7|issue=3|pages=335–342|doi=10.1016/j.ijppaw.2018.09.005|pmc=6154467|pmid=30258780}}</ref>▼
The Pacific black duck is mainly vegetarian, feeding on seeds of aquatic plants. This diet is supplemented with small crustaceans, molluscs and aquatic insects. Food is obtained by 'dabbling', where the bird plunges its head and neck underwater and upends, raising its rear end vertically out of the water. Occasionally, food is sought on land in damp grassy areas.<ref name=TAM/>▼
==Conservation status==
▲The
It was assumed that far more mallard drakes mate with grey duck females than vice versa based on the fact that most hybrids show a mallard-type plumage, but this is not correct; it appears that the mallard phenotype is [[Dominance (genetics)|dominant]], and that the degree to which species contributed to a hybrid's ancestry cannot be determined from the plumage.<ref name=Rhymer1994/> The main reasons for displacement of the grey duck seem to be physical dominance of the larger mallards, combined with a marked population decline of the grey duck due to overhunting in the mid-20th century.<ref name=Williams/>
▲==Feeding==
▲The Pacific black duck is mainly vegetarian, feeding on seeds of aquatic plants. This diet is supplemented with small crustaceans, molluscs and aquatic insects. Food is obtained by 'dabbling', where the bird plunges its head and neck underwater and upends, raising its rear end vertically out of the water. Occasionally, food is sought on land in damp grassy areas.<ref name=TAM/>
==Various views and plumages==
<gallery>
File:Pacific Black Duck - AndrewMercer - DSC07814.jpg|Female with ducklings
File:PBD mallard hybrid (1).jpg|mallard x pacific black duck hybrid, Hobart
File:PBD hybrid.jpg|mallard x pacific black duck hybrid, Richmond Tasmania
</gallery>
==See also==
* ''[[Dalvirus anatis]]''
==References==
{{Reflist|refs=
<ref name=Gillespie>{{cite journal|last=Gillespie|first=Grant D|year=1985|title=Hybridization, introgression, and morphometric differentiation between Mallard (''Anas platyrhynchos'') and Grey Duck (''Anas superciliosa'') in Otago, New Zealand|journal=The Auk|volume=102|issue=3|pages=459–469|url=http://sora.unm.edu/sites/default/files/journals/auk/v102n03/p0459-p0469.pdf|doi=10.1093/auk/102.3.459}}</ref>
<ref name=Madge>{{Cite book|
<ref name=Rhymer1996>{{cite journal|author1=Rhymer, Judith M. |author2=Simberloff, Daniel |name-list-style=amp |year=1996|title=Extinction by hybridization and introgression|journal=Annual Review of Ecology and Systematics|volume=27|doi=10.1146/annurev.ecolsys.27.1.83|pages=83–109}}</ref>
<ref name=Rhymer1994>{{cite journal|author1=Rhymer, Judith M. |author2=Williams, Murray J. |author3=Braun, Michael J |name-list-style=amp |year=1994|title=Mitochondrial analysis of gene flow between New Zealand Mallards (''Anas platyrhynchos'') and Grey Ducks (''A. superciliosa'')|journal=The Auk|volume=111|issue=4|pages=970–978|url=http://sora.unm.edu/sites/default/files/journals/auk/v111n04/p0970-p0978.pdf|doi=10.2307/4088829|jstor=4088829 }}</ref>
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<ref name=TAM>{{cite web |url=https://australianmuseum.net.au/learn/animals/birds/pacific-black-duck/ |title=Pacific Black Duck |publisher=The Australian Museum |access-date=March 6, 2019}}</ref>
}}
{{Taxonbar|from=Q757271}}
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[[Category:Birds of the Pacific Ocean]]
[[Category:Birds described in 1789]]
[[Category:Taxa named by Johann Friedrich Gmelin]]
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